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Chapter 2
Lecture Outline
Tissues
Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display.
Outline
Organs and Tissues
Meristematic Tissues
• Apical Meristems
• Lateral Meristems
• Intercalary Meristems
Tissues Produced by Meristems
• Simple Tissues
• Complex Tissues
Organs and Tissues
Plants have three or four major groups of organs:• Roots• Stems• Leaves• Flowers
Each organ is composed of tissues.
• A tissue is a group of cells performing a similar function.
• There may be more than one tissue per organ.
Meristematic TissuesApical Meristems
Meristems - Permanent regions of growth and active cell division
Apical Meristems - Found at the tips of roots and shoots• Roots and shoots increase in length as the apical
meristems produce new cells (= primary growth).
Meristematic TissuesApical Meristems
• Primary meristems develop from apical meristems.– Protoderm
– Ground Meristem
– Procambium
• Primary meristems produce primary tissues.
Meristematic TissuesLateral Meristems
Lateral Meristems - Produce secondary tissues that increase the girth of roots and stems (= secondary growth)
• Vascular Cambium - Produces tissues that function primarily in support and conduction.– Composed of a thin cylinder of brick-shaped cells that
extends the length of stems and roots
• Cork Cambium - Lies outside vascular cambium just inside the outer bark– Produces bark
Meristematic TissuesIntercalary Meristems
Grasses and related plants do not have vascular cambium nor cork cambium.
• They have intercalary meristems.
• Intercalary Meristems– In vicinity of nodes (leaf attachment area)
– Add to stem length
Tissues Produced By MeristemsSimple Tissues
Parenchyma - Composed of parenchyma cells• Parenchyma cells:
– Thin, pliable walls– Usually 14-sided at maturity– Living cytoplasm, often
containing large vacuoles and various secretions
– May remain alive a long time– Have spaces between them
Parenchyma cells
Simple Tissues - Consist of only one kind of cell
Tissues Produced By MeristemsSimple Tissues
• Types of parenchyma tissue and cells:– Aerenchyma - Parenchyma tissue with extensive
connected air spaces, usually in aquatic plants
– Chlorenchyma - Parenchyma cells containing chloroplasts that function in photosynthesis
– Transfer cells - Develop irregular extensions of inner wall that greatly increase surface area of plasma membraneo Nectaries of flowers
Tissues Produced By MeristemsSimple Tissues
Collenchyma tissue - Contains collenchyma cells
• Collenchyma cells:– Living cytoplasm
– May remain alive a long time
– Cell walls thick, with uneven thickness
– Pliable and strong, thus providing flexible support
Collenchyma cells
Tissues Produced By MeristemsSimple Tissues
Sclerenchyma tissue - Contains sclerenchyma cells• Sclerenchyma cells:
– Thick, tough, secondary walls, normally impregnated with lignin
– Dead at maturity– Function in support– Two types:
o sclereids o fibers
Tissues Produced By MeristemsSimple Tissues
o Sclereids – Stone Cells
– Scattered in tissue
– Cells as long as wide
Tissues Produced By MeristemsSimple Tissues
o Fiberso Much longer that wide and contain lumen (tiny cavity)
Fibers in cross section and longitudinal section
Tissues Produced By MeristemsComplex Tissues
Complex tissues - Composed of two or more kinds of cells
• Vascular tissues include xylem and phloem.
Xylem - Chief conducting tissue for water and minerals that are absorbed by the roots
• Composed of parenchyma cells, fibers, vessels, tracheids and ray cells
Tissues Produced By MeristemsComplex Tissues
• Vessels – Long tubes made of vessel elements– Vessel elements:
o Thick secondary cell walls
o Open at both endso Secondary walls
develop irregularlyo Perforation plate
between end walls
Vesselselements
Spiral thickenings on vessel
walls
Tissues Produced By MeristemsComplex Tissues
• Tracheids – Tapered at the ends with pairs
of pits that allow water to pass from cell to cello Pits - Areas without
secondary cell– Dead at maturity– Thick secondary cell walls– May have spiral thickenings on
cell walls
• Rays - Function in lateral conduction and food storage– Composed of long-lived parenchyma cells
Tracheids
Tissues Produced By MeristemsComplex Tissues
Phloem - Conducts dissolved food materials produced by photosynthesis throughout plant • Composed of sieve tube members, companion cells,
fibers, parenchyma cells and ray cells
Tissues Produced By MeristemsComplex Tissues
• Sieve Tube Members:– Lack secondary cell walls and nuclei– Lay end to end to form sieve tubes– Walls have sieve plates with small
pores– Callose forms callus plug - Prevents
leaking of sieve tube contents when cell injured
Phloem
• Companion cells - Aid in conduction of food
Tissues Produced By MeristemsComplex Tissues
Epidermis- Protective layer that is one cell-layer thick covering all plant organs
• Composed mostly of parenchyma cells, guard cells of stomata, secretory glands and hairs
Leaf cross section
Tissues Produced By MeristemsComplex Tissues
• Cutin - Fatty substance on the surface of outer walls of epidermis that forms cuticle– Wax secreted on cuticle
– Cuticle and wax prevent water loss by evaporationo Resistant to bacteria and other disease organisms
• Root epidermal cells produce root hairs.– Increase absorptive area of root surface
• Leaves have stomata bordered by pairs of guard cells.
Tissues Produced By MeristemsComplex Tissues
• Constitutes outer bark
• Primarily composed of cork cells– Dead at maturity
– While still alive, cytoplasm secretes suberin (fatty substance) into walls.
o Makes cork cells waterproof and helps protect phloem
Periderm - Replaces epidermis when cork cambium begins producing new tissue
– Lenticels - Loosely arranged pockets of parenchyma cells formed by cork cambium that protrude through the surface of periderm. Allows for gaseous exchange.
Tissues Produced By MeristemsComplex Tissues
Secretory Cells and Tissue
• Secretory cells may function individually or as part of a secretory tissue.– Flower nectar
– Citrus oils
– Glandular hair mucilage
– Latex
– Resins
Review
Organs and Tissues
Meristematic Tissues
• Apical Meristems
• Lateral Meristems
• Intercalary Meristems
Tissues Produced by Meristems
• Simple Tissues
• Complex Tissues