N. Jb . Geol. Paläont. Mh. 1996, H. 5 293-308 Stuttgart , Mai 1996

Remarks on the genera Striatoporella and “Riphaeolites” (Devonian Tabulata)

By K lem en s O eken torp and Tong-D zuy Thanh

With 4 figures in the text

O e k e n t o r p , K. & T o n g - D z u y T h a n h (1996): Remarks on the genera Striatoporella and “Riphaeolites” (Devonian Tabulata). - N . Jb . Geol. Paläont. M h., 1996 (5): 2 93-308 ; Stuttgart.

A bstrac t : Restudy o f Mesofavosites schouppei P o t t h a s t & O e k e n t o r p 1988 (Emsian-Eifelian boundary in SE Morrocco) and o f Pachyfavosites pseudoseptatus O e k e n t o r p 1975 (Emsian o f A sturias/N Spain) shows that M . schouppei has to be assigned to Striatoporella R u k h i n 1938 because o f the typical growth form, the peripheral thickenings o f the walls and the developm ent o f spines. P. pseudosep­tatus has been assigned to Pachyfavosites on the basis o f its strikingly thick walls. Diagenetic processes may result in secondary microstructures, for example the arrangement o f carbonate fibers. Therefore, wall thickness is useless as a diagnostic feature. In this view, P. pseudoseptatus corresponds very well with other representatives o f the genus Riphaeolites.

Z u sa m m e n fa s su n g : Neuuntersuchungen von Genera tabulater Korallen u. a. in Hinblick a u f mögliche diagenetische Mikrostruktur-Überprägungen ergaben neue Erkenntnisse über die systematische Zuordnung von Mesofavosites schouppei P o t t h a s t & O e k e n t o r p 1988 aus dem Grenzbereich Em sium -Eifelium SE Marokkos und von Pachyfavosites pseudoseptatus O e k e n t o r p 1975 aus dem Em sium von A sturien /N Spanien. - Aufgrund der charakteristischen Wuchsform, der Zunahme der Wandverdickungen zur Peripherie hin und der Ausbildung von D ornen wird M . schouppei der Gattung Striatoporella R u k h i n 1938 zugeschrieben.- Die aufgrund der stark verdickten Wände bisher der Gattung Pachyfavosites zugewiesene Art P. pseudoseptatus erweist sich als Vertreter von Riphaeolites Y a n e t

1955. Die neuerliche Untersuchung ergab, daß die Wandverdickungen das Resultat frühdiagenetischer Umkristallisationsprozesse darstellen. Kenn­zeichnend für die abiogene Natur sind Kristallvergrößerungen mit einherge­hender stärkerer Aufhellung und die Ausbildung neuer, unregelmäßiger, fächer­förmiger Kristallanordnungen. Diese können aus dem primären Skelett heraus­wachsen und damit Skelettverdickungen vortäuschen. Es wird hier dann zwar der architektonische Rahmen des Skeletts weitgehend beibehalten, jedoch in seinen Maßen und in seinem mikrostrukturellen Feinbau völlig überprägt. Damit entfallt die Anwendungsmöglichkeit der Mikrostruktur und der Wandver­dickungen für systematische Belange. Aufgrund der übrigen Merkmale ergab sich somit die Zuordnung des spanischen Materials zu Riphaeolites.

0028-3630 /96/1996-0293 $ 4.001 996 E . S ch w eiz erb a rt’ sche V e rla g sb u c h h a n d lu n g , D -70176 S tu ttg ar t

2 9 4 Klem ens O ekentorp and Tong-D zuy Thanh

Introduction

During a three month stay at the Forschungsstelle für Korallenpaläo- zoologie, University Münster/Westf., Germany, the junior author disco­vered some interesting features in tabulate corals stored in the collec­tions o f the Forschungsstelle. These concern a species o f Striatoporella previously described as “Mesofavosites” schouppei P o t t h a s t & O e k e n t o r p

(1988) from the upper Lower Devonian o f Morrocco as well as the enigmatic genus “Riphaeolites”, described by Ya n e t (1955). To the latter one we like to assign the species “Pachyfavosites” pseudoseptatus O e k e n t o r p 1975, from the Emsian o f Asturias, Northern Spain. Restudy o f the Spanish material showed significant diagenetic microstructures, which call for a critical discussion o f the genus “Riphaeolites”. The junior author wishes to express his gratitude to the Deutsche Akademische Austauschdienst (DAAD) for financing his research stay in Germany and to the Vietnamese Program for Fundamental Research in the Field o f Natural Science for their kind support. Additionally, he thankfully remembers the Forschungsstelle für Korallenpaläozoologie o f the Westfälische Wilhelms-Universität, Münster.

Fam ily Favosit idae D a n a , 1846

Genus Striatoporella R u k h i n , 1938

1938 Striatoporella R u k h i n , p. 62 1869 Striatoporella - D u b a t o l o v , p. 80

e.p. 1955 Thamnopora - S o k o l o v , p. 175 1959 Favosites - H a m a d a , p. 201

e.p. 1962 Thamnopora - S o k o l o v , 229 e.p. 1963 Favosites - S o k o l o v & T e s a k o v , 128

1964 Rudakites L e l e s h u s , p. 46 1967 Squameopora P r e o b r a z h e n s k i y , p. 8

e.p. 1968 Favosites - C h e k h o v i c h , p. 43 e.p. 1969 Favosites - D u b a t o l o v , p. 53 e.p. 1971 Thamnopora - S o k o l o v in O r l o v , p. 229

1981 Striatoporella - H i l l in M o o r e , R o b i s o n & T e i c h e r t , p. 546 1988 Mesofavosites - P o t t h a s t & O e k e n t o r p , p. 76 1988 Squameopora - T o n g - D z u y , T h a n h et al., p. 61

e.p. 1990 Favosites - D u b a t o l o v , p. 56

T y p e s p e c i e s : Striatoporella multiporifera R u k h i n . - The holotype o f thespecies is lost. A neotype has been chosen by V. N. D u b a t o l o v 1969 (Page 80;№ 109, coll. 337; Lower D evonian ; Kolym a basin, Northeast Sibiria. Conserved at the Institute o f Geology and Geophysics Novosibirsk, Russia).

D i a g n o s i s : Branching and nodulose corallum with coarse cylindrical outgrowing. Corallite polygonal in the transverse section, directed longitudi­nally in the axial zone and regularly curving outward to the surface o f the

Remarks on the genera Striatoporella and “Riphaeolites“ 2 9 5

corallum. Walls slightly thickened to the peripheral zone. Mural pores may be situated to the corner o f the corallites. Septal spines, represented by short, coarse spines and squamulae, are irregularly developed in the axial and more frequent in the peripheral zone. Tabulae may be thickened in the periphery o f the corallum.

R em arks

1. Branching and nodulose coralla o f favositids with cylindrical outgrowings are widespread in the U pper Silurian and particularly in the Lower Devonian. Although m ost o f them have been described as species o f the genus Favosites, their taxonomic positions attract the attention o f palaeontologists. It seems that these forms show a transition from branching favositids to pachyporids (Thamnopora) but it is difficult to find diagnostic features for separating Favosites-like Thamnopora from Thamnopora-Yike. Favosites species ( P r e o b r a z h e n s k i y 1967, H a m a d a 1959) (compare L e c o m p t e 1936, 1939).

2. Describing the genus Striatoporella with a nodulose branching corallum L. B. R u k h i n noted its affinities with favositids but not pachy- porids. The illustrations in R u k h i n ’ s descriptions (1938) are not clear enough and, unfortunately, all his material was lost during the Second World War. As a result, the genus Striatoporella has been forgotten for long time or was regarded as a junior synonym o f Thamnopora ( S o k o l o v

1955, 1962).

3. Em phasiz ing the presence o f squamulae in the peripheral part in a large number o f branching favositids в. V. P r e o b r a z h e n s k i y (1967) established the genus Squameopora and pointed out its affinities to the genus Squameofavosites a n d Emmonsia. Regrettably, a s D. H i l l remarked, the presence o f squamulae in Favosites hidensis (chosen as type species o f Squameopora by в. V. P r e o b r a z h e n s k i y ) is not mentioned in the publi­cation o f T. H a m a d a (1959). So diagnostic characters o f Squameopora are still missing. In any case, favositid corals with cylindrical outgrowings and thickened walls in the peripheral parts o f corallites (with or without squamulae) can be regarded as the representatives o f a genus different from all other genera o f the family Favositidae.

4. O n the basis o f abundant tabulate specimens from R u k h i n ’ s

(1938) localities in the Lower Devonian o f the Kolyma River (Northeast Russia), V. N. D u b a t o l o v (1969) has proved the priority o f the valid name Striatoporella. According to the detailed description o f two species, including the type-species Striatoporella multiporifera in D u b a t o l o v ’ s

m onograph (1969: 80-82, PI. XLVI, Fig. 1-4, PI. XLVII, Fig. 1-6), essential m orphological traits o f this genus can be characterized as follows:

2 9 6 Klem ens O ekentorp and Tong-Dzuy Thanh

- N odulose with cylindrical outgrowings and branching corallum.- Corallite walls thicken towards the peripheral zone.- Irregular development o f septal spines in the axial zone and association o f spines and squamula in the peripheral zone.- Large mural pores, sometimes situated at the com er o f the corallites. These features were not pointed out in the description o f D u b a t o l o v ,

but one o f us (T.D.T.) had the opportunity to observe it in thin sections o f the type-species stored in the Institute o f Geology and Geophysics in Novosibirsk. It can also be seen in D u b a t o l o v ’ s illustrations (1969: PI. XLVI and XLVII).

5. Com posit ion and distribution o f Striatoporella. Representatives o f Striatoporella are widespread from the Upper Silurian to the Middle Devonian, particularly in the Lower Devonian. Apart from typical species described from Lower Devonian o f Northeast Russia, the following branching favositids should be assigned to Striatoporella:

- A great number o f described branching favositids, including the groups o f Favosites socialis S o k o l o v & T e s a k o v , F. polaris C h e k h o v i c h , F. hidensis ( K a m e i ) , F. kozlowskyi ( S o k o l o v ) a. o.- D e s c r i b e d s p e c i e s o f Squameopora, for e x a m p l e s. magnissima, s. vukhuci ( T o n g - D z u y T h a n h e t a l . 1 9 8 8 ) a . o .

Fig. 1. Striatoporella schouppei (P o t t h a s t & O e k e n t o r p , 1988). Emsian-Eifelian boundary; H am ar Laghdad/Tafilalt , SE M orocco.

1: Cross section o f a cylindrical colony with thin walls in axial part, but thickened ones at the periphery. N um erous spines and squamulae are developed.- Section G IM -M A 30—1; X1 .6 .2: Longitudinal section o f a cylindrical outgrowth with walls thickening towards the periphery. Pores and numerous spines are well developed, cross sections o f the latter often appearing as dark dots. - Section G IM -M A 88-1 ; X 1.6.3: Cross section o f the cylindrical outgrowth (Fig. 1.2) with walls thickening towards the periphery o f the colony. - Section G IM -M A 88 -1 ; X2 .3 .4: Longitudinal section showing part o f a wall (left) and one o f the rare squamulae. - Section G IM -M A 88 -1 ; X 20 .5: Longitudinal section with a pore and strong spines, directed upwards. - Section G IM -M A 89 -2 ; X20.6: Longitudinal section with wall pores and pores situated in the corner o f the wall (“Mesofavosites” -zppearance). - Section G IM -M A 94 -1 ; X 20 .7: Detail o f a cross section showing moderately thickened walls with distinct boundary to the corallite cavity. The dark middle line is still preserved. - Section G IM -M A 94 -1 ; X20 .8: Detail o f a recrystallized wall. The original boundary o f the wall is lost due to grain growth, producing a saw-like border with irregular indentations. The latter ones represent scalenohedrons due to crystal growth. - Section G IM -M A 88 -1 ; X 48 .

Remarks on the genera Striatoporella and “Riphaeolites” 2 9 7

- The genus Rudakites L e l e s h u s (1964) from Lower Devonian o f Central Asia might be also regarded as a younger synonym o f Striatoporella. Judging from L e l e s h u s ’ s description (1964) apart from the branching shape o f the corallum the distinguishing trait o f Rudakites is the poor development o f septa, which occur only in the periphery o f the corallum in the form o f obtuse spines. The latter are similarly observed in the type species o f Striatoporella ( D u b a t o l o v 1969: 80-81).

2 9 8 Klem ens O ekentorp and Tong-D zuy Thanh

- Mesofavosites schouppei described by P o t t h a s t & O e k e n t o r p (1988) from the Emsian-Eifelian boundary strata o f SE M orocco represents a very interesting species o f Striatoporella.

N o tes on Striatoporella schouppei

New observations in well preserved polyparies stored in the Forschungsstelle für Korallenpaläozoologie, University Münster (Germany), show that the main m orphological elements o f these corals correspond to the diagnosis o f Striatoporella. These are (1) branching and nodulose coralla with cylindrical outgrowings (Fig. 1.1-1.3), (2) thickened walls toward the surface o f the corallum (Fig. 1 .1-1.3; compare Fig. 1.7) and (3) rare squamulae (Fig. 1.4) associated with coarse spines, especially in the peripheral zones o f the corallum (Fig. 1.2, 1.5).

Fig. 2. Riphaeolites pseudoseptatus ( O e k e n t o r p 1975). Ferrones Crinoidean l imestone, Em sian ; Ensenada de Banugues, Asturias, Northern Spain.

1: Longitudinal section showing moderately thickened walls and zonality. - Section G IM -B 2 -3 2 2 .2 ; X2.2 .2: Longitudinal section through an encrusting colony. The corallites grew along the substrate before bending upwards. - Section G IM -B 2 -1 6 .1 ; X 1 .3 .3: In the basal part o f the section corallites are thin-walled but become extremely thick on top. - Section G IM -B 2 -3 2 4 .1 ; X2 .2 .4: Cross section with most corallites having extremely thickened walls. Partly, diagenetic carbonate fills the whole cavity. - Section G IM -B 2 -3 2 4 .2 ; X 2 .2 .5: Longitudinal section showing the zonality o f wall thickenings; the corallites o f the basal parts remain thin-walled. - Section G IM -B 2-327 .4 ; X2 .4 .6: Detail o f a longitudinal section showing the spiral tube o f a com m ensal worm, Helicosalpinx asturiana O e k e n t o r p 1969. - Section G IM -B 2 -3 2 2 .2 ; X 45 .7: Detail o f a cross section. The polygonal corallites are limited by thick walls o f knotty appearance. These “paratrabecula” are caused by diagenesis. The dots in the large corallites (lower right) are cross-cut-spines. - Section G IM -B2-16.3 ; X 45.8: Detail o f a longitudinal section. The walls in a zone o f thickening show calcitic fibers in fan-like arrangement. In the upper part the walls are thin and may represent the original thickness (compare with Fig. 4.5). - Section G IM -B 2 - 327.4; X 45.9: Enlarged cross section showing different wall structures and thicknesses. In the lower part , some o f the thin walls still show the dark middle line. In the upper part the walls are thickened by coarser and longer fibers arranged in fan-systems (comp. Fig. 4.4). The latter indicates a diagenetic origin, for which the lightened color o f the carbonate material is an additional evidence. - Section G IM -B 2 -3 2 4 .1 ; X27.

Remarks on the genera Striatoporella and “Riphaeolites* 2 9 9

Distinguishing features o f this species are: 1. The profusedevelopment o f septal spines (Fig. 1.1-1.2). In the axial zone, they are numerous and represented as long and acute elements (Fig. 1.5). In the peripheral zone, some squamulae (Fig. 1.4) and squamulae-like spines occur in association with obtuse spines. 2. In addition to the typical mural pores, many pores are situated at the edge o f the walls (Fig. 1.5, Fig. 1.6), in the corner o f corallites, e. g. in Mesofavosites.

3 0 2 Klem ens O ekentorp and Tong-D zuy Thanh

2. In cross section, the thin (e.g. Favosites-Mkz) walls normally show a clear dark middle line and two framing zones o f dark grey, fine-grained carbonate (Fig. 4.4). Carbonate fibers have rarely been observed. The border to the visceral chambers is usually pronounced. Sometimes the middle line has been recrystallized to form a brightened zone with indistinct borders. Wall thicknesses in the Favosites-like parts range between 0.14 to 0.18 mm.

3. In Cleistopora-like parts o f the colony the walls are partially thickened, ranging from 0.1 to 0.33 mm, and are normally very light and coarsely fibrous. The fibers are arranged in fans o f irregular shape and extension without any uniformity (Figs. 2.8, 2.9, 3 .1-5 , 4 .1-6). Within the fans a fine lamellation o f the fibers can be noticed. This lamellation is due to the diagenetic growth o f crystals. The same feature has been observed and described by O e k e n t o r p (1989) in Pleistocene scleracti- nians from the Sinai Peninsula, Egypt. The boundary to the visceral chambers generally variable but mostly saw-like in section, e. g. the indentations, are caused by the m ode o f growth o f the crystals resp. the fans (Figs. 3.3, 3.4, 4 .1-3 , 4 .5-6). These thickenings are developed in more or less regular zones and on the same level throughout the colony. Additionally, they alternate with darker zones, which are often thinner than the lighter ones (Fig. 3.2).

Fig. 3 . Riphaeolites pseudoseptatus ( O e k e n t o r p 1 9 7 5 ) . Ferrones Crinoidean l imestone, Em sian; Ensenada de Banugues, Asturias, Northern Spain.

1: Longitudinal section through a zone o f thickened wall. The fibers are arranged in fan-systems. - Section G IM -B 2 -3 2 2 .2 ; X 20 .2: Variable thickening o f the walls in a longitudinal section. Partly the walls are extremely swollen by diagenetic fan-systems. - Section G IM -B 2 -1 6 .1 ; X20.3: Enlarged wall with several fans o f coarse and elongated fibres in longitudinal section. The fans are the so called “ St irnzonen”, e.g. arches o f recrystallization fronts, proving the diagenetic nature o f these structures - (Fig. 4.2, but also Figs. 4.1, 4.3, 4.4, 4.5, 4.6). - Section G IM -B 2 -1 6 .5 ; X45.4: Enlarged longitudinally-cut wall showing a very clear fibrous fan, which has replaced the primary wall structure. Moreover, the fibers grew beyond the limitation o f the primary wall into the cavity o f the corallite, giving the wall a club-shaped appearance with an irregular border (Com pare Fig. 4.6). - Section G IM -B2-16.5 ; X 45.5: A club-shaped thickened part o f a wall built o f coarse and elongated fibres, becom es much thinner and again thickened. The thin part obviously represents the primary wall (Com pare Fig. 4.5) - Section G IM -B 2 -1 6 .1 ; X 45 .6: Recrystallized wall and tabula. The infilling o f the corallite cavity contains some coarse crystals o f dolom ite , which show that diagenesis o f the skeleton began very early, e. g. before the corals were em bedded into the sediments, under the influence o f meteoric water. - Section G IM -B 2 -1 6 .6 ; X 45 .

Remarks on the genera Striatoporella and “Riphaeolites” 303

4. Cross sections also show peculiarities o f the wall structure, even though details (such as the fan systems) are lacking. The thickened walls look partially nodulose, resulting in a so-called “paratrabecular” m icro­structure (T o n g - D z u y 1966, 1988 and Fig. 2.7) or in a “ coenenchyme”- like appearance. This resembles similar structures in specimens o f the Silurian genus Thecia, e. g. Thecia (Theda) expatiata ( L o n s d a l e 1839)

3 0 4 Klem ens O ekentorp and Tong-D zuy Thanh

Fig. 4. Riphaeolites pseudoseptatus ( O e k e n t o r p 1975). Ferrones Crinoidean limestone, Em sian ; Ensenada de Banugues, Asturias, Northern Spain. Drawings o f wall structures to demonstrate more clearly the different appearance o f m icro­structures and the variable thicknesses o f the walls. The thickness is caused by reorganization o f the original microstructure controlled by the polyps. The reorganisation results in grain growth, brightening o f the carbonate and fan-like crystal growth o f the “rebuilt”, or new aggregates. During these processes the fibers grew beyond the limitation o f the primary walls resulting in an irregular border and extreme thickening. The original wall microstructure and wall thickness is retained only locally, for instance in Fig. 4.5 (upper part) or Fig. 4.4

(lower part). - Figure 4 not to scale.

Remarks on the genera Striatoporella and “Riphaeolites” 305

d e s c r i b e d a n d f i g u r e d b y N i c h o l s o n , 1879 o r A m s d e n , 1949 ( s .

O e k e n t o r p 1970: p i . 1, f ig . 2 a n d 3; c o m p a r e f u r t h e r p i s . ) . A s i n Thecia, w h e r e t h e d i a g e n e t i c m i c r o s t r u c t u r e s h a v e b e e n m o d i f i e d b y t h e

e x i s t e n c e o f s e p t a , s i m i l a r c o n d i t i o n s , e . g . t h e d e v e l o p m e n t o f s p i n e s ,

c a u s e d a c o m p a r a b l e m i c r o s t r u c t u r e in Riphaeolites.

In cross sections, a granular microstructure can be observed depending on the direction o f fibers in the fan-systems in longitudinal cuts. The material appears dark grey in thin walls but lighter in the recry­stallized parts (Figs. 3.2, 3.5). This color change is a clear evidence for a diagenetic origin. The middle line is often obscure or partially brownish; but a dark middle line can be found, where thin walls happen to be preserved.

These features, especially the light and coarse fibrous fan-system s com bined with their irregularity and the irregular bordering to the visceral cham bers, are clearly the results o f post-m ortem diagenetic processes. The appearance o f m icrostructural details (shape o f the fans; the dim ensions o f fibers; the increased transparency o f the carbonate; indentation o f the borders, etc.) corresponds to observations in Pleistocene corals from the Sinai Pen insula/Egypt ( O e k e n t o r p 1989) and Permian rugose corals from Indonesia ( O e k e n t o r p 1980, 1984). M oreover - especially in Permian corals but also in those from other periods - the zonality o f thickening is conspicuous. It has to be con si­dered as a progressive phase-like “ recrystallisation” during early diage­nesis in shallow m arine environm ents, resulting in so-called “Stirn- fronten” (Schouppe & S tacul 1966; O e k e n t o r p 1980, 1984; Potthast 1992). Shallow m arine conditions and influence o f m eteoric water during fossilization is evidenced by the existence o f dolom ite crystals in the cavity in fillings as well as in cem ents (Fig. 3.6).

According to these observations and their interpretation, the thicke­nings are secondary, diagenetic features rather than skeletal properties o f taxonomic significance. Therefore, “Pachyfavosites” pseudoseptatus O e k e n t o r p , 1975 can no longer be considered as a representative o f the genus Pachyfavosites, but resembles the genus Riphaeolites, whose wall structure is the same as in the Spanish material.

A m ong the few known species o f Riphaeolites, the one nearest to the type species (R. sokolovi) is the Emsian form from Northern Spain described as Pachyfavosites pseudoseptatus (O ekentorp 1975: 68-72; PI. XII, Fig. 4; PI. XIII, Fig. 4; PI. XIV, Fig. 2-3). The similarity o f these two species is clearly expressed in the crustose, lamellar shape o f the corallum, the manner o f rapidly changing corallite shape and the wall structure. Their similarity is also expressed even in the coexistence with

2 0 * N. Jb . Geol. Palaont. Mh. 1996

3 0 6 Klem ens O ekentorp and Tong-D zuy Thanh

specific symbionts in the form o f com mensal worms. However, Riphae- olites pseudoseptatus stands out by its peculiar septal structure. In the adult stages o f corallites, disjointed, ridge-like septa are observed in addition to coarse, spine-like septa ( O e k e n t o r p ) 1975: Pl. XII, Fig. 4; PI. XIV, Fig. 1). By these distinguishing m orphological elements, R. pseudo­septatus differs from all known species o f Ripbaeolites. Moreover, the Spanish species differs from the type-species by the small size o f the corallites and the thin walls.

While a detailed description and illustration o f Ripbaeolites pseudosep­tatus has been provided by O e k e n t o r p ( 1 9 7 5 ) , this publication adds the illustration o f some important morphological elements that improve the diagnosis o f the species.

References

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D u b a t o l o v , V. N . & S p a s s k i y , N. Y. A. (1964): Nekotorye novye korally iz devona S ov ’ etskogo Soyusa. - [In:] Stratigraficheskiy i geograficheskiy obzor devonskikh korallov S S S R : 112-137, pi. 1-11 ; Moskva (Akademia Nauk SSSR ). [Some new corals from D evonian o f the Soviet Union]

F l ü g e l , E. (1978): Mikrofazielle U ntersuchungsm ethoden von Kalken. - 454 pp., 68 figs., 57 tbls. , 33 pis . ; Berlin-Heidelberg-New York (Springer).

H a m a d a , T. (1959): O n the taxonom ic posit ion o f Favosites hidensis and its Devonian age. - Ja p . J . Geol. G eophys., 30: 2 01-213 , pi. 16.

H i l l , D. (1981): Tabulata. - [In:] T e i c h e r t , C . (Ed.) : Treatise on Invertebrate Paleontology. F. Coelenterata, Suppl. 1.2: 430-762 ; B o u lder/C ol . - Lawrence/Kans.

L e l e s h u s , V. L . (1964): Novye devonskiy rod Rudakites (Tabulata) iz Tadzhiki- stana. - Paleont. Zhurnal, 1964 (4): 4 5 -4 8 ; Moskva (Akademia N auk SSSR). [New Devonian genus Rudakites (Tabulata) from Tadzhikistan]

O e k e n t o r p , K l . (1969): Kom m ensalism us bei Favositiden. - Münster. Forsch. Geol. Paläont., 12: 165-216, 10 Abb., Taf. 13-16; Münster.

- (1970): Revision des Genus Thecia E d w a r d s & H a i m e 1849 (Coelenterata, Tabulata). - N . Jb . Geol. Paläont. Abh., 137 (1): 103-173, 12 A bb .; Stuttgart.

- (1975): Beschreibung und Systematik devonischer Favositidae Asturiens und Beobachtungen zur Biogeographie nordspanischer Korallenfaunen. - Münster. Forsch. Geol. Paläont., 37: 1-129, 10 Abb., 1 Tab., 14 Taf.; Münster.

Remarks on the genera Striatoporella and “Riphaeolites“ 3 0 7

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T o n g - D z u y T h a n h ; N g u e n D u c K h o a ; K h r o m y k h , V . G.; N g u y e n H u u H u n g ;

N g u y e n T h e D z a n & T a H o a P h u o n g ( 1 9 8 8 ) : S t r a t i g r a f i a i t s e l e n t e r a t y

D e v o n a V i e t n a m a , 2 T s e l e n t e r a t y . - 2 4 8 p p . , 6 4 p l . [ D e v o n i a n s t r a t i g r a p h y

a n d c o e l e n t e r a t a o f Vietnam, 2 Coelenterata]

3 0 8 K. O ekentorp and Tong-D zuy Thanh, Remarks

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Revidiertes Manuskript bei der Tübinger Schriftleitung eingegangen am19. Dezem ber 1995.

A n s c h r i f t e n d e r V e r f a s s e r :

K l e m e n s O e k e n t o r p , Forschungsstelle für Korallenpaläozoologie , Geol.-Paläont.Institut und M useum , Westfälische Wilhelms-Universität, Pferdegasse 3,D - 4 8 1 4 3 Münster/Westf.; T o n g - D z u y T h a n h , Department o f Geology, Vietnam

National University Hanoi, 90 Nguen Trai Road, Dong-Da. Hanoi, Vietnam.