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Processing Food and Processing Food and NutritionNutrition
Chapter 46Chapter 46
Learning Objective 1Learning Objective 1
• Describe food processing, including Describe food processing, including ingestioningestion, , digestiondigestion, , absorptionabsorption, and , and egestionegestion or or eliminationelimination
• Compare the digestive system of a Compare the digestive system of a cnidarian (such as cnidarian (such as HydraHydra) with that of an ) with that of an earthworm or vertebrateearthworm or vertebrate
NutritionNutrition
• The process of taking in and using foodThe process of taking in and using food
Food ProcessingFood Processing
• FeedingFeeding • selection, acquisition, and selection, acquisition, and ingestioningestion of food of food
• DigestionDigestion • breaking down food mechanically, chemically breaking down food mechanically, chemically
• Absorption Absorption • nutrients pass from digestive tract into bloodnutrients pass from digestive tract into blood
• EgestionEgestion (elimination)(elimination)• undigested, unabsorbed food discharged undigested, unabsorbed food discharged
Cnidarians and FlatwormsCnidarians and Flatworms
• Food digested in Food digested in gastrovascular cavitygastrovascular cavity • only one opening only one opening • serves as both mouth and anusserves as both mouth and anus
Gastrovascular Gastrovascular CavitiesCavities
Fig. 46-2a, p. 992
Wastes
Food
Tentacle
Mouth
Food Enzymes secreted by inner layer
Gastro- vascular cavity Food
absorbed
Gastrodermis(a) Hydra
(b) Exhalation.
Fig. 46-2b, p. 992
Gastrovascular cavityEpidermis
Food particles
Pharynx
Food absorbed
Mouth
EnzymesWastes
Lining of the intestine Food
(b) Flatworm
Complex Invertebrates and Complex Invertebrates and VertebratesVertebrates
• Digestive tract is complete tubeDigestive tract is complete tube• with opening at each end with opening at each end
• As food passes through tube As food passes through tube • digestion takes placedigestion takes place
• Parts of digestive tract are specializedParts of digestive tract are specialized• to perform specific functionsto perform specific functions
Complete Digestive TractComplete Digestive Tract
Fig. 46-3, p. 992
Crop Gizzard
EsophagusIntestine
Pharynx
Mouth Anus
Wastes
Food
Explore various digestive Explore various digestive systems by clicking on the figure systems by clicking on the figure
in ThomsonNOW.in ThomsonNOW.
KEY CONCEPTSKEY CONCEPTS
• Food processing includes ingestion, Food processing includes ingestion, digestion, absorption, and eliminationdigestion, absorption, and elimination
Adaptations for Obtaining FoodAdaptations for Obtaining Food
KEY CONCEPTSKEY CONCEPTS
• Many animal adaptations are associated Many animal adaptations are associated with mode of nutritionwith mode of nutrition
Learning Objective 2Learning Objective 2
• Trace the pathway traveled by an ingested Trace the pathway traveled by an ingested meal in the human digestive systemmeal in the human digestive system
• Describe the structure and function of Describe the structure and function of each organ involvedeach organ involved
Human Digestive System 1Human Digestive System 1
• Mouth Mouth • Mechanical, enzymatic digestion of carbohydratesMechanical, enzymatic digestion of carbohydrates
• Mammalian teethMammalian teeth• incisorsincisors for biting for biting • caninescanines for tearing food for tearing food • premolars,premolars, molarsmolars for crushing and grinding for crushing and grinding
Teeth and DietTeeth and Diet
Fig. 46-6a, p. 994
Incisors
CaninesPremolars Molars
(a) Carnivore.
Fig. 46-6b, p. 994
Incisors
Canine
Premolars
Molars
(b) Herbivore.
Fig. 46-6c, p. 994
Canines
Incisors
Premolars
Molars
(c) Omnivore.
Tooth StructureTooth Structure
Fig. 46-7, p. 995
Enamel GumCrown
Pulp cavity
PulpNeck Dentin
CementumRoot canal
RootSpongy bone
Nerve
Vein
(a) Human lower molar. Sagittal section showing the crown, neck, and root.
Artery (b) X-ray of a healthy tooth.
Human Digestive System 2Human Digestive System 2
• Three pairs of Three pairs of salivary glandssalivary glands • secrete saliva (enzyme secrete saliva (enzyme salivary amylasesalivary amylase
digests starch)digests starch)
• PharynxPharynx and and esophagusesophagus• carry food to stomach carry food to stomach
Human Digestive System 3Human Digestive System 3
• PeristalsisPeristalsis• waves of muscular contractionwaves of muscular contraction• pushes pushes bolusbolus of food along digestive tract of food along digestive tract
• StomachStomach • mechanical digestion by vigorous churningmechanical digestion by vigorous churning• enzyme enzyme pepsinpepsin in gastric juice digests in gastric juice digests
proteinsproteins
PeristalsisPeristalsis
Fig. 46-8, p. 995
EsophagusRelaxed muscle layer
Circular muscles contract, constricting passageway and pushing bolus ahead
Longitudinal muscles contract, shortening passageway ahead of the bolus
Relaxed muscle layer
Food bolus
Sphincter closed
Sphincter open
Stomach StomachA bolus is moved through the esophagus by peristaltic contractions.
When the sphincter (ring of muscle) at the entrance of the stomach opens, food enters the stomach.
Human Digestive System 4Human Digestive System 4
• RugaeRugae • folds in stomach wallfolds in stomach wall• expand as stomach fills with food expand as stomach fills with food
• Gastric glandsGastric glands secrete secrete • hydrochloric acidhydrochloric acid • pepsinogen (pepsinogen (precursor of precursor of pepsin)pepsin)
Stomach StructureStomach Structure
Fig. 46-9a, p. 996
Visceral peritoneum
Esophagus
Sphincter
Circular muscle layer
Longitudinal muscle layer
Oblique muscle layer
Pyloric sphincter
Duodenum Rugae
Fig. 46-9b (1), p. 996
Openings into gastric glands
EpitheliumGastric glands
Lymph nodule
Gastric mucosa
Fig. 46-9b (2), p. 996
Chief cell Parietal cell
Nuclei
Surface epithelium
Chief cells
Parietal cellsGastric glands
Gastric glands
Human Digestive System 5Human Digestive System 5
• ChymeChyme • soup of partly digested foodsoup of partly digested food• leaves stomach through leaves stomach through pyloruspylorus• enters enters small intestinesmall intestine in spurts in spurts
• DuodenumDuodenum • location of most enzymatic digestionlocation of most enzymatic digestion• produces several digestive enzymesproduces several digestive enzymes• receives secretions from liver and pancreasreceives secretions from liver and pancreas
Structure of Digestive TractStructure of Digestive Tract
Fig. 46-5, p. 993
VISCERAL PERITONEUM
Lymph nodule
Villi
Blood vessels
SUBMUCOSA
Nerve fibersMUCOSA
MUSCLE LAYER Inner circular fibers Outer longitudinal fibers
Human Digestive System 6Human Digestive System 6
• LiverLiver produces produces bilebile• which emulsifies fats which emulsifies fats
• PancreasPancreas releases enzymes releases enzymes• digest protein, lipid, carbohydrate, RNA, DNA digest protein, lipid, carbohydrate, RNA, DNA
• TrypsinTrypsin and and chymotrypsinchymotrypsin • digest polypeptides to dipeptides digest polypeptides to dipeptides
Human Digestive System 7Human Digestive System 7
• Pancreatic lipasePancreatic lipase • degrades fatsdegrades fats
• Pancreatic amylasePancreatic amylase • digests complex carbohydratesdigests complex carbohydrates
Liver and PancreasLiver and Pancreas
Fig. 46-11, p. 998
StomachInferior vena cava
Right lobe of liver
Right hepatic duct
PancreasCommon bile duct Hepatic
portal vein
Pancreatic duct
GallbladderDuodenum
Human Digestive System 8Human Digestive System 8
• Large intestineLarge intestine• cecum, colon, rectum, anuscecum, colon, rectum, anus • eliminates undigested wastes eliminates undigested wastes • incubates bacteria (produce vitamin K, certain incubates bacteria (produce vitamin K, certain
B vitamins)B vitamins)
Human Digestive SystemHuman Digestive System
Fig. 46-4, p. 993
Parotid salivary gland
PharynxSublingual salivary gland
EsophagusSubmandibular salivary gland
Liver Stomach
DuodenumGallbladder
Pancreas
Ascending colon Transverse colon
Jejunumlleum Descending colon
CecumSigmoid colon
Vermiform appendix Anus
Rectum
Insert “Human digestive Insert “Human digestive system”system”
human_digestive_m.swfhuman_digestive_m.swf
Learn more about human Learn more about human digestion by clicking on the digestion by clicking on the
figures in ThomsonNOW.figures in ThomsonNOW.
KEY CONCEPTSKEY CONCEPTS
• Various parts of the vertebrate digestive Various parts of the vertebrate digestive system are specialized to perform specific system are specialized to perform specific functions; accessory glands (liver, functions; accessory glands (liver, pancreas, and salivary glands) secrete pancreas, and salivary glands) secrete fluids and enzymes important in digestionfluids and enzymes important in digestion
Learning Objective 3Learning Objective 3
• Describe the step-by-step digestion of Describe the step-by-step digestion of carbohydratecarbohydrate, , proteinprotein, and , and lipidlipid
Carbohydrate DigestionCarbohydrate Digestion
• Polysaccharides Polysaccharides • digested to disaccharide digested to disaccharide maltosemaltose by by salivarysalivary
and and pancreatic amylasespancreatic amylases
• MaltaseMaltase in small intestine in small intestine• splits splits maltosemaltose into into glucoseglucose (main product of (main product of
carbohydrate digestion)carbohydrate digestion)
Protein DigestionProtein Digestion
• Proteins are split Proteins are split • by by pepsinpepsin in stomach in stomach• by proteolytic enzymes in pancreatic juiceby proteolytic enzymes in pancreatic juice
• DipeptidasesDipeptidases • split small peptides into split small peptides into amino acidsamino acids
Lipid DigestionLipid Digestion
• Lipids are emulsified by Lipids are emulsified by bile saltsbile salts • then hydrolyzed by then hydrolyzed by pancreatic lipasepancreatic lipase
Learning Objective 4Learning Objective 4
• What structural adaptations increase the What structural adaptations increase the surface area of the digestive tract?surface area of the digestive tract?
Surface Area of the Small IntestineSurface Area of the Small Intestine
• Greatly expanded by Greatly expanded by • folds in wall (folds in wall (rugaerugae) ) • intestinal intestinal villivilli (projections of mucosa) (projections of mucosa)• microvillimicrovilli (plasma membrane projections of (plasma membrane projections of
epithelial cells of villi)epithelial cells of villi)
Villi and MicrovilliVilli and Microvilli
Fig. 46-10a, p. 997
Fig. 46-10b, p. 997
VilliEpithelial cells lining villus
Capillary network
Nerve fiber
Mucosa Openings of intestinal glands
Goblet cells
Intestinal glands
Submucosa Lacteal
Lymph vessel
Muscle layer
Visceral peritoneum
(b)
Fig. 46-10c, p. 997
Learning Objective 5Learning Objective 5
• Compare lipid absorption with absorption Compare lipid absorption with absorption of other nutrientsof other nutrients
Nutrient AbsorptionNutrient Absorption
• Nutrients are absorbed through thin walls Nutrients are absorbed through thin walls of intestinal villi of intestinal villi
• Hepatic portal veinHepatic portal vein transports transports amino acidsamino acids and and glucoseglucose to liver to liver
Lipid Absorption 1Lipid Absorption 1
• Fatty acids and monoacylglycerolsFatty acids and monoacylglycerols• enter epithelial cells in intestinal liningenter epithelial cells in intestinal lining• are reassembled into triacylglycerols are reassembled into triacylglycerols • then packaged into then packaged into chylomicron chylomicron dropletsdroplets
Lipid Absorption 2Lipid Absorption 2
• ChylomicronsChylomicrons • also contain cholesterol and phospholipidsalso contain cholesterol and phospholipids• are covered by a protein coat are covered by a protein coat
• Lymphatic systemLymphatic system• transports chylomicrons to blood circulationtransports chylomicrons to blood circulation
Learning Objective 6Learning Objective 6
• Summarize the nutritional requirements for Summarize the nutritional requirements for dietary carbohydrates, lipids, and proteinsdietary carbohydrates, lipids, and proteins
• Trace the fate of glucose, lipids, and Trace the fate of glucose, lipids, and amino acids after their absorptionamino acids after their absorption
A Balanced DietA Balanced Diet
• Humans and other animals requireHumans and other animals require• carbohydrates carbohydrates • lipidslipids• proteins proteins • vitaminsvitamins• minerals minerals
Carbohydrates 1Carbohydrates 1
• Most are ingested as polysaccharidesMost are ingested as polysaccharides• starch and cellulosestarch and cellulose
• Polysaccharides are Polysaccharides are complex carbohydratescomplex carbohydrates
• FiberFiber • mixture of cellulose, other indigestible mixture of cellulose, other indigestible
carbohydratescarbohydrates
Carbohydrates 2Carbohydrates 2
• CarbohydratesCarbohydrates• used mainly as energy source used mainly as energy source
• Glucose concentration in bloodGlucose concentration in blood• is carefully regulated is carefully regulated
• Excess glucoseExcess glucose• is stored as glycogenis stored as glycogen• or converted to fator converted to fat
Lipids 1Lipids 1
• Used to Used to • provide energy provide energy • form components of cell membranes form components of cell membranes • synthesize steroid hormones, other lipid synthesize steroid hormones, other lipid
substances substances
• Most lipids are ingested as triacylglycerols Most lipids are ingested as triacylglycerols
Lipids 2Lipids 2
• Fatty acids are converted to molecules of Fatty acids are converted to molecules of acetyl coenzyme A acetyl coenzyme A
• which enter citric acid cycle which enter citric acid cycle
• Excess fatty acids Excess fatty acids • are converted to triacylglycerolare converted to triacylglycerol• stored as fatstored as fat
Lipids 3Lipids 3
• Lipids are transported as large molecular Lipids are transported as large molecular complexes complexes (lipoproteins)(lipoproteins)
• Low-density lipoproteins (LDLs)Low-density lipoproteins (LDLs)• deliver cholesterol to cells deliver cholesterol to cells
• High-density lipoproteins (HDLs)High-density lipoproteins (HDLs)• collect excess cholesterolcollect excess cholesterol• transport it to livertransport it to liver
Fat Fat MetabolismMetabolism
Fig. 46-13, p. 1002
Fat cell Fat
Fatty acids +
Glycerol
Transported in blood
Liver Fatty acid + Glycerol
Enters cellular
respirationUsed to
make triacylglycerols
Acetyl CoA G3P
GlucoseConverted to other lipidsKetone bodies
Other cellsPyruvate or
acetyl CoA
Converted to other lipids
Enters cellular
respiration
Enters cellular
respiration
Fig. 46-13, p. 1002
Fat cell Fat
Fatty acids +
Glycerol
Transported in blood
+ Glycerol
Enters cellular respirationG3P
Glucose
Liver Fatty acid
Used to make
triacylglycerols
Acetyl CoA
Converted to other lipidsKetone bodies
Enters cellular respiration
Other cellsPyruvate or
acetyl CoA
Converted to other lipids
Enters cellular respiration
Stepped Art
Proteins 1Proteins 1
• Proteins serve as enzymesProteins serve as enzymes• and essential structural components of cells and essential structural components of cells
• Best distribution of Best distribution of essential amino acidsessential amino acids• in complete proteins of animal foods in complete proteins of animal foods
• Excess amino acidsExcess amino acids• are deaminated by liver cells are deaminated by liver cells
Proteins 2Proteins 2
• Amino groups are converted to ureaAmino groups are converted to urea• excreted in urineexcreted in urine
• Remaining keto acids are Remaining keto acids are • converted to carbohydrate and used as fuel converted to carbohydrate and used as fuel • converted to lipid and stored in fat cellsconverted to lipid and stored in fat cells
Protein Protein MetabolismMetabolism
Fig. 46-14, p. 1003
AMINO ACIDS
Catabolism Anabolism
Excess amino acids Structural proteins, hemoglobin, myosin, actin, enzymes, plasma proteins
Deamination
NH3 (ammonia) + -keto acids
FatUrea Pyruvate, -
ketoglutarate
Acetyl CoA
Ketone bodies
To kidneys Acetyl CoA
Fatty acids + glycerol
Triacylglycerol
Storage in fat cells
Liver
Enters cellular
respiration
Learning Objective 7Learning Objective 7
• Describe the nutritional functions of Describe the nutritional functions of vitaminsvitamins, , mineralsminerals, and , and phytochemicalsphytochemicals
VitaminsVitamins
• Organic compounds required in small Organic compounds required in small amounts for biochemical processes amounts for biochemical processes
• components of coenzymes components of coenzymes
• Fat-soluble vitaminsFat-soluble vitamins• vitamins A, D, E, Kvitamins A, D, E, K
• Water-soluble vitaminsWater-soluble vitamins • B and C vitaminsB and C vitamins
MineralsMinerals
• Inorganic nutrients ingested as saltsInorganic nutrients ingested as salts• dissolved in food and water dissolved in food and water
• Trace elementsTrace elements • minerals required in small amounts minerals required in small amounts • (< 100 mg/day)(< 100 mg/day)
PhytochemicalsPhytochemicals
• Plant compounds that promote health Plant compounds that promote health • many are many are antioxidantsantioxidants that destroy that destroy oxidantsoxidants
• OxidantsOxidants • free radicals,free radicals, other reactive molecules other reactive molecules• damage DNA, proteins, unsaturated fatty damage DNA, proteins, unsaturated fatty
acids by snatching electronsacids by snatching electrons
KEY CONCEPTSKEY CONCEPTS
• Most animals require the same basic Most animals require the same basic nutrients: carbohydrates, lipids, proteins, nutrients: carbohydrates, lipids, proteins, vitamins, and mineralsvitamins, and minerals
Learning Objective 8Learning Objective 8
• Contrast Contrast basal metabolic ratebasal metabolic rate with with total total metabolic ratemetabolic rate
• Write the basic energy equation for Write the basic energy equation for maintaining body weight, and describe the maintaining body weight, and describe the consequences of altering it in either consequences of altering it in either directiondirection
Measuring MetabolismMeasuring Metabolism
• Basal metabolic rate (BMR)Basal metabolic rate (BMR) • body’s cost of metabolic livingbody’s cost of metabolic living
• Total metabolic rateTotal metabolic rate • BMR plus energy used to carry on daily BMR plus energy used to carry on daily
activitiesactivities
EnergyEnergy
• If energy (kcal) input equals energy output If energy (kcal) input equals energy output • body weight remains constant body weight remains constant
• If energy output exceeds energy input If energy output exceeds energy input • body weight decreases body weight decreases
• If energy input exceeds energy outputIf energy input exceeds energy output• body weight increasesbody weight increases
Learning Objective 9Learning Objective 9
• In general terms, describe the effects of In general terms, describe the effects of malnutritionmalnutrition, including both , including both undernutritionundernutrition and and overnutritionovernutrition
UndernutritionUndernutrition
• Form of malnutrition Form of malnutrition • causes fatigue causes fatigue • depresses immune function depresses immune function
• Essential amino acidsEssential amino acids• nutrients most often deficient in dietnutrients most often deficient in diet
Protein DeficiencyProtein Deficiency
OvernutritionOvernutrition
• ObesityObesity • serious form of malnutrition serious form of malnutrition • excess fat accumulates in adipose tissuesexcess fat accumulates in adipose tissues• factor in heart disease, diabetes mellitus, factor in heart disease, diabetes mellitus,
other disorders other disorders
• Person gains weight taking in more energy Person gains weight taking in more energy (kilocalories) than expended in activity(kilocalories) than expended in activity
KEY CONCEPTSKEY CONCEPTS
• Basal metabolic rate is the body’s cost of Basal metabolic rate is the body’s cost of metabolic living metabolic living
• When energy (kilocalories) input equals When energy (kilocalories) input equals energy output, body weight remains energy output, body weight remains constantconstant
Learning Objective 10Learning Objective 10
• Summarize current hypotheses about the Summarize current hypotheses about the regulation of food intake and energy regulation of food intake and energy homeostasis, including the roles of homeostasis, including the roles of leptinleptin and and neuropeptide Yneuropeptide Y
LeptinLeptin
• Hormone produced by fat cellsHormone produced by fat cells• in proportion to body fat in proportion to body fat
• Signals brain about status of energy stores Signals brain about status of energy stores
Effect of LeptinEffect of Leptin
Neuropeptide Y (NPY)Neuropeptide Y (NPY)
• Neurotransmitter produced in hypothalamusNeurotransmitter produced in hypothalamus• increases appetite increases appetite
• Slows metabolism when leptin levels and Slows metabolism when leptin levels and food intake are lowfood intake are low
Insert “Chronology of Insert “Chronology of leptin research”leptin research”
leptin_research.swfleptin_research.swf
Learn more about leptins and Learn more about leptins and body weight by clicking on the body weight by clicking on the
figure in ThomsonNOW.figure in ThomsonNOW.