of ~engthl weight for Relationships for Research Report 88 ... 43.pdf · Lengthmeight Relationships for 88 Species of Fish Encountered in the North East ... (Ammodytes maximus Raitt

Department of Agriculture and Fisheries for Scotland

~ength l weight Relationships for 88 Species of Fish Encountered in the North East Atlantic

K A Coull A S Jermyn A W Newton G I Henderson and W B Hall

Scottish Fisheries Research Report Number 43 1989 ISSN 0308 8022

Lengthmeight Relationships for 88 Species of Fish Encountered in the North East Atlantic

by

K A Coull, A S Jermyn, A W Newton, G I Henderson and W B Hall

Marine Laboratory, Aberdeen

Introduction As a necessary adjunct to monitoring fish stocks the Marine Laboratory has obtained a large volume of lengthlweight data for al l species of major commercial importance to Scottish fishermen. In addition a limited amount of information has also been obtained on a variety of minor species. This paper summarises the data currently available.

Material Material was collected from the middle and northern North Sea, the west coast of Scotland, Rockall and, to a lesser extent, Faeroese and Icelandic waters.

Information has been obtained from four separate sources:

a. Samples were collected during research vessel surveys. Prior to 1981 fish were deep frozen a t sea, transported back to the Laboratory and measured and weighed after defrosting. Since 1981 fish have been weighed a t sea using a Mettler PC8000 electronic balance linked to a micro-computer (Newton, 1982).

b. Samples were obtained while monitoring discards aboard commercial fishing vessels (Jermyn and Hall, 1978). These samples were kept on ice until they could be processed in the Laboratory, usual1 y three or four days after capture.

c. Rare fish records dating back to 1900 were used to provide information on species not encountered on a regular basis. It is not possible to estimate how long after death these specimens were weighed or, for the earlier part of the century, how they were preserved prior to weighing. Later specimens were preserved on ice.

d. The bulk of the data for the most common commercial species were obtained from samples purchased a t local markets. These fish had been caught 1 - 10 days before weighing, and had been kept on ice until processed.

Methods The lengths of most species were determined by measuring the distance from the tip of the snout to the extreme tip of the caudal fin. However, for skates and rays the term length refers to the distance from wing tip to wing tip. Prior to 1967 measurements were recorded to the nearest centimetre, but from January 1967 measurements have been made to the centimetre below. In order to standardise measurements 0.5 cm was deducted from the length of fish measured prior to 1967.

The total ungutted weight of each fish was recorded in grammes and for many species the gutted weight was also obtained.

Programs provided by the Laboratory's Computing and Statistics Section were used in data preparation and analysis. Where length (L), total weight (MI) and gutted weight (GW) were recorded, relationships and plots have been produced for lengthltotal weight, lengthlgutted weight and total weightlgutted weight. In the case of species for which there were no gutted weight data, only a lengthltotal weight relationship was calculated. For relationship between length and weight (total or gutted) a function of the form:

was fitted to the data, where W = weight (g) and L = length (cm).

For a relationship between total weight and gutted weight a function of the form:

was fitted to the data, where TW = total weight and GW = gutted weight.

Results The weight a t length of many species is known to vary seasonally, and wherever possible lengthlweight relationships have been prepared for each calendar month. A desirable secondary division is into the relevant sampling areas (see Appendix I, Figs 1-4) so as to make allowance for any regional variation in growth rates. Monthlylarea relationships are available only for herring (Clupea harengus (L.) 1, mackerel (Scomber scombrus (L.)), Raitt 's sandeel (Ammodytes maximus Raitt 1 934) and sprat (Sprattus sprattus (L.)) (Tables I, I I and I I I). For the main demersal species, viz cod (Gadus morhua ( L.) I,, haddock (Melanogrammus aeglefinus ( L.) 1, whiting (Merlangius merlangus ( L.) 1, sai t he (Pollachius virens ( L.) ) , lemon sole (Microstomus kik (Walbaum)) and plaice (Pleuronectes platessa (L.)), data from all areas were combined to give a monthly relationship (Table IV). Quarterly relationships are provided for spurdogs (Sgualus acanthias (L.)) (Table V) whilst for porbeagles (Lamna nasus (Bonnaterre)) an annual relationship for each sex is provided (Table VI). For all other species the available data were sufficient to provide only an annual relationship.

Plots for species where relationships are available for lengthltotal weight, lengthlgutted weight and total weightlgutted weight are shown in Appendix I I, Figures 5 to 43. Plots for species for which there are only sufficient data for a lengthltotal weight relationship are shown in Appendix I I - Figures 44 to 81. All the plots are accompanied by a table indicating the source area and number of observations. For the majority of species the source area is based on demersal market sampling areas (Fig. 1 and Appendix I) which, in turn, are associated with biological andlor geographical boundaries.

An index of species, cross-referenced by common and scientific names, is provided in Appendix 1 1 1.

Table I Herring (Clupea harengus) - total weightllength relationship

Annual value for 'b', monthly and annual mean values for 'a' are given below

Area North Rona Hebrides NW Ireland North and all North and South M inch

Sea Minch

Area Code 2,3,4,5, 1 7,9 8 6,11,12

'b' Value 3.0904 2.9776 3.0083 3.0256

'a' Values

Jan Feb Mar A P ~ May Jun Jul AwJ S ~ P oct Nov Dec

Mean Annual Values 0.00603 0.00863 0.00791 0.00742

Irish Sea and

Clyde

See Figure 4 for herring sampling areas and Appendix I for area codes.

Table II Total weightllength relationships

Annual values for 'b', monthly values for 'a' and annual mean values for 'a' are given below for mackerel (Scomber scombrus) and Raitt's sandeel (Ammodytes marinus) by sampling area

Species Mackerel Raitt's sandeel

Areas Clyde Others Shetland West Coast

Area Code 11 - -

'b' Value 3.21 3.29 3.32 3.32

'a' Values

Jan Feb Mar A P ~ May Jun Jul Aug S ~ P Oct Nov Dec

Mean Annual Values 0.0038 10 0.003001 0.001243 0.001317

See Figure 2 for mackerel sampling areas and Appendix I for area codes.

Table Ill Sprat (Sprattus spratms) - total weightllength relationship

TW = a ~ ~

Annual values for 'b', monthly and annual mean values for 'a' are given below

Area North Sea areas West Scotland Clyde

Area Code 1,2,3,4 5 6

'b' Value 3.4746 3.4746 3.4788

'a' Values

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Mean Annual Values "'

1

See Figure 3 for sprat bhlpling areas and Appendix I for area codes.

Table IV I, ;, Gutted weighthernth nlationlript

G W = ~ L ~ ( T W = G W x R F I

Annual values for 'b',mnt###atueD for 'a' and annual mean values for 'a' are given below each species. Raising factors (RF) for converting gutted weight t o total weight are also given

Cod Haddock

'b' Value 2.8571' 28268

'a' Values

Jan Feb Mar A P ~ May Jun Jul Aug S ~ P Oct Nov Dec

Mean Annual Values

Whiting

2.9456

0.0100 0.0094 0.0090 0 .0088 0.0088 0.0089 0.0090 0.0092 0.0095 0.0096 0.0097 0.0097

0.0093

1.13

Saithe

2.7374

0.0241 0.0222 0.0219 0.0227 0.0229 0.0238 0.0236 0.0243 0.025 1 0.0252 0.0255 0.0246

0.0238

1.19

Lemon Plaice Sole

Table V

Weightnength relationship, spurdog - Squalus scanthias (from Shanks, 1981 1

Females: Quarters 1 ,2 ,4 TW(g) = 0.00108 Quarter 3 TW(g1 = 0.00595 ~(cm)2 .8~9

Males: Annual TW(g) = 0.00576 ~ ( c m ) ~ . ~ ~ ~

Table VI

Weighttlength relationship, porbeagle - Lamna nasus (from Shanks, 1988)

Male lW(kg) = O.OOl'922 L ~ . ~ ~ Male GW(kg) = 0.0001 14 ~ 2 . ~ ~ ~

Female TW(kg) = 0.000315 L ~ - ~ ~ ~ Female GW(kg) = 0.000047 L ~ . ~ ~ ~

Acknowledgemcmnts Our thanks are due to many members, both past and present, of the Laboratory's staff for their efforts in collecting the samples on which this paper is based. Our thanks are also due to D.C. Emslie for his advice on analysis and presentation.

References Jsrmyn, AS. and Hall, W.B. 1978. Sampling procedures for estimating haddock and whiting discards in the North Sea by Scottish fishing vessels in 1976 and 1977. ICES CM 1978/0:9.

Newton, A.W. 1982. Weighing at sea. DAFS Marine Laboratory Internal Report.

Shanks, A.M. 1981. Further analysis of total weightllength data for spur dogfish. DAFS Marine Laboratory lnternal Report.

Shanks, A.M. 1988. Whole weightllength and gutted weightllength relationships for porbeagles. DAFS Marine Laboratory Internal Report.

Appendix I. Sampling Areas and Codes (Figs 14)

North Sea

01 Shetland 02 Viking 03 Moray Firth 04 Buchan 05 Forties 06 Central 07 Danish Coast 08 Humber 09 West Orkney 10 German Bight 11 Thames 12 IJmuiden 13 Utsire

Mackerel

01 Northwest North Sea (N) 02 Northeast North Sea (N) 03 Northwest North Sea (S) 04 Northeast North Sea (S) 05 West Central 06 East Central 07 Outer Hebrides 08 Cape Butt 09 Northwest lreland 10 Minch 11 Clyde

DEMERSAL SPECIES

West Coast of Scotland North Coast

Solan 09 West Orkney Rising Ground 40 Solan Butt of Lewis Inner Hebrides Outer Hebrides South Minch Clyde Northern lreland lrish Sea

PELAGIC SPECIES

Herring

01 Hebrides 02 North Rona 03 West Shetland 04 East Shetland 05 Moray Firth 06 Northeast North Sea 07 Northwest lreland 08 North Minch 09 South Minch 10 Clyde 11 South Buchan 12 Central North Sea 13 lrish Sea

Sprat

01 Shetland 02 Moray Firth 03 East Scotland ,04 Northeast England 05 West Scotland 06 Clyde 07 Northern lreland 08 Northern North Sea 09 Central North Sea 10 Northwest Scotland

Figure 1. Demersal Market Sampling Areas

Figure 2. Mackerel Sampling Areas

Figure 3. Sprat Sampling Areas

Figure 4. Herring Sampling Areas

1 O W 8W 6 W 4W 2W 0 2E 4E 6E

Appendix I I . Figures 5-81 showing relationships between length, total weight, and gutted weight for various species

Figure 5 . Cuckoo Ray - Raja naevus Mul,ler & Henle

Length Ccm)

0 500 1000 1500 2000

Gutted Weight Cg)

8 16 24 32 40

Length C c m l

Areas collected : Shetland

Buchan

Butt of Lewis

Data source : Discard trips

Date collected : 1981 - 1984 How measured : cm below Cwingspsn)

Annual Relationship

L/TW L/GW GW/TW

obs used 173 173 173

min L 8 8 - max L 30 30 - min TW 19 - 19

max TW 1974 - 1974

min GW - 17 17

max GW 1832 1832

Figure 6 . Blue Skate - RaJa batis Linnaeus

20 30 40

Length Ccml Length Ccml

Are- collected : Inner Hebrides

Morw Fir th

South Hi nch

Data swrce : Diecard tripe

Date collected : 1982 - 1985 cm below How measured :

obs used

min L

max L

min TW

max TW

min GW

max GW

Relationship

L/GW GW/TW

e e ie - 49 - - 88

- 1686

79 79

1713 1713

Gutted Weight Cg)

Figure 7. Spotted Ray - Raja montagui Fowler

Length Ccml Length (cml

0 500 1000 1500 2000

Gutted Weight (gl

Areas col lected : Shetland

North Coast

West Coast

of Scotland

Data source : Research vessels

Discard t r i p s

Date collected : 1977 - 1985 How measured : cm below

Annua l

L/TW

obs used 87

min L 12

max L 42

rnin TW 27

max TW 1890

rnin GW - rnax GW

Relationship

L/GW GW/TW

87 87

12 - 42 - - 27

- 1890

25 25

1794 1794

Figure 8. Starry Ray - Raja radiata Donovan

8 16 24 32 40

Length Ccrnl

9 15 21 27 33

Length (cml

Areas collected : North Sea

Data source : Discard t r i p s

Data collected : 1977 - 1984 HOW measured : cm below

obs used

min L

max L

rnin TW

max TW

rnin GW

max GW

Annual Relationship

L/TW L/GW GW/TW

376 85 85

9 9 -

33 31 -

27 - 27

1160 - 745

- 24 24

- 766 766

Gutted Weight (91

Figure 9. Thornback Ray - Raja clavata Linnaeus

24.5 27.0 29.5


How measured :


Moray Fir th

Buchan

South Min*,,)i % L -..

Data source : Discard ripe

Date collected : 1981 - 1 84

Annua l

L/TW

obs used 12

min L 22

max L 31

min TW 197

max TW 572

min GW - max GW

Gutted Weight Cgl

Figure 10. St ing Ray - Dasyatis pastinaca Linnaeus

20 24 28 32 23.5 25.0 26.5 28.0 29.5

Length Ccml Length tcml

Areas collected : Moray Firth

Buchan

Butt of Lewis

Data source : Rare fish records

Date col lected : 1960 - 1965

9.' How measured :

/ Annua l

L/TW

obs used 5

min L 16.5

max L 28.5

min TW 131

mex TW 790

300 I I I I min GW - 300 400 500 600 700 max GW

cm below Cwingspanl

Relationship

L/GW GW/TW

4 4

23.5 - 28.5 - - 350

- 790

319 319

696 696

Gutted Weight Cgl

Figure 11. Eagle Ray - Myl iobatis aqui la Linnaeus


Areas collected : Buchan

Data source : Rare f i sh records

Date collected : 1957 - 1970

How measured : cm below

Annual Relationship

L/TW L/GW GW/TW .'

obs used 12 3 3

min L 27.5 27.5 - max L 58.5 51.5 - min TW 612 - 612 7

max TW 3690 - 2054

min GW - 554 554

max GW

Gutted Weight Cgl

I' F igure 12. Sea Lamprey - Petromyzon mar i nus L innaeus I

40 60 80

Length Ccm)

0 300 600 900 1200

Gutted Weight Cgl

45 60 75

Length Ccm)


Clyde

Faroe



Annua I

L/TW

obs used 25

min L 31.5

max L 93

min TW 41

max TW 1310

min GW - mex GW

Relationship

L/GW GW/TW

13 13

31.5 - 84.5 - - 41

- 1169

34 34

1113 1113

Figure 13. Three-bearded Rockl ing - Gaidropsarus v u l g a r i s Cloquet

Length Ccml

32 36 40

Length Ccml


Buchan

West Coast of

Scot l and


Research vessels


Annua l Re l at i onsh i p

L/TW L/GW GW/TW

obs used 17 I1 11

min L 29.5 29.5 - max L 47 43.5 - min TW 230 - 230

max TW 972 - 764

min GW - 210 210

max GW 726 726

Gutted Weight Cgl

Figure 14. Tadpole F i s h - Raniceps raninus (Linnaeusl

Length Ccml

TW = GW * 1.0845

o ! I I I I

15 20 25 30 35

Length Ccml


West Coast

of Scotland


Date collected : 1954 - 1982 How measured : crn below

Annual Relationship

L/TW L/GW GW/TW

obs used 21 10 10

min L 13.5 15.5 - max L 30.5 30.5 - min TW 24 - 47

max TW 465 - 465

min GW - 39 39

rnax GW 430 430

Gutted Weight Cgl

Figure 15. Skipper - Scornberesox saurus CWalbauml

Length Ccml

804 I I I I

100 130 160 190 220

Gutted Weight Cgl

Length Ccm)


West Coast

of Sco

Data source : Rare f

Date collected : 1932 - How measured :

Annua l

L/TW

obs used 22

min L 17

max L 40.5

min TW 13

max TW 233

min GW - rnax GW

t l and

ish records

1983

Relationship

L/GW GW/TW

13 13

30.5 - 40.5 -

1 F igure 16. Red Sea Bream - Pagel lus bogaraveo CBrunnichl

35 40 45

Length Ccml

400 800 1200 1600 2000

Gutted Weight Cgl

30 35 40 45 50

Length Ccml


Buchan

Outer Hebrides



Annua l

L/TW

obs used 6

min L 31.5

max L 47

min TW 520

max TW 2050

min GW - max GW

Relationship

L/GW GW/TW

6 6

31.5 - 47 - - 520

- 2050

490 490

1787 1787

Figure 17. Trigger fish - Balistes carolinensis (Gmelinl

25 30 35


Areas collected : Butt of Lewia

Inner Hebrides

Data source : Rare f ish records

Date collected : 1984


obs used

min L

mex L

min TW

max TW

min GW

max GW

Annual Relationship

Gutted Weight Cgl

Figure 18. Tub Gurnard - T r i g l a lucerna Linnaeus

3200

2400 a, V

4 s a - 1600 3: - (0 4

eoo

0 30 42 54 66

Length C c m l


Buchan

Inner Hebrides

Dete source : Discard t r ips



Annual Relationship

L/TW L/GW GW/TW

obs used 23 6 6

min L 17 23 - max L 62.5 62.5 - min TW 40 - 146

max TW 2944 - 2944

min GW - 0 600 n

128 1200 1800

128

max GW 2368 2368

Gutted Weiaht (a)

Figure 19. Black Fish - Centrolophus niger (Gmelinl

Length (cml Length Ccml


West Coast

of Scotland

Faroe

Ice l and


Date collected : 1931 - 1984 How measured I cm below

Annue I

L/TW

obs used 33

min L 39

max L 65.5

min TW 409

max TW 2462

min GW - max GW

Relationship

L/GW GW/TW

19 19

39 - 65.5 - - 409 - 2380

368 368

2110 2110

Figure 20. Bass - Dicentrarchus labrax (Linnaeusl

36 44 52



Clyde

Inner Hebrides


Date collected : 1955 - 1977 How measured

obs used

min L

max L

min TW

max TW

min GW

max GW

Annua l

L/TW

32

29.5

58

275

2050

-

Relationship

L/GW GW/TW

21 21

29.5. - 54.5 - - 275

- 2041

259 259

1906 1906

Gutted Weight Cgl

Figure 21. Ling - Molva molva (Linnaeus)

30 42 54

Length Ccm)

32 40 48 56 64

Length Ccm)


Data source

Date collected :

How measured

Annua l

L/TW

obs used 19

min L 20

max L 60

min TW 33

max TW 1358

min GW - max GW

Research vessels

Discard trips

1978 - 1985 cm be l ow

Relationship

L/GW GW/TW

16 16

33 - 60 - - 125

- 1358

117 117

1194 1194

Gutted Weight Cgl

Figure 22. Dover Sole - Solea solea (Linnaeusl

3

15 20 25

Length Ccml

- 0 60 120 180 240

Gutted Weight Cgl

Length Ccml t

Areas collected : Moray Firth

German Bight

Clyde


Discard trips

Date collected :

How measured

Annua l

L/TW

obs used 13

min L 11

max L 29

min TW 9

max TW 243

min GW - max GW

Relationship

L/GW GW/TW

13 13

11 - 29 - - 9

- 243

8 8

238 238

Figure 23. Gar f i sh - Belone belone (Linnaeusl

Length Ccm) Length Ccml

Areas col lected : North Sea

West Coast

of Scotland



How measured

Annua l

L/TW

obs used 20

min L 30.5

max L 76.5

min TW 24

max TW 609

min GW - max GW

Relationship

L/GW GW/TW

9 9

30.5 - 71 - - 24 - 529

22 22

494 494

Gutted Weight Cgl

Figure 24. Greater Fork-beard - Phycis blennoides ( B r ~ n i c h l

Length Ccml

30 42 54 66

Length Ccml


Swth Minch

Clyde


Discard trips

Date collected : 1926 - 1982 HOW measured

obs used

min L

max L

min TW

max TW

0 500 1000 1500 2000 m-in GW

max GW

.Gutted Weight Cgl

cm be l ow

Relationship

Figure 25. Hake - Merluccius merluccius CLinnaeusl

Length Ccml

Gutted Weight Cgl

15 30 45 60 75

Length Ccml


West Coast

of Scotland

Data source I Discard t r ipe

Date collected : 1977 - 1992 HOW measured :

obs used

min L

mex L

min TW

max TW

min GW

msx GW

Annual

L/TW

134

16

66

25

1773 -

cm below

Relationship

Figure 26. Greater Weever - Trachinus draco Linnaeus



Moray F i r t h

Buchan


Discard t r i p s


Annua l

L/TW

obs used 29

min L 24.5

max L 35

min TW 73

max TW 344

min GW - max GW

Relationship

L/GW GW/TW

14 14

25 -

31.5 - - 90 - 244

84 84

231 231

Gutted Weight Cg)

Figure 27. Grey Mullet - Mugi l cephalus Linnaeus

-

20 30 40 50

Length Ccml

- 24 32 40 48 56

Length Ccml

Areas collected : Moray F i r t h

Buchan

West Coast

of Scot I and




Annual Relationship - L/TW L/GW GW/TW

obs used 32 8 8

min L 26.5 26.5 - max L 56.5 51 , - min TW 220 - 220

mex TW 2314 - 1603

Gutted Weight Cgl

Figure 28. Angler - Lophius piscatorius Linnaeus

10 20 30 40 50

Length Ccml

0 250 500 750 1000

Gutted Weight Cgl

10 20 30 40

Length Ccml


West Coast

of Scotland


Discard t r i p s



Annua l

L/TW

obs used 371

min L 14

max L 44

min TW 37

max TW 1257

min GW - max GW

Relationship

L/GW GW/TW

371 371

14 - 44 - - 37 - 1257

32 32

967 967

Figure 29. Black Sea Bream - Spondyl iosoma cantharusCLinnaeus1

Length Ccm)

Gutted Weight Cg3

Length Ccml


West Coast

of Scotland

Data source : Rare f i s h records


H o w measured : cm below

Annual Relationship

L/TW L/GW GW/TW

obs used 43 16 16

min L 18.5 23.5 - max L 41.5 41.5 - min TW 106 - 209

msx TW 1268 - 1268

m i n GW AW+'-%, b , 198 ' *

m x ow 4 wd ' ' ' 1174

Figure 3 0 . Catfish - Anarhichas lupus L-innaeus

45 60 35 90

Length Ccm)

Gutted Weight Cgl

Length Ccm)


Moray F i r th

k h a n


Discard t r ips


obs used

min L

max L

min TW

max TW

min GW

max GW

cm be l ow

Relationship

Figure 31. B r i I I - Scophthalmus rhombus CLinnaeus)

0 0 25 30 35 40 45



Clyde

Data emume I Resesrch vessels

Diacsrd t r ips



AmKlsI

L/TW

obs used 7

min L 25

msx L 43

min TW 242

max TW 1562

min GW - msx GW '

Relationship

L/GW GW/TW

4 4

25 - 43 - - 242

- 1562

228 228

1529 1529

Gutted Weight Cgl

Figure 32. Blue-Mouth - He l icolenus dactylopterusCDelaroche3

21 27 33 39


Areas col lected : Moray F i r t h

Buchan

For t i es

Data source : Rare f i s h records

Date col lected : 1956 - 1968 How measured : cm below

Annua l Re l a t i onsh i p

L/TW L/GW GW/TW

obs used 6 6 6

rnin L 15.5 15.5 - max L 38.5 38.5 - m i n TW 68 - 68

mex TW 1044 - 1044

m i n GW - 63 63

max GW 954 954

Gutted Weight Cgl

Figure 33. Bib - Trisopterus luscus CLinnaeus3

Length C c m l Length Ccml



Discard t r ips


Annua l

L/TW

obs used 147

min L 18

max L 42

min TW 61

rnax TW 1106

min GW - max GW

Relationship

L/GW GW/TW

86 86

21 -

42 -

- 109

- 1106

101 101

915 915

Gutted Weight Cgl

!

F igure 34. All i s Shad - Alosa alosa CLinnaeusl



West Coast

of Scotland



Annua l

L/TW

obs used 42

min L 31.5

max L 64.5

min TW 260

max TW 2340

min GW - max GW

Relationship

L/GW GW/TW

18 18

34.5 - 57.5 - - 317

- 2261

296 296

2003 2003

Gutted Weight Cgl

Figure 35. Cuckoo Wrasse - Labrus mixtus Linnaeus

16 24 32 40 48 20 24 28 32 36



West Coast

TW = GW * 1.0653 of Scotland



How measured

Annua l

L/TW

obs used 80

min L 18

max L 43

min TW 65

max TW 1318 I I I 1 I min GW - 60 180 300 420 540

max GW

cm be l ow

Relationship

L/GW GW/TW

9 9

20.5 - 32.5 - - 107

- 522

102 102

506 506

Gutted Weight Cgl

Figure 36. Witch - Glyptocephalus cynoglossus CLinnaeusl

Length Ccml

Gutted Weight Cgl

15 21 27 33 39

Length Ccml


West Coast

of Scotland

Data source I Discard t r ipe

Date col lected : 1977 - 1984


Annual Relationship

obs used

min L

max L

min TW

max TW

mln GW

mex GW

Figure 37. Red Mul le t - MU1 1 . ~ 6 swrnuletus Linnabs

24 32 40

Length Ccml

Gutted Weight Cgl

Length Ccm)


North Coast

West Coast

of Scotland




Annual Relationship

L/TW L/GW GW/TW

obs used 49 32 32

min L 20.5 20.5 - . max L 46.5 30.5 - min TW 112 - . 112

max TW 1272 - 940

min GW - 100 100

max GW 866 ' 866

' L

Figure 38. Topknot - Zeugopterus punctatus (Bloch)

12 16 20



Moray Firth

Buchen

So l an

Clyde


Date collected : 1958 - 1984 How measured I cm below

Annual Relationship

L/TW L/GW GW/TW

obs used 28 8 8

min L 11.5 15.5 - max L 23.5 23.5 - min TW 28 - 90

max TW 308 - 282 60 ! 1 I 1 1

min GW - 120 180 240 300 86 86 60

max GW 275 275

Gutted Weight Cgl

Figure 39. Turbot - Scophthalmus maximus (Linnaeusl

Length Ccrnl

48 56 64 72 80

Length Ccrnl



TW = GW * 1.0643 Oate collected : 1901 - 1985 loo00 -


n 8000 - Annual Relationship 0) u L/TW L/GW GW/TW

4 s obs used 4 4 4 m .- 6000 - min L 50 50 - al 3 mex L 75 75 - -

m i n TW 2457 - 2457

mex TW 9720 - 9720

min GW - 2317 2317

max GW 9184 9184

2000 4000 6000 8000 loo00

Gutted Weight Cgl

Figure 40 . Twaite Shad - Alosa f a l l a x (Lacepedel I

33 39 45



West Cmst

TW = GW * 1.1362 of Scotland 0 . Data source I Rare f ish records


00 How measured : cm below

Annual Relationship

L/TW L/GW GW/TW

obs used 80 58 58

min L 27.5 27.5 - max L 50.5 50.5 - min TW 167 - 168

max TW 1335 - 1335

min GW - 156 156 150

0 300 600 900 1200 max GW 1165 1165

Gutted Weight Cg)

Figure 41. Red Band-fish - Cepois rubescens Linnaeus

40 50 60

Length Ccml

30 40 . 50 60 70

Length Ccml

Areas collected : South Minch

Clyde

I r i s h Sea

Date source : Rare f i sh records


How measured :

Annua I

L/TW

obs used 12

min L 33.5

mex L 60

min TW 34

mex TW 142

min GW - max GW

Relationship

L/GW GW/TW

6 6

33.5 - 68 - - 42

- 142

38 38

136 136

Gutted Weight Cg)

Figure 4 2 . Common Dab - Limanda l imanda CLinnaeusl

Length Ccm) Length Ccm)


Moray F i r th

Buchen

Swth Minch

Dsts source : Discard t r ips


How measured r cm below

Annual Relationship

L/TW L/GW GW/TW

obs used 575 575 575

min L 9 9 - msx L 37 37 - min TW 7 - 7

mex TW 608 - 608

min GW - 7 7

max GW 567 567

Gutted Weight Cgl

Figure 43. Bat tan Wrasse - Labus bergytte Ascentijs

Length Ccm) Length Ccm)


West Coast

of Scotland

Data source r Rare f i sh records


1800 - a, u

4 Annual Relationship

L 0 L/TW L/GW GW/TW . 1200 - 3

obs ueed 67 43 43

- min L 16.5 16.5 - (0 4 max L 47.5 47.5 - f 600 - min TW - 82 163

max TW 2106 4, 2106

min GW - 150 5 a 150 0 8

I 1 I 1

0 500 1000 1500 2000 max GW 19S5 1995

Gutted Weight Cgl

- -igure 44. Rabbitfish - Chimaera monstrosa Linnaeus

Length Ccml

Areas collected: Shetland West Orkney Rocka l I Faroe

Data source : Rare fish records Date collected : 1958 - 1965 How measured : To the cm below

Annual Relationship

obs used 6 min L 80.5 max L 97.5 min TW 1435 max TW 1860

Figure 45. Horse Mackerel - Trachurus trachurus (Linnaeusl

16 24 32 40 48

Length Ccml

Areas col lected: North Sea West Coast of Scotland

Data source : Research vessels Discard t r i p s

Date col lected : 1977 - 1984 How measured : To the cm below

Annual Relationship

obs used 283 min L 16 max L 41 min TW 26 max TW 724

Figure 46. Angel fish - Squatina squatina CLinnaeusI

60 80 100 120 140

Length Ccml

Areas collected: North Sea West Coast of Scotland

Data source : Rare f i sh records Date collected : 1956 - 1984 How measured : To the cm below

Annual Relationship


Figure 47. Bogue - Boops boops CLinnaeusl

1.6484

23.4 24 .O 24.6 25.2 25.8

Length Ccm3

Areas collected: Buchan Humber South Minch


Annual Relationship


Figure 48. Bull Rout - Myoxocephalus scorpius CLinnaeusl

12 18 ?4 30 36

Length Ccml

Areas col lected: Moray F i r t h Buchan Danish Coast

Data source : Research vessels Discard t r i p s


Annual Relationship

obs used 311 m i n L 12 max L 35 min TW 32 max TW 751

72 84 96 108 120

Length Cmm)

Areas cot lected: Moray Firth Buchan

Data source : Research vessels' Date collected : 1984 - 1985 How measured : To the mm below

Annual Relationship

obs used 8 min L 7 msx L 11 min TW 4.8 max TW 21.1

-igure 50. Sunfish - Mola mola CLinnaeusl

Areas collected: Danish Coast Humber North Coast


Annual Relationship

obs used 13 m i n L 46 max L 213 min TW 4.90 max TW 762.00

8 .

Figure 51. Ray's Bream - Brama brama CBonnaterreI

32 40 48 56 64

Length Ccml

Areas col lected: North Sea South Minch


Annual Relationship


Figure 52. Lumpsucker - Cyclopterus lumpus Linnaeus

Length Ccml

Areas collected: Buchan Centra l

Data source : Research vessels Discard trips

Date collected : 1982 - 1985 How measured : To the cm below

Annual Relationship


Figure 53. Meagre - Argyrosomosus regium (Ass03

60 85 110 135 160

Length Ccml

Areas col lected: Buchan Data source : Rare fish records Date collected : 1939 - 1962 How measured : To the cm below

Annual Relationship

obs used 4 min L 62.5 rr ,

max L 152.5 min TW 2154 max TW 26989

Figure 54. Norway Haddock - Sebastes viviparus Kroyer

Areas collected: Shetland So l an



Annual Relationship


Figure 55. Electric Ray - Torpedo nobi l iana Bonaparte

20 40 60 80 100

Length Ccml

Areas collected: North Sea West Coast of Scotland


Annual Relationship

obs used 8 min L 20 max L 83.5 min TW 277 max TW 25855

Figure 5 6 . Great Pipefish - Syngnathus acus Linnaeus

3.5270

25.0 27.5 30 .O 32.5 35.0

Length Ccml

Areas collected: Moray Firth Data source : Research vessels Date collected : 1985 How measured : To the cm below

Annual Relationship

obs used 4 min L 25 max L 33 min TW 5.1 max TW 13.4

Figure 57. Hook-nose - Agonus cataphractus CLinnaeusl

90 110 130 150 170

Length Cmml

Areas collected: Moray Firth Buchan

Data source : Discard trips Date collected : 1977 - 1985 How measured : To the mm below

Annual Relationship

obs used 82 min L 9 max L 16 min TW 5.6 max TW 29.6

Figure 58. Wreckfish - Po lypr ion arnericanus CBloch & Schneider:

42 54 66 78 90

Length .Ccml

Areas collected: West Coast of Scotland Rocka I I


Annual Relationship

obs used 6 min L 46.5 max L 87 m i n TW 1870 max TW 10433

Figure 59. Sturgeon

40 120 200 280 360

Length Ccml

Areas co l lec ted: North Sea West Coast o f Scotland

Data source : Rare f i s h records Date co l lec ted : 1931 - 1984 How meas. : To the cm below

Annual Relat ionship

obs used 72 min L 75.5 max L 352.5 min TW 4.80 max TW 317.50

Figure 60. Opah - * Lsmpris gutkatus CBrunnichl

Areas collected: Shetland Buchen West Coast of Scotland

Data source : Rere f i sh recdrds Date cotlected : 1948 - 1983 How measured : To the cm below

Annual Relationship

obs usea 13 min L 77 mex L 136.5 min TW 20900 max TW 63500

Length Ccm)

Figure 61. ,Deal f i sh - Trachipterus arcticus (Brunnich)

04 6 I

105 135 165 195 225

Length (cml

Areas cot lected: North Sea West Coast of Scotland Rocka l I Faroe

Data source : Rare f i s h records Date col lected : 1963 - 1983 How measured : To the cm below

Annua I , Re I a t i onsh i p

obs used 18 min L 111:5 max L 218 , ' ". . m i n TW 1500 max TW M O O 0

-

Figure 62. Snake Blenny - Lumpenus lampretaeforrni:~ ~ W ~ l b e w l

L

Areas collected: Buchen South Minch

Date source : Discard t r ips Date collected 1 1982 - 1984 How measured : To the cm below

Annual Relationship

obs used 11

min L 16 mex L 27 min TW 6 mex TW 19

Length Ccml

Figure 63. Bu t te r f i sh - Phol is ~ u n n e l l u s (Linnaeusl

Areas collected: Buchen Data swrce : Discard t r ips Date collected : 1984 How measured : To the mm below

Annual Relationship

obs used 59

min L 15 max L 25 min TW 14.5 max TW 73.9

Length Cmml

I Figure 64. . ,

Long Rough Dab - H i ppog l osso i des p l atesso i des CFabr i c i us)

Areas collected: Shetland Viking Moray Firth Buchan



Annual Relationship

obs used 534 min L 10 mex L , 36 min TW 7 max TW 444

8 16 24 32 40

Length Ccml

Figure 65. Flounder - Platichthus flesus CLinnaeus)

Areas collected: Moray Firth Buchan

Data source : Discard trips Date collected : 1979 - 1985 How measured : To the cm below

Annual Relationship


8 16 24 32 40

Length Ccml

Figure 66. John Dory - Zeus faber Linnaeus

2.9343

Areas cot lected: North Coast Northern Ireland

Data swrce : Rere f i sh records Discard t r i p s

Date collected : 1977 - 1385 How meesured : To the cm below

Amual Relationship

obs used 56 min L 13.5 mex L 61 min TW 50 mex TW 3778

10 25 40 55 70

Length C c m l

Figure 67. Boar-fish - Capros aper CLinnaeusl

Areas collected: Shetland Bu t t of Lewis

Data swrce : Rare f i sh records Date collected : 1962 - 1970 How measured : To the cm below

Annual Relationship


Length Ccm)

Figure 68. Lesser Argentine - Argentina sphyraena Linnaeus - '' - -1 -

3.0534

120 TW = 0.0053CLl

Areas collected: North Sea Inner Hebrides Outer Hebrides

Data ewrce : Research vessels 95 Date collected : 1963 - 1984

How measured : To the cm below

Annual Relationship 70

obs used 157 min L 15.5 max L 25.5 min TW 21

45 max TW 110

20 15 18 21 24 27

Length Ccml

Figure 69. Greater Argentine - Argentina s i l us Ascanius

3.2027

Areas collected: Rockall Ice l and

Data source : Research vessels Date collected : 1362 - 1963 How measured : To the cm below

Annual Relationship


Length Ccml

Figure 70. Poor Cod - Trisopterus minutus CLinnaeusl

Areas collected: North Sea Inner Hebrides Clyde

Date swrce : Research vessels 150 -

0 Discard trips u Date collected : 1971 - 1984 c, How measured : To the cm below s a, .;; 100 - Annual Relationship

3 - obs used 452 do min L 9 4 max L 26

50 - min TW 6 max TW 172

0 1 I

7.5 12.5 17.5 22.5 27.5

Length Ccm3

Figure 71. Four-bearded Rockling - Rhinonemus cimbrius (Linnaeusl

Areas collected: North Sea Date swrce : Research vessels

0 Discard trips

P Date collected : 1982 - 1985 How measured : To the cm below

I AMwel Relationship

obs uaed 63 min L 11 max L 29 min TW 5 max TW 134

Length C c m l

Figure 72. Grey Gurnard - Eutrigla gurnardus CLinnaeusl

3.1003

800 TW = 0.0062CLl Areas col lected: North Sea

West Coast of Scotland

600 Data source : Research vessels

Discard t r i p s Date col lected : 1976 - 1984 How measured : To the cm below

400 Annual Relat ionship

obs used 747 m i n L 7

200 max L 44 m i n TW 3 max TW ' 776

0 5 15 25 35 45

Length Ccml

Figure 73. Red Gurnard - Aspitrigla cuculus CLinnaeusl

Areas col lected: West Coast of Scotland


900 D i scard tr i ps

a, u

Date co l lec ted : 1977 - 1982 How measured : To the cm below

c, L a, Annual Relat ionship

600 3: obs used 480 - min L m

12

c, max L 46 c" 300 min TW 14

max TW 1062

0 10 20 30 40 50

Length Ccml

Figure 74. Esmark's Eelpout - Lycodes esmarki i Col l e t

Areas col lected: Iceland Data source : Rare f i s h records Date col lected : 1958 - 1965 How measured : To the cm below

Annual Relationship


Length Ccml

Figure 75. Viviparous Blenny - Zoarces viviparus (Linnaeusl

Areas col lected: Buchan Data source : Discard t r i p Date col lected : 1982 How measured : To the cm below

Annual Relationship


19 21 23 25 27

Length (cml

Figure 76. Dragonet - Cal l ionymus lyra Linnaeus

Length Ccm1

Areas collected: Moray F i r t h Buchan South Minch

Data swrce : Research vessels Discard t r i p s


Annual Relationship

obs used 417 min L 8 max L 28 min TW 4 max TW ' 132

Figure 77. Spotted Dragonet - Cal l ionymus maculatus Rafinesque-Schmaltz

Areas collected: Morsy F i r t h Buchan

Date swrce : Research veesele Discard t r ips

Date col lected : 1977 - 1983 How measured To the cm below

Annual Relationship

obs used 66 m i n L 10 max L 16 min TW 6 max TW 20

Length Ccml

Figure 78. Boni to - Sarda sarda

3.1000 TW = O.OO51CLl

CB loch)

Areas collected: Buchen I r i s h Sea

Data source : Rare f i s h records Date col lected : 1938 - 1969 How meesured : To the cm below

Amual Relationship

obs used 5 m i n L 60.5 m x L 76.5

. inin TW 2.400 hsxl TW 5.695

Length Ccml

Figure 79. B l u e f i n Tunny - Thunnus thynnus (Linnaeus)

Areas collected: Shetland I Morey F i r t h

South Minch

240 - Data source : Rare f i s h records Date col lected : 1951 - 1959 How measured I To the cm below

Annual Relationship 160 -

obs used 5 m i n L 58.5 max L 263.5

80 - min TW 3.402 mex TW 254.010

0 I

30 90 150 210 270

Length Ccml

Figure 80. Solenette - Buglossidium luteum (Rissol

Areas col lected: Danish Coast Data swrce : Research vessels Date col lected : 1985 How measured : To the mm below

Annual Relationship

obs used 3 inin L 7 max L 8 m i n TW 4.14 max TW 4.75

Length Cmml

Figure 81. Thickback Sole - Microchirus variegatus (Donovan1

Areas col lected: Moray F i r t h Buchan

Data source : Research vessels Date col lected : 1985 How measured : To the mm below

Annual Relationship

obs used 5 min L 13 max L 18 m i n TW 23.2 max TW 64.5

Length Cmml

Appendix I I I. Alph&mtical i*x of spmim

Table Figure

Acipenser sturio Agonus cataphrttctus Allis shad Alosa alosa Alosa fallax Amrnodytes marinus Anarhichas lupus Angel ray Angler Argyrosomosus regium Argentina silus Argentina sph yraena Aspitrigla cuculus

Balistes carolinensis Ballan wrasse Bass Belone belone Bib Black fish Black sea bream Bluefin tunny Bluemouth Blue skate Boar fish Bogue Bonito Boops boops Brama brama Brill Buglossidium luteum Bull rout Butterfish

Callionymus lyra Callionymus maculatus Capms aper Catfish Cen trolophus niger Cepola rubescens Chimaera monstrosa Clupea harengus Cod Common dab Cuckoo ray Cuckoo wrasse Cyclopterus lumpus

Dasyatis pastinaca Deal fish Dicentrarchus labrax Dover sole Dragonet

Table Figure

Eagle ray Electric ray Esmark's eelpout Eutrigla gurnardus

Flounder Four-bearded rockling

Gadus morhua Gaidropsarus vulgaris Garfish Glyptocephalus cynoglossus Greater argentine Greater fork-beard Greater weever Great pipefish Grey gurnard Grey mullet

Haddock Hake Helicolenus dactylopterus Herring Hippoglossoides platessoides Hooknose Horse mackerel

John Dory

Labrus bergylta Labrus mixtus Lamna nasus Lampris guttatus Lemon sole Lesser argentine Limanda limanda Ling Long rough dab Lophius pisca torius L umpenus lampretae formis Lumpsucker L ycodes esmarkii

Mackerel Meagre Melanogrammus aegle finus Merlangius merlangus Merluccius merluccius Microchirus variegatus Microstomus kit t Mola mola Molva molva Mugil cephalus Mullus surmuletus Mylioba tis aquila Myoxocephalus scorpius

Table Figure

Norway haddock Norwegian topknot

Opah

Pageflus bogaraveo Petromyzon marinus Pholis gunnellus Phrynorhombus norvegicus Phycis blennoides Plaice Platichth ys flews Pleuronectes platessa Pollachius virens Polyprion americanus Poor cod Probeagle

Rabbitfish Raitt's sandeel Raja batis Raja clavata Raja montagui Raja naevus Raja radiata Raniceps raninus Ray's bream R e d band-fish Red gurnard Red mullet Red sea bream Rhinonemus cimbrius

Saithe Sarda sarda Scomber scombrus Scomberesox saurus Scophthalmus maximus Scophthalmus rhombus Sea lamprey Sebastes viviparus Skipper Snake blenny Solea solea Solenette Spondyliosoma cantharus Spotted dragonet Spotted ray Sprat 1 1 1 Sprattus sprattus 111 Spurdog V Squalus acanthias V Sguatina sguatina Starry ray Sting ray Sturgeon Sunfish Syngnathus acus

Tadpole fish Thickback sole Thornback ray Three-bearded rockiing Thunnus thynnus Topknot Torpedo nobiliana Trachinus draco Trachipterus arcticus Trachurus trachurus Trigger-f ish Trigla lucerna Trisopterus luscus Trisopterus minutus Tub gurnard Turbot Twaite shad

Viviparous blenny

Whiting Witch Wreckfish

Zeugopterus punctatus Zeus faber Zoarces viviparus

Table Figure

Printed by HMSO Reprographic Unit, Edinburgh 7/89 750 (025212)

Documents

of ~engthl weight for Relationships for Research Report 88 ... 43.pdf · Lengthmeight Relationships for 88 Species of Fish Encountered in the North East ... (Ammodytes maximus Raitt