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Northwest and Alaska F i s h e r i e s Center Processed Report;\
RECREATIONALLY IMPORTANT MARINE FISHES OF PUGET SOUND, WASHINGTON
Percy M. Washington
U.S. DEPARTMENT OF COMMERCE National Oceanic and Atmospheric Administrat ion
Nat ional Marine F i s h e r i e s Serv ice Northwest and Alaska F i s h e r i e s Center
Resource Ecology and Fishery Management Division
9: This r e p o r t does not c o n s t i t u t e a pub l i ca t ion and i s f o r information only. A l l d a t a h e r e i n a r e t o be considered provis ional .
ABSTRACT
Thirty-four species of non-salmonid fishes were determined to be of
recreational importance in Puget Sound, Washington. A review of the litera-
ture provided such information as local names, physical features used in
their recognition, world and Washington state distribution, habitat, utiliza-
tion, and size. Additional information on Puget Sound distribution, eating
qualities, and baits and lures were gathered during NMFS-Marine Recreational
Fisheries surveys (1973-1976) and was added to provide a 'handy recreational
marine fishery resource guide.
CONTENTS
Page No . INTRODUCTION . . . . . . . . . . . . . . . . . . . . . . . . 1
Codfishes and hakes - Gadidae . . . . . . . . . . . . . . . 5
Pacific eod . . . . . . . . . . . . . . . . . . . . . . . 7
Pacific hake . . . . . . . . . . . . . . . . . . . . . . . 9
Tomcod . . . . . . . . . . . . . . . . . . . . . . . . . 11
Walleye pollock . . . . . . . . . . . . . . . . . . . . . 13
Surfperches - Embiotocidae . . . . . . . . . . . . . . . . . 15
Shiner perch . . . . . . . . . . . . . . . . . . . . . . . 16
Striped seaper, ch . . . . . . . . . . . . . . . . . . . . . 18
Pile perch . . . . . . . . . . . . . . . . . . . . . . . . 20
. . . . . . . . . . . . . . . . . Rockfishes - Scorpaenidae 22
Brownrockfish . . . . . . . . . . . . . . . . . . . . . . 24
Copper rockfish . . . . . . . . . . . . . . . . . . . . . 26
Greenstriped rockfish . . . . . . . . . . . . . . . . . . 28
Yellowtail rockfish . . . . . . . . . . . . . . . . . . . 30
Quillback rockfish . . . . . . . . . . . . . . . . . . . . 32
. . . . . . . . . . . . . . . . . . . . . . Black rockfish 34
Bocaccio . . . . . . . . . . . . . . . . . . . . . . . . . 36
Canary rockfish . . . . . . . . . . . . . . . . . . . . . 38
Redstripe rockfish . . . . . . . . . . . . . . . . . . . . 40
Yelloweye rockfish . . . . . . . . . . . . . . . . . . . a 42
Sablefishes - Anoplopomatidae . . . . . . . . . . . . . . . 44
Sablefish . . . . . . . . . . . . . . . . . . . . . . . . 46
iii
Page No . Greenlings . Hexagrammidae . . . . . . . . . . . . . . . . . . . 48
Kelpgreen l ing . . . . . . . . . . . . . . . . . . . . . . . . 50
Whitespot tedgreenl ing . . . . . . . . . . . . . . . . . . . . 52
Lingcod . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 4
Sculpins . Cottidae . . . . . . . . . . . . . . . . . . . . . . 56
Red I r i s h Lord . . . . . . . . . . . . . . . . . . . . . . . . 58
. . . . . . . . . . . . . . . . . . . P a c i f i c staghorn sculpin 60
. . . . . . . . . . . . . . . . . . . . . . . . . . . Cabezon 6 2
Lefteye f lounders . Bothidae . . . . . . . . . . . . . . . . . . 6 4
. . . . . . . . . . . . . . . . . . . . . . . P a c i f i c sanddab 66
. . . . . . . . . . . . . . . Righteye f lounders Pleuronectidae 68
Arrowtooth flounder . . . . . . . . . . . . . . . . . . . . . 70
P e t r a l e so le . . . . . . . . . . . . . . . . . . . . . . . . . 72
F l a t h e a d s o l e . . . . . . . . . . . . . . . . . . . . . . . . 7 4
P a c i f i c ha l ibu t . . . . . . . . . . . . . . . . . . . . . . . 76
Rock so le . . . . . . . . . . . . . . . . . . . . . . . . . . 78
English sole . . . . . . . . . . . . . . . . . . . . . . . . . 80
Sta r ry flounder . . . . . . . . . . . . . . . . . . . . . . . 82
C-0 so le . . . . . . . . . . . . . . . . . . . . . . . . . . . 84
Sand sole . . . . . . . . . . . . . . . . . . . . . . . . . . 86
ACKNOWLEDGMENTS . . . . . . . . . . . . . . . . . . . . . . . . 88
LITERATURE CITED . . . . . . . . . . . . . . . . . . . . . . . . 89
LIST OF FIGURES
Fig. 1 Photos of (A) P a c i f i c cod; (B) P a c i f i c hake; ( C ) P a c i f i c tomcod; (Dl walleye pollock.
Fig. 2 Known d i s t r i b u t i o n of P a c i f i c cod, Gadus macrocephalus, i n Puget Sound.
Fig. 3 Known d i s t r i b u t i o n of P a c i f i c hake, Merluccius productus, i n Puget Sound.
Fig. 4 Known d i s t r i b u t i o n of P a c i f i c tomcod, Microgadus proximus, i n Puget Sound.
Fig. 5 Known d i s t r i b u t i o n of wal leye po l lock , Theragra chalcogranma, i n Puget Sound.
Fig. 6 Photos of (A) s h i n e r perch; (B) s t r i p e d perch; (C) p i l e perch.
Fig. 7 Known d i s t r i b u t i o n of sh ine r perch, Cyrnatogaster aggrega ta , i n Puget Sound.
Fig. 8 Known d i s t r i b u t i o n of s t r i p e d perch , Embiotoca l a t e r a l i s , i n Puget Sound.
Fig. 9 Known d i s t r i b u t i o n of p i l e perch, Rhacochilus vacca, i n Puget Sound.
Fig. 10 Photos of (A) known r o c k f i s h ; (B) copper rock f i sh ; (C) g r e e n s t r i p e r o c k f i s h ; (D) y e l l o w t a i l rock f i sh ; (E) b lack rock f i sh ; (F) qu i l l back r o c k f i s h ; (G) bocaccio; (H) canary rock f i sh ; (I) yelloweye rock f i sh ; and (J) r e d s t r i p e rock f i sh .
Fig. 11 Known d i s t r i b u t i o n of brown r o c k f i s h , Sebas tes a u r i c u l a t u s , i n Puget Sound.
Fig. 12 Known d i s t r i b u t i o n of copper r o c k f i s h , Sebas tes cau r inus , i n Puget Sound.
Fig. 13 Known d i s t r i b u t i o n of g r e e n s t r i p e r o c k f i s h , Sebas tes e longa tus , i n Puget Sound.
Fig. 14 Known d i s t r i b u t i o n of y e l l o w t a i l r o c k f i s h , Sebas tes f l a v i d u s , i n Puget Sound.
Fig. 15 Known d i s t r i b u t i o n of q u i l l b a c k r o c k f i s h , Sebas tes ma l ige r , i n Puget Sound.
Fig. 16 Known d i s t r i b u t i o n of b l ack r o c k f i s h , Sebas tes melanops, i n Puget Sound.
LIST OF FIGURES, cont.
Fig. 17 Known distribution of bocaccio, Sebastes paucispinis, in Puget Sound.
Fig. 18 Known distribution of canary rockfish, Sebastes pimiger, in Puget Sound.
Fig. 19 Known distribution of redstripe rockfish, Sebastes proriger, in Puget Sound.
Fig. 20 Known distribution of yelloweye rockfish, Sebastes ruberrimus, in Puget Sound.
Fig. 21 Photograph of a sablefish.
Fig. 22 Known distribution of sablefish,~Anoplopoma fimbria, in Puget Sound.
Fig. 23 Photograph of (A) female kelp greenling; (B) male kelp greenling; (C) whitespotted greenling; and (D) lingcod.
Fig. 24 Known distribution of kelp greenling, Hexagrammos decagrammus, in Puget Sound.
Fig. 25 Known distribution of whitespotted greenling, Hexagrammos stellari, in Puget Sound.
Fig. 26 Known distribution of lingcod, Ophiodon elongatus, in Puget Sound.
Fig. 27 Photograph of (A) red Irish lord; (B) Pacific staghorn sculpin; and (C) cabezon.
Fig. 28 Known distribution of red Irish lord, Hemilepidotus hemilepidotus, in Puget Sound.
Fig. 29 Known distribution of Pacific staghorn sculpin, Leptocottus armatus, in Puget Sound.
Fig. 30 Known distribution of cabezon, Scorpaenichthys rnarmoratus, in Puget Sound.
Fig. 31 Photograph of a Pacific sanddab.
Fig. 32 Known distribution of Pacific sanddab, Citharichthys sordidus, in Puget Sound.
Fig. 33 Photographs of (A) arrowtooth flounder; (B) petrale sole; (C) flat- head sole; (D) Pacific halibut; (E) rock sole; (F) English sole, (GI starry flounder; (H) C-0 sole; and (I) sand sole.
Fig. 34 Known distribution of arrowtooth flounder, Atheresthes stomias, in Puget Sound.
LIST OF FIGURES, cont.
Fig. 35 Known distribution of petrale sole, Eopsetta jordani, in Puget Sound.
Fig. 36 Known distribution of flathead sole, Hippoglossoides elassodon, in Puget Sound.
Fig. 37 Known distribution of Pacific halibut, Hippoglossus stenolepis, in Puget Sound.
Fig. 38 Known distribution of rock sole, Lepidopsetta bilineata, in Puget Sound.
Fig. 39 Known distribution of English sole, Parophrys vetulus, in Puget Sound.
Fig. 40 Known distribution of starry flounder, Platichthys stellatus, in Puget Sound.
Fig. 41 Known distribution of C-0 sole, Pleuronichthys coenosus, in Puget Sound.
Fig. 42 Known distribution of sand sole, Psettichthys melanostictus, in Puget Sound.
LIST OF TABLES
Table 1.--A listing of the common and scientific names of the recreationally important marine fish other than salmon and smelt in Puget Sound, Wash- ington (Delacy et a1 1972; Buckley 1967, 1968, and 1969).
INTRODUCTION
While numerous marine species have been the object of intensive recrea-
tional fisheries elsewhere along the Pacific Coast, few have received much
attention from anglers in the Puget Sound region. Here, salmon alone is
king-that is, until recently. In addition, of the 45 or so species in the
Sound that may be of potential interest to anglers, few other than salmon
are the target of commercial fisheries. Possibly for these reasons relatively
little has been known of the rockfishes and numerous other "bottom" fish
species. That, however, is also changing. For example, a National Marine
Fisheries Service (NMFS) program, begun in 1973, has now gained considerable
information on the abundance, distribution, and life history of the marine
fish species of recreational importance.
This report constitutes a compilation of data on Puget Sound marine
fishes of potential interest to anglers. Included is information on abundance,
distribution, and life history of bottomfish species--as well as charts
which inform the reader where to find them--compiled from NMFS survey records
and from a review of fishery publications. In this report, Puget Sound is
defined as the marine waters inside or east of a line drawn from Partridge
Point on Whidbey Island to Point Wilson on Quimper Peninsula, and south
of a line drawn from Strawberry Point on Whidbey Island to Utsalady Point
on Camano Island.
Common and s c i e n t i f i c names a r e those provided by t h e American F i s h e r i e s
Socie ty (Bailey e t a 1 1970). A phylogenet ic l i s t i n g of t h e f a m i l i e s and
an a l p h a b e t i c a l l i s t i n g of names of spec ie s i s given i n Table 1; t h e t a b l e
a l s o i n d i c a t e s t h e number of t h e page on which t h e d e s c r i p t i o n of t h e spec ie s
begins. For each spec ie s , a l i s t of r e fe rences has been included by author
and da te of pub l i ca t ion , wi th a comprehensive b ib l iography beginning on
Page 51. These were added i n hopes of making t h i s a more complete r e fe rence
work f o r Puget Sound a n g l e r s a s we l l a s o t h e r s i n t e r e s t e d i n f i s h and f i sh ing .
Table 1.--A listing of the common and scientific names of the recreationally important marine fish other than salmon and smelt in Puget Sound, Wash- ington (Delacy et a1 1972; Buckley 1967, 1968, and 1969).
Codfishes and hakes - Gadidae Pacific cod, Gadus macrocephalus
Pacific hake, Merluccius productus
Tom cod, Microgadus proximus
Walleye pollock, Theragra chalcogramna
Surfperches - Embiotocidae Shiner perch, Cymatogaster aggregata
Striped seaperch, Embiotoca lateralis
Pile perch, Rhacochilus vacca
Rockfishes - Scorpaenidae Brown rockfish, Sebastes auriculatus
Copper rockfish, Sebastes caurinus
Greenstriped rockfish, Sebastes elongatus
Yellowtail rockfish, Sebastes flavidus
Quillback rockfish, Sebastes rnaliger
Black rockfish, Sebastes melanops
Bocaccio, Sebastes paucispinis
Canary rockfish, Sebastes pinni~er
Redstripe rockfish, Sebastes proriger
Yelloweye rockfish, Sebastes ruberrimus
Sablefishes - Anoplopomatidae Sablefish, Anoplopoma fimbria
Greenlings - Hexagrammidae Kelp greenling, Hexagramnos decagrammus
Whitespotted greenling, Hexagrammos stelleri
Lingcod, Ophiodon elongatus
Sculpins - Cottidae Red Irish Lord, Hemilepidotus hemilepidotus
Pacific staghorn sculpin, Leptocottus armatus
Cabezon, Scorpaenichthys rnarmoratus
Lefteye flounders - Bothidae Pacific sanddab, Citharichthys sordidus
Righteye flounders - Pleuronectidae Arrowtooth flounder, Atheresthes stomias
Petrale sole, Eopsetta jordani
Flathead sole, Hippoglossoides elassodon
Pacific halibut, Hippoglossus stenolepis
rock sole, Lepidopsetta bilineata
English sole, Parophrys vetulus
Starry flounder, Platichthys stellatus
C-0 sole, Pleuronichthys coenosus
Sand sole, Psettichthys melanostictus
CODS AND HAKE
The fishes in this family, Gadidae, are real cod and quite common in
the waters of Puget Sound. Included in this group are walleye pollock, Pacific
hake, tomcod, and Pacific cod. Tomcod and Pacific cod are regarded highly
by experienced anglers who know quality foodfish. Pollock is of lesser value,
and hake is regarded as a nuisance to anglers.
Pacific cod, Gadus macrocephalus Pacific hake, Merluccius productus Pacific torncod, Microgadus proximus Walleye pollock, Theragra chalcogramma
NOTE: A l l photos have been de le t ed from t h i s r e p o r t
but w i l l be a v a i l a b l e i n t h e pub l i ca t ion .
' ,
Pacific cod, Gadus macrocephalus
Local Names: True cod, gray cod, cod, Alaska cod.
Recognition: Three separate dorsal fins; the anus is located ventrally below the second dorsal; "whiskert1 below lower jaw is as long as or longer than eye. Brown to gray on back. Numerous brown spots and vermi- culated coloration on back and sides.
Distribution: Common from southern California to Alaska. 14,21
Washington Distribution: Very common in almost all marine waters of this state. Polnell Point, Holmes Harbor, Possession Point, Port Madison, Port Orchard, Rich Passage, Dalco Passage, and Hale Passage are very good areas.
Habitat: Occurs over a wide variety of bottom types (generally mud silt) from shallow to deep water. During winter months, forms large spawning schools; at times found off bottom.
Utilization: Important in commercial catch; 1-2 million pounds are landed annually from Puget Sound. Eagerly sought by a growing number of sport anglers in Washington.
Size: To 39 inches. 14
Usually smaller (to 28 inches--71.3 cm) in Puget Sound, but individuals over 10 pounds are not uncmon.
Eating Qualities: Excellent--firm white flesh; 3-7 pound individuals highest quality and less parasitized. Highly recommended in recipes for (1) fish and chips, (2) poached fish, (3) steamed fish, and (4) smoked fish.
Baits and Lures: Herring, jigs, rubber worms.
Additional References:
1. Alderdice and Forrester 1971. 2. Alverson 1960. 3. Alverson, Pruter, and Ronholt 1964. 4. Andriashev 1937. 5. Barraclough, Robinson, and Fulton 1968. 6. Cobb 1927. 7. DeLacy, Miller, and Borton 1972. 8. Fish. Res. Board Canada 1966. 9, 10. Forrester 1964a, b. 11. Forrester and Ketchen 1955. 12. Grinols 1965. 13, 14. Hart 1949, 1973. 15. Haw and Buckley 1971. 16. Kennedy 1970. 17, 18. Ketchen 1961, 1964. 19. Larkin and Richer 1964. 20. Matsubara 1955. 21. Miller and Lea 1972. 22. Moiseev 1953. 23. Nikolfskii 1961. 24. Okada 1955. 25, 26, 27. Phillips 1951, 1953, 1958. 28. Pinkas 1967. 29. Popov 1933. 30. Rass 1953. 31. Schultz and Welander 1935. 32. Schmidt 1950. 33, 34. Thompson 1962, 1963. 35. Wilimovsky, Peden and Peppar 1967.
Figure 2 Known distribution of Pacific cod, Gadus macrocephalus, in Puget Sound.
P a c i f i c hake, Merluccius productus
Local Names: S i l v e r cod, white f i s h , haddock, b u t t e r f i s h , mellusa.
Recognition: Two d o r s a l f i n s wi th t h e second d o r s a l and t h e ana l f i n both long and notched. There i s no "whisker1' under t h e jaw. The mouth i s l a r g e wi th sharp t ee th . Dark o r s i l v e r y gray on back. Black i n s i d e mouth.
D i s t r ibu t ion : From Baja C a l i f o r n i a t o Alaska. 13,17
Washington Dis t r ibu t ion : C ~ o n i n south Puget Sound, t h e S e a t t l e and Evere t t a r e a , and Hood Canal. Very abundant i n Por t Susan dur ing t h e winter . Found i n much smaller numbers along t h e S t r a i t of Juan de Fuca and San Juans. Puget Sound s tocks a r e known t o be d i s c r e t e from c o a s t a l stocks.
Habi ta t : Usually found over a nonrocky bottom i n moderate water depths. Often found well of f bottom.
U t i l i z a t i o n : A l a r g e corrmercial f i s h e r y e x i s t s during t h e winter around t h e Evere t t a rea . Almost a l l f i s h caught a r e used f o r animal food. Large f o r e i g n f l e e t s t ake g r e a t numbers of hake annual ly o f f t h e coas t of North America. Sold i n Russia a s f resh- f rozen product.
13 Size: To 3 f e e t along c o a s t , somewhat smaller i n Puget: Sound. Specimens
t o 27 inches (68.6 cm) have been c o l l e c t e d i n Puget Sound.
B a i t s and Lures: Herr ing, j i g s .
Eat ing Q u a l i t i e s : F lesh i s very s o f t i f no t cared f o r proper ly (b leed , ev i s - c e r a t e , and i c e immediately). Smaller i nd iv idua l s wi th fewer p a r a s i t e s .
Addi t ional References:
1. Ahlstrom and Counts 1955. 2. Alton and Nelson 1970. 3. Alverson, P r u t e r , and Ronholt 1964. 4. Barraclough 1967a. 5. Barraclough, Robinson, and Ful ton 1968. 6. Best 1963a. 7. Conway 1967. 8. DeLacy, M i l l e r , and Borton 1972. 9. Ginsburg 1954. 10. Gotsha l l 1969. 11. Gr inols 1965. 12. Gr inols and Tillman 1970. 13. Hart 1973. 14. Haw and Buckley 1971. 15 , 16. MacGregor 1966, 1971. 17. M i l l e r and Lea 1972. 18. Mi l l ikan and P a t t i e 1970. 19. Nelson and Larkins 1970. 20, 21. Robinson, Barraclough, and Ful ton 1968a, b , 22. Shippen and Alton 1967.
Figure 3 Known distribution of Pacific hake, Merluccius productus, in Puget Sound.
Pacific tomcod, Microgadus proximus
Local Names: Tomcod, and piciata.
Recognition: Three dorsal fins, with the anus under the first dorsal; the "whiskert' on the lower jaw is about as long as the diameter of the eye. Olive green on back with creamy white sides.
Distribution: From central California to Alaska. 4
Washington Distribution: Found in all marine waters of Washington. Common in San Juan-Bellingharn area as well as around Seattle, Port Orchard, and south Puget Sound. Less common in Hood Canal and the Strait of Juan de Fuca. 2
Habitat: Found over mud silt bottoms. Often found in shallow water in summer.
Utilization: Not the specific target of a commercial fishery.
Size: To 12 inches (30 cm).
Baits and Lures: Herring, rubber worms, jigs.
Eating Qualities: Excellent--firm white flesh. Smaller sizes require a little extra work in cleaning and filleting.
Additional References:
1. Cobb 1927. 2. DeLacy, Miller, and Borton 1972. 3, 4. Hart 1949, 1973. 5. Haw and Buckley 1971. 6. Isaacson 1965. 7. Lamb 1969. 8. Miller and Lea 1972. 9. Wilimovsky 1964.
F i g u r e 4 Known d i s t r i b u t i o n of P a c i f i c tomcod, Microgadus proximus, i n Puget Sound.
Walleye Pollock, Theragra chalcogramma
Local Names: Pollock, whiting.
Recognition: Three well separated dorsal fins. The anus is below the space between first and second dorsals. IWhisker1' on lower jaw small or absent. The eyes are relatively large. Dark green to brown on back (often blotched), silvery on sides.
Distribution: Central California to Alaska. 10
Washington Distribution: Found in all marine waters of Washington. Very common in Strait of Juan de Fuca, central and south Puget Sound. Less com- mon elsewhere.
Habitat: Prefers soft bottoms in moderate depths.
Utilization: An important commercial species in the North Pacific and Bering Sea. Small amounts are landed locally for animal food.
Size: Length to 3 ft. Usually smaller in Puget Sound (22 inches--56 cm).
Baits and Lures: Herring, j igs.
Additional References:
1. Alverson, Pruter, and Ronholt 1964. 2. Armstrong and Winslow 1968. 3. Barraclough 1967a. 4. Barraclough, Robinson, and Fulton 1968. 5. Clemens and Wilby 1961. 6. Cobb 1927. 7. DeLacy, Miller, and Borton 1972. 8. Grinols 1965. 9, 10. Hart 1949, 1973. 11. Haw and Buckley 1971. 12. Larkin and Richer 1964. 13. Matsubara 1955. 14. Miller and Lea 1972. 15. Nikollskii 1961. 16. Okada 1955. 17, 18. Phillips 1942, 1943. 19, 20. Robinson, Barraclough, and Fulton 1968a, b. 21. Wilimovsky, Peden, and Peppar 1967.
Figure 5 Known distribution of walleye pollock, Theragra chalcogramma, in Puget Sound.
SURFPERCHES
I n Puget Sound, t h e sur fperches , family Embiotocidae, have t h r e e repre-
s e n t a t i v e s t h a t a r e important i n t h e spor t ca tch . These inc lude t h e sh ine r
perch, s t r i p e d seaperch, and p i l e perch. This family i s an important group
t o t h e shore-bound a n g l e r , being c l o s e l y a s soc ia t ed wi th barnacle- and mussel-
encrus ted p i l i n g s . I n a d d i t i o n , t hey a r e important forage spec ie s f o r numerous
p reda to r f i s h species .
A r a t h e r unique group, t h e eggs a r e f e r t i l i z e d and develop i n t e r n a l l y ,
being r e l eased a s min ia tu re r e p l i c a s of t h e parent . A pregnant female taken
i n e a r l y summer w i l l very l i k e l y abor t s eve ra l dozen o f f sp r ing .
Shiner perch, Cyrnatogaster aggreaa ta S t r i p e d seaperch, Embiotoca l a t e r a l i s P i l e perch , Rhacochilus vacca
NOTE: A l l photos have been de le t ed from t h i s r e p o r t
but w i l l be a v a i l a b l e i n t h e pub l i ca t ion .
Shiner perch, Cymatogaster aggregata
Local Names: Shiner seaperch, yellow s h i n e r , s h i n e r , bayperch, poggie.
Recognition: Large sca l e s . There i s no f o l d of s k i n a t t ach ing lower jaw t o l i p . S i lve ry . May have t h r e e yellow v e r t i c a l bands on s ides .
D i s t r ibu t ion : From nor the rn Baja C a l i f o r n i a t o southern Alaska. 11
Washington Dis t r ibu t ion : Found i n a l l marine waters of t h i s s t a t e . Very common throughout Puget Sound, Hood4 Canal, and t h e San Juans. Less abundant i n t h e S t r a i t of Juan de Fuca.
Hab i t a t : During summer and f a l l , found i n schools i n shal low water. During t h e winter they apparent ly move i n t o deeper water.
U t i l i z a t i o n : Sometimes used f o r b a i t . There i s a l imi t ed market f o r human consumption.
S ize : Length t o 6 inches. 11
B a i t s and Lures: Clams, mussels , shrimp, worms.
Eat ing Q u a l i t i e s : Occasional ly ea t en d r i e d o r pickled. Considered a de l i cacy by Chinese.
Addi t ional References:
1. Anderson and Bryan 1970. 2. Arai 1967. 3. C lo th ie r 1950. 4. DeLacy, M i l l e r , and Borton 1972. 5. Earnest and Benvi l le 1972. 6. Eigenmann 1894. 7. Gordon 1965. 8. Hart 1973. 9. Haw and Buckley 1971. 10. Hubbs 1917, 1921. 11. Mi l l e r and Lea 1972. 12. Roedel 1953. 13. Shaw, Al len , and Stone 1974. 14. Smith 1969. 15. Suornela 1931. 16. Tarp 1952. 17. Turner 1952. 18, 19. Wiebe 1968a, b.
Figure 7 Known distribution of shiner perch, Cymatogaster aggregata, in Puget Sound.
Striped seaperch, Embiotoca lateralis
Local Names: Blue perch, crugnoli.
Recognition: Several bright blue stripes running along body. Relatively short dorsal fin. Copper color on back with about 15 blue stripes run- ning along body.
Distribution: Pt. Cobras, Baja California to southeast Alaska (Port rang el). 5 ,a
Washington Distribution: Very common in southern an9 central Puget Sound. Less common in San Juans and Strait of Juan de Fuca.
Habitat: During summer and fall, found in shallow water in schools. Found over rocks or often in kelp and eel grass beds.
Utilization: Small amounts caught commercially in this area. Small fishery exists in northern California. Important to shore-bound angler.
Size: Length to 15 inches. 5
Baits and Lures: Clams, mussels, shrimp, worms, small spinners, and flies.
Eating Qualities: Good.
Additional References:
1. Arai 1967. 2. DeLacy, Miller, and Borton 1972. 3. Edwards 1970. 4. Fraser 1923. 5. Hart 1972. 6. Haw and Buckley 1971. 7. Grose 1968. 8. Miller and Lea 1972. 9. Roedel 1953. 10. Smith 1969. 11. Swedberg 1965. 12. Tarp 1952.
Figure 8 Known distribution of striped perch, Embiotoca lateralis, in Puget Sound.
P i l e perch, Rhacochilus vacca
Local Names: P i l e seaperch, dusky seaperch, po rg ie , white seaperch, s i l v e r perch.
Recognition: The d o r s a l f i n h a s a low spiny s e c t i o n followed by a much higher s o f t rayed sec t ion . The t a i l i s deeply forked. Dark gray o r brown on back. S i l v e r y on s ides.
D i s t r ibu t ion : Northern Baja C a l i f o r n i a t o southern Alaska. 6
Washington DiBSribution: Common throughout a l l of Puget Sound. Less common elsewhere.
Habi ta t : P r e f e r s rocky bottoms i n shallow water.
U t i l i z a t i o n : Small amounts caught commercially i n t h e P a c i f i c Northwest. There i s a commercial f i s h e r y i n nor thern Ca l i fo rn ia . Important t o shor ebound angler .
Size: Length t o 17 i n ~ h e s . ~ Ind iv idua l s have been r epor t ed over 19 inches (about 5 l b ) from Puget Sound.
B a i t s and Lures: Clams, mussels , shrimp, worms, smelt , c r abs , small sp inners , and f l i e s .
Ea t ing Qua l i t i e s : Good
Addit ional References:
1. Arai 1967. 2. DeLacy, M i l l e r , and Borton 1972. 3 . Eigenrnann and Ulrey 1894. 4. Hart 1973. 5. Haw and Buckley 1971. 6. Mi l l e r and Lea 1972. 7. Morgan 1961. 8. Randolph 1898. 9. Roedel 1953. 10. Smith 1969. 11. Tarp 1952. 12. Wales 1929. 13. Wares 1968. 14. Wilimovsky 1954.
Figure 9 Known distribution of pile perch, Rhacochilus vacca, in Puget Sound.
ROCKFISHES
The rockf i shes a r e by f a r t h e most important marine f i s h group i n Puget
Sound. There a r e known t o be 21 members of t h e family Scorpaenidae i n t h e
1 "Sound" ; however, f i v e of them (brown, copper, qu i l lback , b l ack , and yellow-
t a i l ) make up 85-90% of t h e t o t a l bot tomfish catch--both i n c i d e n t a l ( t o
salmon) and s p e c i f i c ca t ches , while some o t h e r s a r e sel.dom caught .2 Local
names f o r t h i s spec ie s wrongly a s s o c i a t e them with t h e c:ods, sea bass , and
snapper ( a t r o p i c a l deep-water spec ies) . I n no c a s e i s t h e r e any r e l a t i o n .
The va r ious spec ie s resemble each o t h e r i n body shape but may d i f f e r r a d i c a l l y
i n co lor .
Rockfish, l i k e t h e sur fperches , have an unusual l i f e h i s t o r y i n t h a t
t hey g ive b i r t h t o l i v e young. Unlike t h e sur fperches , t h e young a r e born
a s l a rvae which a r e i n i t i a l l y pe lagic . A female r o c k f i s h may c a r r y 1,000,000
o r more eggs, o r l a r v a e , t h e number varying widely between species . The
time of spawning v a r i e s by spec ie s but i s u sua l ly between winter and summer.
Some spec ies (rasphead and copper r o c k f i s h ) have been found t o con ta in f e r t i -
l i z e d eggs i n summer, which probably means a minor spawning per iod occurs
i n f a l l a l so .
Brown r o c k f i s h , 2. a u r i c u l a t u s Copper r o c k f i s h , 2. caur inus Greens t r ipe r o c k f i s h , 2. e longatus Yel lowta i l r o c k f i s h , S. f l a v i d u s Quil lback rockf i sh , 5; ma1 i g e r Black r o c k f i s h , 2. melanops Bocaccio , 2. p a u c i s p i n i s Canary r o c k f i s h , 2. p i n n i g e r Reds t r ipe r o c k f i s h , 2. p r o r i g e r Rasphead r o c k f i s h , 2. ruberr imus
1 DeLacy, M i l l e r , and Borton 1972. 2 Buckley 1970.
NOTE: All photos have been deleted from this report
but will be available in the publication.
Brown r o c k f i s h , Sebas tes a u r i c u l a t u s
Local Names: Bol ina, sand b a s s , rock cod.
Recogni t ion: S imi l a r t o copper rock f i sh . Brown body wi th dark spot on g i l l cover. Dark t o l i g h t brown, head and body wi th dark vague b a r s a c r o s s back.
D i s t r i b u t i o n : H i p o l i t o yay, Baja C a l i f o r n i a ~ t o sou theas t Alaska. Not common n o r t h of Washington.
Washington D i s t r i b u t i o n : F a i r l y c o T o n i n Puget Sound. Rare ly found i n San Juans , o r S t r a i t of Juan de Fuca.
Hab i t a t : Widely spread i n shal low water ( t o 180 f t ) .
U t i l i z a t i o n : Caught c o m e r c i a l l y bu t no t sought s p e c i f i c a l l y .
4 S ize : To 20 inches. Uncommon over 15 inches , bu t a specimen 17 inches (43.2
cm) and 4 pounds was taken i n Puget Sound.
B a i t s and Lures: Her r ing , squ id , c r ab , clams, a r t i f i c i a l worms, j i g s .
Ea t ing Q u a l i t i e s : Excel lent--s teamed, baked, o r f r i e d .
Add i t i ona l References:
1. DeLacy and Dryfoos 1962. 2. DeLacy, H i t z , and Dryfoos 1964. 3. DeLacy Miller, and Borton 1972. 4. Hart 1973. 5. H i t z 1965. 6. H i t z and DeLacy 1961. 7. Miller and Lea 1972. 8. P h i l l i p s 1957. 9. Roche and Hals tead 1972. 10. Wilimovsky 1954.
3b' 1230 30' ~. 1220 --
Figure 11 Known distribution of brown rockfish, Sebastes auriculatus, in Puget Sound.
Copper rockfish, Sebastes caurinus
Local Names: Whitebelly rockfish, white gopher, barriga blanca, palermotana, copper cod, northern rockfish, bracea, rock cod.
Recognition: Coppery brown coloration, dark fins, light-ly colored along posterior two-thirds of lateral line. Black and dark brown to olive brown, with some copper color; occasionally yellow patches.
Distribution: From Monterey, California, to Kenai Peninsula, Alaska. 3
Washington Distribution: Very common throughout Puget Sound, San Juans, and S rait. of Juan de Fuca. One of the most commorl rockfish in Puget Sound. 5
Habitat: Prefers rocky bottoms or kelp beds. Most often found in shallow water (less than 100 ft).
Utilization: The copper rockfish makes up a significant portion of the commer- cial catch in Puget Sound. Salmon anglers often catch this fish while fishing in water less than 100 feet deep.
Size: To 20 in~hes.~ A number of individuals have been collected from Puget Sound that exceed 20-314 inches (52.7 cm).
Baits and Lures: Herring, squid, crab, clams, worms, artificial lures.
Eating Qualities: Excellent--steamed, baked, or fried.
Additional References: -. 1. DeLacy, Hitz, and Dryfoos 1964. 2. DeLacy, Miller, and Borton 1972. 3. Hart 1973. 4. Haw and Buckley 1971. 5, 6. Hitz 1961, 1965. 7, 8. Hitz and DeLacy 1960, 1965. 9. Miller and Lea 1972. 10. Patten 1973. 11. Schultz and DeLacy 1936. 12. Westrheim 1966a.
Figure 1 2 Known d i s t r i b u t i o n of copper r o c k f i s h , Sebas tes cau r inus , i n Puget Sound.
Greens t r i pe r o c k f i s h , Sebas tes e longa tus
Local Names: Strawberry r o c k f i s h , s e r ena , r ima, p o i n s e t t a , s t r i p e d rock f i sh .
Recogni t ion: Dis t inguished by i t s s l ende r body and green h o r i z o n t a l s t r i p e s on body. Body i s p i n k i s h r e d wi th f o u r h o r i z o n t a l s t r i p e s .
D i s t r i b u t i o n : Cedros I s l a n d , Baja C a l i f o r n i a t o t h e Montague I s l a n d , ~ l a s k a . ~ , ~
Washington D i s t r i b u t i o n : Most abundant i n Hood Canal. 3
Habi ta t : Not known. P r e f e r s deep water ( g r e a t e r t han 100 f t ) .
U t i l i z a t i o n : Small p o r t i o n of i n c i d e n t a l cormnercial ca tch .
4 S ize : To 15 inches. Larges t s p e c i m e n f r o m p u g e t S o u n d 1 3 - 3 / 4 i n c h e s (34.9
cm).
B a i t s and Lures: Her r ing , a r t i f i c i a l l u r e s .
Ea t ing Q u a l i t i e s : Good.
Addi t iona l References:
1. Alverson, P r u t e r , and Ronholt 1964. 2. DeLacy, H i t z , and Dryfoos 1964. 3. DeLacy, M i l l e r , and Borton 1972. 4. Har t 1973. 5. H i t z 1962. 6. M i l l e r and Lea 1972. 7. P h i l l i p s 1957. 8. Roche and Hats tead 1972. 9. Smith 1936. 10. Westrheim 1966. 11. Westrheim, Har l i ng , Davenport and Smith 1968.
Figure 13 Known d i s t r i b u t i o n of g r e e n s t r i p e r o c k f i s h , Sebas tes e longa tus , i n Puget Sound.
Yel lowta i l rock f i sh , Sebastes f l a v i d u s
Local Names: Green snapper, g i o l a , g i a l o t a , chune, yel low rockf ish .
Recognition: Body co lo r greenish t o o l i v e green. T a i l i s a d e f i n i t e yellow, vaguely s t reaked wi th brown.
Di s t r ibu t ion : San Diego, C a l i f o r n i a , t o Kodiak I s l a n d , Alaska. 6 , 9
Washington Dis t r ibu t ion : Found throughout Puget Sound, Hood Canal, and San Juans. Very common along t h e coas t .
Habi ta t : Younger f i s h form schools of f bottom i n shallow water , while o l d e r f i s h can be found i n both deep ( t o 900 f t during day) and shallow water.
U t i l i z a t i o n : Of cons iderable importance (up t o 3 m i l l i o n l b annual ly) t o commercial f i s h e r y , a long t h e Washington coas t . A very Eine s p o r t f i s h .
6 Size: To 26 inches; i n Puget Sound uncommon over 15 inches o r 2 pounds.
B a i t s and Lures: Herr ing, squid, p lugs , spoons, f l i e s , rubber worms.
Eat ing Qua l i t i e s : Excel len t .
Addit ional References:
1. Alverson, P r u t e r , and Ronholt 1964. 2. Carlson and Haight 1972. 3 . DeLacy, H i t z , and Dryfoos 1964. 4. DeLacy, M i l l e r , and Borton 1972. 5. Gr ino l s 1965. 6. Hart 1973. 7. Haw and Buckley 1971. 8. H i t z 1962. 9. Mi l l e r and Lea 1972. 10. Pereyra, Pearcy, and Karvey 1969. 11, 12. P h i l l i p s 1957, 1964. 13. Westr- heim 1966.
Figure 14 Known distribution of yellowtail rockfish, Sebastes flavidus, in Puget Sound.
Quillback rockfish, Sebastes maliger
Local Names: Orange spotted rockfish, yellow backed rockfis:h, brown rockfish, speckled rockfish, rock cod.
Recognition: Very similar to the copper rockfish. Can be distinguished by high fin on back, yellowish blotch on back, dark fins, and freckles. Brown with yellow or tan areas behind head and brownish freckles on jaw. Fins are dark.
Distribution: From central California (point Sur) to Gulf of Alaska. 4,6
Washington Distribution: Common throughout Puget Sound, San uans, and the Strait of Juan de Fuca. Most common rockfish in Puget Sound. 5
Habitat: Prefers rocky bottoms and obstructions on bottom--such as sunken logs, ships or rock piles. Found in 10 to over 500 feet of water.
Utilization: Quillbacks make up a significant portion of the commercial catch of rockfish in Puget Sound. Many are taken by sports fishermen, a large portion incidental to salmon.
4 Size: To 24 inches ; uncommon over 15 inches in Puget Sound (to 15-518 in- ches--39.8 cm or 2-3/4 lb).
Baits and Lures: Herring, candlefish, squid, worms, crabs, plugs, spoons, flies, jigs, rubber worms.
Eating Qualities: Excellent.
Additional References:
1. Alverson, Pruter, and Ronholt 1964. 2. DeLacy, Miller, and Borton 1972. 3. Grinols 1965. 4. Hart 1973. 5. Haw and Buckley 1971. 6. Miller and Lea 1972. 7. Phillips 1957. 8. Roche and Holstead 1972. 9. Wilimovsky 1954.
Figure 15 Known d i s t r i b u t i o n of q u i l l b a c k r o c k f i s h , Sebas tes ma l ige r , i n Puget Sound.
Black r o c k f i s h , Sebas tes melanops
Local Names: Black snapper , b l ack b a s s , ne ro , cherna , c o a s t a l b lack r o c k f i s h , Columbia River r o c k f i s h , blackcod, sea bass .
Recognition: Has l a r g e eyes. When t h e mouth i s c lo sed , t h e upper jaw extends a t l e a s t t o t h e r e a r of t h e eye ( d i s t i n g u i s h e s from b l u e r o c k f i s h ) , b lack over e n t i r e head and body except f o r w h i t i s h b e l l y .
D i s t r i b u t i o n : From southern C a l i f o r n i a t o t h e Gulf of Alaska. 3 ,4 ,6
Washington D i s t r i b u t i o n : Common i n a l l ma i n e waters of Washington except f o r sou th Puget Sound where it i s absent . 5
Habi ta t : Kelp beds , rocky sho re l i ne s . I n summer it i s found i n shal low water. These f i s h a r e o f t e n found o f f t h e bottom.
U t i l i z a t i o n : A small number a r e t aken a s an i n c i d e n t a l c a t c h i n t h e commercial f i s h e r y . Very f i n e l i g h t - g e a r s p o r t f i s h .
Size: To 23 i n ~ h e s . ~ Specimens t o 22 inches (56 cm) and 6 pounds i n Puget Sound with, unconfirmed r e p o r t s of f i s h up t o 10 pounds.
B a i t s and Lures: Her r ing , squ id , p lugs , spoons, sp inne r s , f l i e s , rubber worms, j i g s .
Ea t ing Q u a l i t i e s : Exce l l en t .
Addi t iona l References:
1. Alverson, P r u t e r , and Ronholt 1964. 2. DeLacy, M i l l e ~ r , and Borton 1972. 3. Dunn and Hi t z 1969. 4. Hart 1973. 5. Haw and Buckley 1971. 6. M i l l e r and Lea 1972. 7. P h i l l i p s 1957. 8. Wilimovsky 1954.
Figure 16 Known distribution of black rockfish, Sebastes melanops, in Puget Sound.
Bocaccio, Sebas tes p a u c i s p i n i s
Local Names: Salmon grouper , tomcod, rock cod, spo t t ed rock f i sh .
Recogni t ion: Large mouth. Lower jaw ex tends beyond upper jaw. Dull r ed on head and back. May be pink on s i d e s w i t h dark b l o t c h e s on s ides .
D i s t r i b u t i o n : Baja C a l i f o r n i a t o B r i t i s h Columbia. 3 y 4
Washington D i s t r i b u t i o n : R e l a t i v e l y common i n no r the rn Puget Sound and Hood Canal. Less common i n south Puget Sound. 2
Hab i t a t : P r e f e r s deep water (more t h a n 100 f t ) deep. May be found over a v a r i e t y of hard bottom types .
U t i l i z a t i o n : Caught commercially i n Washington and B r i t i s h Columbia. An important cormnercial s p e c i e s i n C a l i f o r n i a . Recent ly , an important s p o r t s p e c i e s i n south Puget Sound.
Size: To 36 i n ~ h e s . ~ Specimens t o 28-314 inches (73 cm) and 13 pounds c o l - l e c t e d from Puget Sound wi th r e p o r t s of l a r g e r specimens 15 pounds unve r i f i ed .
B a i t s and Lures: Her r ing , squ id , j i g s .
Ea t ing Q u a l i t i e s : F a i r . A l i t t l e g r a i n i e r f l e s h t h a n o t h e r rock f i sh . F l e sh e x c e p t i o n a l l y f a t t y f o r a rock f i sh . Smaller f i s h i n 3-5 pound range b e s t .
Add i t i ona l References:
1. Davenport 1966. 2. DeLacy, M i l l e r , and Borton 1972. 3. Hart 1973. 4. M i l l e r and Lea 1972. 5. Moser 1967. 6 , 7. P h i l l i p s 1964. 8. Roche and Hal- s t e a d 1972. 9. Westrheim 1966b.
- - .
Figure 17 Known distribution of bocaccio, Sebastes paucispinis, in Puget Sound.
Canary rockfish, Sebastes pinniger
Local Names: Orange rockfish, yellow snapper, codalargo, filione, fantail, red rock cod, red snapper, bosco.
Recognition: Head and body orange with some dark speckles on back. Fins are bright orange.
Distribution: Baja California to southeast Alaska. 3 y 5
Washington Distribut 'on: Found in northern Puget Sound, Sa11 ~uans , and Strait of Juan de Fuca. i?
Habitat: Prefers deep water (deeper than 150 ft). Found over hard bottom.
Utilization: An important part of the coastal commercial catch. Marketed "red snapper. 'I
Size: To 30 inches.' Specimens to 22 inches (56.4 cm) and 6 pounds collected from Puget Sound.
Baits and Lures: Herring, candlefish, squid, jigs, rubber worms.
Eating Qualities: Excellent.
Additional References:
1. Alverson, Pruter, and Ronholt 1964. 2. DeLacy, Miller, and Borton 1972. 3. Hart 1973. 4. Haw and Buckley 1971. 5. Miller and Lea 1972. 6,7. Phillips 1957, 1964. 8. Waldron 1968. 9. Westrheim 1966b.
Figure 18 Known distribution of canary rockfish, Sebastes pinniger, in Puget Sound.
Reds t r i pe r o c k f i s h , Sebas tes p r o r i g e r
Local Names: L i t t l e r e d rock cod, r e s t r i p e cod.
Recogni t ion: C lea r red s t r i p e running t h e f u l l l eng th of t h e l a t e r a l l i n e cana l . Green s t r i p e s r a d i a t i n g from eyes. Pa l e r e d , mo t t l ed w i t h green and yellow.
D i s t r i b u t i o n : San Diego, C a l i f o r n i a , t o t h e Bering Sea. 4 ,5
Washington D i s t r i b u t i o n : Moderately common i n San Juanq and no r the rn Puget Sound. Also found i n Hood Canal and south Puget Sound.
Hab i t a t : P r e f e r s moderately deep water over rocky bottoms.
U t i l i z a t i o n : An i n c i d e n t a l commercial spec ies .
S ize : Length t o 20 inches4 , uncommon over 12 inches (30 cm) i n Puget Sound. 2
B a i t s and Lures: He r r ing , j i g s , rubber worms.
Ea t ing Q u a l i t i e s : Exce l l en t .
Addi t iona l References:
1. Alverson, P r u t e r , and Ronholt 1964. 2. DeLacy, M i l l e r and Borton 1972. 3. Gr ino l s 1965. 4. Hart 1973. 4. M i l l e r and Lea 1972. 6. hilli ips 1957. 7. Wilimovsky 1954.
Figure 19 Known d i s t r i b u t i o n of r e d s t r i p e r o c k f i s h , Sebas tes p r o r i g e r , i n Puget Sound.
Yelloweye r o c k f i s h , Sebas tes ruberr imus
Local Names: Red snapper , pot b e l l y , drum, tambor, vecchia , red rock cod, cowfish, tu rkey r o c k f i s h , rasphead rock f i sh .
Recognition: Orange r ed co lo ra t ion . The f i n s a r e t i pped wi th black and t h e eye i s a b r i g h t yellow. Orange yel low over head and body. May be some b lack m o t t l i n g around head.
D i s t r i b u t i o n : Baja C a l i f o r n i a t o Gulf of ~ l a s k a . ~ , "
Washington D i s t r i b u t i o n : Common i n c o a s t a l waters . San Juan I s l a n d s , S t r a i t of Juan de Fuca, Puget Sound, and Hood Canal. 5
Hab i t a t : Found over r e e f s i n deep water (deeper than 200 f t ) .
U t i l i z a t i o n : Important commercial spec ies . Usual ly caught by s e t l i n e s . Sold a s "red snapper." Highly p r i zed by most a n g l e r s because of i t s l a r g e average s i z e .
S ize : To 36 inches.8 Specimens t o 27-314 inches (70.1 cm) and 16 pounds have been c o l l e c t e d i n Puget Sound wi th unconfirmed r e p o r t s of l a r g e r specimens.
B a i t s and Lures: Herr ing, squid , j i g s , spoons, spinners .
Ea t ing Q u a l i t i e s : Exce l len t .
Addi t iona l References:
1. Alverson, P r u t e r , and Ronholt 1964. 2. Barsukov 1964. 3. DeLacy, and Dryfoos 1962. 4. DeLacy, H i t z , and Dryfoos 1964. 5. DeLacy, M i l l e r , and Borton 1972. 6. Gr inols 1965. 7 , 8. Hart 1942, 1973. 9. Haw and Buckley 1971. 10. M i l l e r and Lea 1972. 11. P h i l l i p s 1957. 12. Welander and Alverson 1954. 13. Wilimovsky 1954.
Figure 20 Known distribution of yelloweye rockfish, Sebastes ruberrimus, in Puget Sound.
SABLEFISH
S a b l e f i s h belong t o t h e s k i l f i s h f ami ly , Anoplopomatidae, and a r e a l s o
known a s blackcod (no r e l a t i o n t o P a c i f i c cod ) , t h e a d u l t s of t h i s s p e c i e s
a r e seldom, i f e v e r , found i n Puget Sound. S a b l e f i s h a r e a v a i l a b l e t o Puget
Sound a n g l e r s a s I - , 2-, and 3-year-old juveni les - -probably having en t e r ed
t h e Sound a s p e l a g i c eggs o r l a r v a e i n water masses from t h e ocean o r t h e
S t r a i t of Juan de Fuca. The a d u l t female w i l l produce s e v e r a l hundred thousand
f r ee - f l o a t i n g eggs.
Even though much smal le r t han a d u l t s which can grow t o 40 inches i n
l e n g t h and over 126 pounds , t h e youngs te rs found i n t h e "Sound" a r e " a l l ey
f i g h t e r s , " p u t t i n g up a more t han r e s p e c t a b l e b a t t l e from t h e i r bottom home a t
100 o r so f e e t t o t h e sur face . This s p e c i e s i s ga in ing p o p u l a r i t y amongst
bottom a n g l e r s because of t h e f i r m whi te f l e s h and t h e h igh o i l c o n t e n t , which
makes it a prime smoking f i s h .
S a b l e f i s h , Anoplopoma f imbr i a
NOTE: All photos have been deleted from this report
but will be available in the publication.
Sablefish, Anoplopoma fimbria
Local Names: Blackcod, coalfish, blue cod, butterfish.
Recognition: No spines on head or body. Two well-separated dorsal fins, that are almost equal in size. Black or greenish gray on back with a light gray on underside.
Distribution: Baja California to Alaska. 17
Washington Distribution: Found in almost all marine waters of this state. L L
Common in Strait of Juan de Fuca, northern Puget Sound. Puget Sound has only juveniles. Species leaves by the end of its third year.
Habitat: Found over flat mud silt bottoms over a wide variety of depths.
Utilization: An important coastal commercial species, 40 thousand tons are landed annually, mainly by foreign fishing vessels off the coast. Prized in Midwest as Alaska blackcod--a smoked product. ( ~ e careful, the oil will burn up your electric smoker.)
Size: To 40 inches17 along coast; much smaller (23-112 inches--59.4 cm and 4-114 lb) within Puget Sound (juveniles only).
Bait: Herring (plugged or spinner cut best).
Eating Qualities: The flesh is very oily, which results in an excellent smoked product. Good with beer or poached for breakfast.,
Additional References:
1. Alverson 1960. 2. Alverson, Pruter, and Ronholt 1964. 3. Andriashev 1937. 4. Bailey, Carter, and Swain 1952. 5. Bell and Gharett 1945. 6. Bell and Pruter 1958. 7. Bourne and Pope 1969. 8. Brock 1940. 9. Conway 1967. 10. Cox 1948. 11. DeLacy, Miller and Borton 1972. 12. Edson 1.954. 13, 14. Fish- eries Research Board Canada 1966, 1967. 15. Grinols 1965. 16. Grinols and Gill 1968. 17. Hart 1973. 18. Haw and Buckley 1971. 19. Heyarnoto 1963. 20. Heyamoto and Alton 1964. 21. Holmberg and Jones 1954. 22, 23. Kennedy 1968, 1970. 24. Kennedy and Pletcher 1968. 25. Ketchen and Forrester 1954. 26. Larkin and Ricker 1964. 27. Matsubara 1955. 28. Miller and Lea 1972. 29. Nikol'skii 1961. 30. Pasquale 1964. 31, 32. Phillips 1952, 1958a, 33. Phil- lips, Clothier, and Fry 1954. 34, 35. Pruter 1954, 1959. 36. Roedel 1953. 37. Thompson 1941.
30' 123' 30' 122O Figure 22 Known distribution of sablefish, Anoplopoma fimbria, in Puget
Sound.
GREENLINGS
Members of this family, Hexagramidae, vary widely in coloration and
size from the smaller whitespotted greenling to the massive lingcod (no
relation to Pacific cod, rockfish, or sablefish), which have been caught
weighing up to 105 pounds in British Columbia. Only the lingcod is of comer-
cia1 importance to the United States.
Kelp greenling, Hexagrammus decagramos Whitespotted greenling, Hexagrammos stellari Lingcod, Ophiodon elongatus
NOTE: A l l photos have been deleted from t h i s report
but w i l l be avai lable i n the publication.
Kelp g r e e n l i n g , Hexagrammus decagrammos
Recogni t ion : Two p a i r s of " f l e shy f l a p s " above e y e s , long d o r s a l f i n w i t h about 21 s p i n e s and f i v e l a t e r a l l i n e s on body. Color depends on sex. Females have a s l a t e brown o r g r ay body covered w i t h round r e d d i s h spo ts . The males have s l a t e brown o r g r ay bod i e s w i t h l a r g e t u r q u o i s e s p o t s on t h e head and a n t e r i o r p a r t of body.
D i s t r i b u t i o n : From Los Angeles Harbor t o ~ l a s k a . ~ r 7
Washington D i s t r i b u t i o n : Found i n moderate numbers, i n S t r a i t of Juan d e Fuca, San J u a n s , Bell ingham a r e a , c e n t r a l Puget Sound. Not common i n sou th Puget Sound o r Hood Canal.4
Hab i t a t : Found i n sha l low water ove r rocky and sandy bottoms. Most common i n k e l p beds and over r e e f s .
U t i l i z a t i o n : Not caught commercially.
S i z e : Length t o 22 i n ~ h e s . ~ Specimens t o 17-1/2 i nches (44.5 cm) have been c o l l e c t e d from Puget Sound.
B a i t s and Lures: He r r i ng , c a n d l e f i s h , rubber worms.
E a t i n g Q u a l i t i e s : E x c e l l e n t .
A d d i t i i o n a l References:
1, 2 , 3. Bar rac lough 1967a, b , c. 4. DeLacy, M i l l e r , and Borton 1972. 5. Ha r t 1973. 6. Haw and Buckley 1971. 7. Miller and Lea 1972. 8 , 9. Robinson, Bar rac lough , and Fu l t on 1968a, b. 10. Roedel, Roedel 1953. 11. Wilimovsky 1964.
36' 1 230 50' 1220 Figure 24 Known distribution of kelp greenling, Hexagrammos decagrammus,
in Puget Sound.
Whitespotted greenling, Hexagramrnos stellari
Recognition: One small fleshy flap above each eye, small white spots all over body, five lateral lines. Usually mostly green but may vary from pale red to tan with dark bars or spots. White spots over body.
Distribution: Northern California to Alaska. 5 ?a
Washington Distribution: Found in moderate numbers in Strait of Juan de Fuca, San Juans, and Puget Sound.
Habitat: Prefers rocky areas in shallow water.
Utilization: Not caught commercially.
Size: Length to 24 in~hes.~ Specimens to 14-1/2 inches (37 cm) collected from Puget Sound.
Baits and Lures: Herring, candlefish, jigs, rubber worms.
Eating Qualities: Excellent.
Additional References:
1. Barraclough and Fulton 1968. 2. Barraclough, Robinson, and Fulton 1968. 3. DeLac?, Dryfoos, and Miller 1963. 4. DeLacy, Miller, and Borton 1972. 5. Hart 1973. 6. Haw and Buckley 1971. 7. Matsubata 1955. 8. Miller and Lea 1972. 9. Popov 1933. 10. Roedel 1953. 11. Schmidt 1950. 11, 12. Wilimov- sky 1954, 1964.
Figure 25 Known distribution of whitespotted greenling, Hexagrammos stellari, in Puget Sound.
Lingcod, Ophiodon elongatus
Recognition: Elongated body with l a r g e mouth, l a r g e t e e t h , and long d o r s a l f i n . Very va r i ab le . May be green t o dark brown. Color depends on environ- ment.
D i s t r ibu t ion : Baja C a l i f o r n i a t o Alaska. Grea te s t abundance i s i n v i c i n i t y of B r i t i s h ~ o l u m b i a . ~ ~ r ~ ~ ,29
Washington Dis t r ibu t ion : Found i n a l l marine waters of t h e S ta t e . Most common along t h e c o a s t , t h e western S t r a i t of Juan de Fuca and t h e San Juan I s l ands . Less c&on i n c e n t r a l Puget Sound. 15
Habi ta t : P re fe r rocky bottoms where t h e r e i s a s t rong t i d a l cur rent . Usually found i n deep water i n summer and shallow water i n winter .
U t i l i z a t i o n : An important commercial f i s h . Over 2 m i l l i o n pounds of l ingcod a r e landed annually--mostly from c o a s t a l waters.
Size: Length t o 5 f e e t , 2 4 Specimens t o 41 inches (104 cm) and 26 pounds c o l l e c t e d from Puget Sound wi th many unve r i f i ed r e p o r t s of much l a r g e r specimens.
B a i t s and Lures: Herr ing, p lugs , j i g s , spoons, rubber worms.
Eat ing Q u a l i t i e s : Excel len t . The p a l e b lue lgreen co lo r f l e s h t u r n s white during cooking.
Addi t ional References:
1. Alverson 1960. 2. Alverson, P r u t e r , and Ronholt 1964. 3. Bai ley , Ca r t e r , and Swain 1952. 4 , 5 , 6. Barraclough 1967a, b , c. 7 , 8. Barraclough and Ful ton 1967, 1968. 9. Bio logica l Board of Canada 1935. 10, 11, 12, 13. Chat- win 1954, 1965a, b , 1968. 14. C lo th ie r 1950. 15, DeLacy, M i l l e r , and Borton 1972. 16, 17. F i s h e r i e s Research Board Canada 1954, 1967. 18. F i t c h 1949. 19. F o r r e s t e r 1969. 20. Gr inols 1965. 21, 22, 23, 24. Hart 1.943a, b , 1967, 1973. 25. Haw and Buckley 1971. 26. Ketchen 1950a. 27. Larkfn and Ricker 1964. 28. Mi l l e r and Lea 1972. 29. P h i l l i p s 1959. 30, 31. Robinson, Barra- clough, and Ful ton 1968a, b. 32. Roedel 1948. 33. Wendler 1953. 34. Wilby 1937.
30' 1 23O 30' 1 2 2 O Figure 26 Known distribution of lingcod, Ophiodon elongatus, in Puget Sound.
SCULPINS
The fami ly Co t t i dae i s an extremely l a r g e and d i v e r s e group. While
most a r e important a s forage f o r o t h e r f i s h e s , on ly a few spec i e s a r e l a r g e
enough t o be of t r u e r e c r e a t i o n a l importance. While t h e f l e s h of some spec i e s
i s of h igh q u a l i t y , t h e eggs of t h e Cabezon a r e extremely poisonous.
Red I r i s h l o r d , Hemilepidotus hemilepidotus P a c i f i c s taghorn s c u l p i n , Leptocot tus armatus Cabezon, Scorpaenichthys marmoratus
NOTE: All photos have been deleted from this report
but will be available in the publication.
Red Irish lord, Hemilepidotus hemilepidotus
Local Names: Spotted Irish lord, bullhead.
Recognition: Has a single dorsal fin that has three different levels; there is a band of scales, 4-5 scales wide, surrounding dorsal fin. Variable, mainly red with mottling all over body. Not always red--some may be brownish.
Distribution: From central California to Bering Sea. y 7
Washington Distribution: Common in Puget Sound; not recorded from Hood Canal. 4
Habitat: Occurs in shallow water over rocky bottoms.
Utilization: Not caught commercially.
Size: Length to 20 inches (50.8 cm). 5
Baits and Lures: Herring, jigs, rubber worms.
Eating Qualities: Good.
Additional References:
1. Arai 1969. 2, 3. Barraclough 1967a, b. 4. DeLacy, Miller, and Borton 1972. 5. Hart 1973. 6. Haw and Buckley 1971. 7. Miller and Lea 1972. 8. Peden 1964. 9. Popov 1933. 10, 11. Wilimovsky 1954, 1964.
Figure 28 Known distribution of red Irish lord, Hemilepidotus hemilepidotus, in Puget Sound.
P a c i f i c s taghorn scu lp in , Leptocottus armatus
Local Names: Staghorn scu lp in , bu f fa lo scu lp in , bu f fa lo f i s h , bullhead.
Recognition: NO s c a l e s on body. There i s a l a r g e sp ine on each g i l l cover and a dark spot on t h e r e a r po r t ion of t h e spinous do r sa l f i n . Olive gray t o green wi th some yellow on back. Orange-yellow t o white on stomach.
D i s t r i b u t i o n : Baj a C a l i f o r n i a t o t h e Gulf of ~ l a s k a . 2 7 4
Washington Dis t r ibu t ion : Found i n a l l marine watefs of Washington. Very abundant throughout Puget Sound and t h e San Juans.
Habitat : Usually found i n very shallow water over nonrocky bottoms but may occur i n water over 100 f e e t deep. Often e n t e r s lower p o r t i o n s of r i v e r s and streams.
U t i l i z a t i o n : Not caught commercially. This spec ie s i s one of t h e more common ca tches f o r t h e shore o r p i e r angler . Mainly thought of a s a p e s t , but f i l l e t s a good b a i t . Try l i v e f o r l i n g cod.
Size: Length t o 18 inches (45.7 cm). 2
Ea t ing Q u a l i t i e s : Good.
B a i t s and Lures: Herr ing , j i g s , rubber worms.
Addit ional References:
1. DeLacy, M i l l e r , and Borton 1972. 2. Hart 1973. 3. Haw and Buckley 1971. 4. M i l l e r and Lea 1972. 5. Roedel 1948. 6. Wilimovsky 1954.
Figure 29 Known d i s t r i b u t i o n of P a c i f i c s taghorn s c u l p i n , Leptocot tus armatus, i n Puget Sound.
Cabezon, Scorpaenichthys marmoratus
Local Names: Giant marbled sculpin, bullhead, blue cod.
Recognition: Flaplike fleshy stalk on snout with a mottled appearance over head and body. Green to brown or gray over entire body, mottled with large pale areas. Color is very variable.
Distribution: Baja California to south ~laska. 417
Washington Distribution: Abundant in area of Port Orchard. Found throughout Washington shallow marine waters. 3
Habitat: Prefers shallow water (3-80 ft) over rocky and sandy bottoms.
Utilization: Caught commercially occasionally, but eaten by only the knowlege- able few.
4 Size: Length to 30 inches. Specimens to 24-112 inches (68.1 cm) and 9-114
pounds from Puget Sound with unconfirmed reports of individuals much larger.
Baits and Lures: Squid, ghost shrimp, crabs, jigs, rubber wolms.
Eating Qualities: Excellent. (Caution: the eggs are poisonous.) One of the better eating fish in Puget Sound. Don't let the greenish flesh scare you off (it turns white when cooked).
Additional References:
1. Barraclough 1967a. 2. Clothier 1950. 3. DeLacy, Miller, and Borton 1972. 4. Hart 1973. 5. Haw and Buckley 1971. 6. Hubbs and Wick 1951. 7. Miller and Lea 1972. 8. O'Connell 1953. 9. Pillsbury 1957. 'LO. Quast 1968. 11. Roedel 1948.
Figure 30 Known distribution of cabezon, Scorpaenichthys marmoratus, in Puget Sound.
SANDDABS - LEFT EYE FLATFISHES
The sanddabs ( f ami ly ~ o t h i d a e ) a r e ve ry abundant i n Puget Sound found
over sandy bottoms i n moderate t o shal low wa te r s , 30-80 f e e t . Dabs a r e so
vorac ious a t t imes t h a t they can be p e s t s . Included i n t h i s group a r e t h e
speckled and t h e P a c i f i c sanddab. The P a c i f i c i s t h e l a r g e r and t h e more
commonly caught i n Puget Sound. The speckled i s t o o smal l t o be cons idered
of r e c r e a t i o n a l importance.
P a c i f i c sanddab, C i t h a r i c h t h y s sord idus
NOTE: A l l photos have been d e l e t e d from t h i s r e p o r t
b u t w i l l be a v a i l a b l e i n t h e pub l i ca t ion .
P a c i f i c sanddab, C i tha r i ch thys sordidus
Local Names: Mottled sanddab, megrim, s o f t f l ounde r , sanddab.
Recognition: Mouth on l e f t s i d e of head. There i s a bony br idge between t h e eyes. The diameter of t h e lower eye i s g r e a t e r than t h e l eng th of t h e snout. On eyed s i d e , d u l l brown. May be mot t led wi th black. Blind s i d e i s whi te t o l i g h t brown.
D i s t r i b u t i o n : Baja C a l i f o r n i a t o Alaska. 8 , lO
Washington D i s t r i b u t i o n : Found i n a l l marine wa te r s of Washington. Espec ia l ly common around S e a t t l e , E v e r e t t , and c e n t r a l Puget Sound,. 5
Hab i t a t : Found i n shal low water ( t o 80 f t ) over a v a r i e t y of bottom types . Most common over g r a v e l , sand, o r mud bottoms.
U t i l i z a t i o n : A small commercial f i s h e r y e x i s t s i n B r i t i s h Columbia and Ca l i - f o r n i a . A ve ry good l i v e b a i t f o r l ingcod.
Size: To 16 inches. 8 Uncommon over 10 inches i n Puget Sound ( t o 10-112 inches--26.5 cm).
B a i t s and Lures: Herr ing (bes t sp inner o r plug c u t ) , po lychae tes , clams, j i g s , sp inne r s , f l i e s .
Ea t ing Q u a l i t i e s : Good--highly regarded i n C a l i f o r n i a and by many people i n t h e Puget Sound a rea . Try skinning i n s t e a d of f i l l e t i n g .
Addi t iona l References:
1. Alverson, P r u t e r , and Ronholt 1964. 2. Arora 1951. 3. B a t t s 1964. 4. C l o t h i e r 1950. 5. DeLacy, M i l l e r , and Borton 1972. 6. Dernory 1971. 7. Gr inols 1965. 8. Hart 1973. 9. Haw and Buckley 1971. 10. M i l l e r and Lea 1972. 11. Roedel 1948. 12. Smith 1936. 13. Wilimovsky 1954.
Figure 32 Known distribution of Pacific sanddab, Citharichthys sordidus, in Puget Sound.
FLOUNDERS
This is one of the larger families (~leuronectidae), in terms of numbers
of species and individuals present in Puget Sound. This is probably due
to the fact that the major portion of Puget Sound habitat is well suited
to flatfishes--sand, mud, or silt bottoms.
Arrowtooth flounder, Atheresthes stomias Petrale sole, Eopsetta jordani Flathead sole, Hippoglossoides elassodon Pacific halibut, Hippoglossus stenolepis Rock sole, Lepidopsetta bilineata English sole, Parophrys vetulus Starry flounder, Platichthys stellatus C-0 sole, Pleuronichthys coenosus Sand sole, Psettichthys melanostictus
NOTE: All photos have been deleted from this report
but will be available in the publication.
Arrowtooth f lounde r , Atheres thes s tomias
Local Names: Long jaw f lounde r , t u r b o t , French s o l e , English f lounde r , bas t a rd h a l i b u t , arrowtooth h a l i b u t .
Recognition: Very l a r g e mouth (ex tends behind e y e s ) , and long, sharp t e e t h . It has a s lender body compared t o most f l a t f i s h . Brown on eyed s ide . Blind s i d e i s whi te and f i n e l y d o t t e d wi th black.
D i s t r i b u t i o n : Cen t r a l C a l i f o r n i a t o Bering Sea. It i s most common i n B r i t i s h Columbia and Alaska. 10
Washington D i s t r i b u t i o n : Found i n a l l marine waters. Found throughout Puget Sound, e s p e c i a l l y near E v e r e t t , Po r t Orchard, and throughout c e n t r a l and southern Puget sound.
Hab i t a t : P r e f e r s moderately deep water (more than 75 f t ) and s o f t bottoms.
U t i l i z a t i o n : Large amounts a r e so ld annual ly f o r mink food. There i s minimal market f o r human consumption.
S ize : Length t o 33 inches (83.8 cm).l0
B a i t s o r Lures: Herr ing.
Ea t ing Q u a l i t i e s : F a i r .
Addi t iona l References:
1. Alverson, P r u t e r , and Ronholt 1964. 2. Andriashev 1937. 3. Barraclough and Ful ton 1968. 4. B a t t s 1964. 5. DeLacy, M i l l e r , and Borton 1972. 6 , 7. F o r r e s t e r 1958, 1968. 8. Gotsha l l 1969. 9. Gr ino l s 1965. 10. Hart 1973. 11. Haw and Buckley 1971. 12. M i l l e r and Lea 1972. 13. N i k o l l s k i i 1961. 14. Roedel 1948. 15. Taylor 1967.
Figure 34 Known distribution of arrowtooth flounder, Atheresthes stomias, in Puget Sound.
P e t r a l e s o l e , Eopset ta jo rdani
Local Names: B r i l l , C a l i f o r n i a s o l e , English s o l e , Jo rdan ' s f lounder .
Recognition: Large mouth (max i l l a ry ex tends under eye) . Two rows of t e e t h i n upper jaw. The s c a l e s a r e small . Eyed s i d e i s an even o l i v e brown. Blind s i d e i s white.
D i s t r i b u t i o n : Northern Baja C a l i f o r n i a t o t h e Bering Sea. 17,20
Washington D i s t r i b u t i o n : Abundant la long t h e coas t of 'Washington, bu t i n l i m i t e d numbers i n Puget Sound.
Hab i t a t : P r e f e r s deep water. I n w in te r , moves i n t o very deep water (400 f t o r more) t o spawn. During t h e r e s t of t h e yea r , from 150 t o 200-ft depths.
U t i l i z a t i o n : A very va luab le commercial spec i e Two m i l l i o n pounds a r e taken annual ly by Washington c o a s t a l t r awle r s . 73
Size : Length t o 27 inches (68.6 cm) , I7 g e n e r a l l y smal le r i n Puget Sound.
B a i t s and Lures: Herr ing , c a n d l e f i s h , rubber worms.
Ea t ing Q u a l i t i e s : Exce l len t . Probably t h e f i n e s t e a t i n g "small" f l a t f i s h of t h e n o r t h e a s t e r n P a c i f i c Ocean.
Addi t iona l References:
1. Alderdice and F o r r e s t e r 1968. 2. Alverson 1960. 3. Alverson and Chatwin 1957. 4. Alverson, P r u t e r , and Ronholt 1964. 5. Barraclough 1954a. 6. Ba t t s 1964. 7. Best 1963a. 8. Cleaver 1949. 9. Clemens and Wilby 1961. 10. DeLacy, M i l l e r , and Borton 1972. 11. DiDonato and Pasquale 1970. 12. Evermann and Goldsborough 1907. 13. F o r r e s t e r 1969a. 14. F o r r e s t e r and Alderdice 1967. 15. F o r r e s t e r and Smith 1971. 16. Gr ino l s 1965. 17. Hart 1973. 18. Harry 1959. 19. Haw and Buckley 1971. 20. M i l l e r and Lea 1972.
Figure 35 Known distribution of petrale sole, Eopsetta jordani, in Puget Sound.
Fla thead s o l e , Hippoalossoides elassodon
Local Names: Paper s o l e , c i g a t e t t e paper , f a l s e h a l i b u t .
Recognit ion: Mouth extends t o p u p i l of eye but not beyond, end of t a i l i s n e a r l y s t r a i g h t and t e e t h on t h e upper jaw a r e i n one row. Gray t o brown on eyed s ide . Dusky b l o t c h e s on f i n s .
D i s t r i b u t i o n : Northern C a l i f o r n i a t o Alaska. 9,13
Washington D i s t r i b u t i o n : Found i n a l l marine waters of Washington S t a t e . 4
Habi ta t : P r e f e r s deep water - -usua l ly wel l over 150 f e e t .
Size: Length t o 18 i n ~ h e s . ~ Genera l ly smaller i n Puget Sound.
U t i l i z a t i o n : Genera l ly used i n r educ t ion and animal food.
B a i t s and Lures: Herr ing , c a n d l e f i s h , rubber worms.
Ea t ing Q u a l i t i e s : F a i r t o poor--too much work f o r a small f l e s h y i e l d .
Addi t iona l References:
1. Alverson, P r u t e r , and Ronholt 2. Andriashev 1937. 3. B a t t s 1964. 19?7*5. English 1966. 6. F o r r e s t e r and 4. DeLacy, M i l l e r , and Borton 1972.-
Alderdice 1968. 7. G i l b e r t and Burke 1912. 8. Gr ino l s 1965. 9. Hart 1973. 10. Haw and Buckley 1971. 11. Matsubara 1955. 12. M i l l e r 1969. 13. M i l l e r and Lea 1972. 14. Mineva 1964. 15. S c h i d t 1950. 16. Smith 1936. 17. Townsend 1936. 18. Wilimovsky, Peden, and Peppar 1967.
11 Eng l i sh , T. S. 1966. Engl i sh s o l e egg s t u d i e s . Contract Rep. Northwest - Pulp Pap. Assoc. 93 p.
Figure 36 Known distribution of flathead sole, Hippoglossoides elassodon, in Puget Sound.
Pacific halibut, Hippoglossus stenolepis
Local Names: Northern halibut, right halibut, alabato.
Recognition: Moderately large mouth with conical teeth, arched lateral line, and smooth scales. Any flatfish larger than 15 pounds in this region is likely to be a halibut. Dark brown or gray on eyed side. Usually white on blind side.
Distribution: Southern California to Bering Sea. Uncommon south of Columbia River. 8,15
Washington Distribution: Most common along the Strait of Juan de Fuca. Also found in San Juan Islands, Hood Canal, and Admiralty Inlet.) Not common in central Puget Sound.
Habitat: Prefers water deeper than 100 feet.
Utilization: An important commercial and sports specie:^. Over 50 million pounds were caught commercially in 1970. In Washington the average sport catch is about 1,200 fish annually, mostly by salmon fishermen who generally can only land fish less than 20 pounds.
Size: To 8 feet, 9 inches and over 500 pounds,8 to 200 pounds in Strait of Juan de Fuca and Puget Sound. Several unconfirmed reports of indi- viduals to 50 pounds in Puget Sound.
Baits and Lures: Herring , j igs, spoons, plugs, spinners. Eating Qualities: Excellent. Smaller fish ("chickens") best eating.
Additional References:
1. Alverson, Pruter, and Ronholt 1964. 2. Bell and Pruter 1958. 3. DeLacy, Miller, and Borton 1972. 4. *English 1966. 5. Gray 1964. 6. Grinols 1965. 7. Hardman and Southward 1965. 8. Hart 1973. 9. Haw and Buckley 1971. 10. International Pacific Halibut Commission 1965. 11. Kask 1936. 12. Ketchen 1956a. 13. Larkin and Ricker 1964. 14. Matsubara 1955. 15. Miller and Lea 1972. 16, 17, 18. Myre 1966, 1967, 1969. 19. Novikov 1969. 20. Peltinen 1969. 21. Phillips 1958b. 22. Popov 1933. 23. Ricker 1963. 24. Schmidt 1934. 25. Schmidt 1950. 26, 27, 28, 29. Southward 1961, 1962, 1967, 1968. 30. Southward and Chapman 1965. 31. Thompson and Freeman 1932. 33. Thompson and Herrington 1930. 34. Thompson and VanCleve 1936. 35. Townsend 1936. 36. VanCleve 1936.
%See footnote 1 page 40.
Figure 37 Known d i s t r i b u t i o n of P a c i f i c h a l i b u t , Hippoglossus s t e n o l e p i s , i n Puget Sound.
Rock s o l e , Lepidopset ta b i l i n e a t a
Local Names: Rough back s o l e , g r a v e l s o l e , double- l ine so l e .
Recogni t ion: The l a t e r a l l i n e has a l a r g e a r c h d i r e c t l y behind g i l l cover. The s c a l e s a r e ve ry rough. On eyed s i d e , g ray o r brown, i r r e g u l a r l y blotched.
D i s t r i b u t i o n : Southern C a l i f o r n i a t o Bering Sea. 9 ,14
Washington D i s t r i b u t i o n : Very common i n a l l marine wa te r s i n s i d e of Cape F l a t t e r y .
Hab i t a t : Found over a wide range of depths .
U t i l i z a t i o n : Highly p r i zed a s a foodf i sh . A t a r g e t s p e c i e s of a l a r g e commer- c i a l f i s h e r y along t h e Washington c o a s t and i n Puget Sound. Landings by Washington t r a w l e r s t o t a l about 650,000 pounds a y e a r , and most a r e t aken o f f t h e c o a s t .
9 S ize : To a l eng th of 23 inches : specimens t o 14-314 inches (37.5 an) from Puget Sound.
B a i t s and Lures: Herr ing.
Ea t ing Q u a l i t i e s : Exce l l en t .
Addi t iona l References:
1. Alverson 1960. 2. Alverson, P r u t e r , and Ronholt 1964. 3. B a t t s 1964. 4. DeLacy, M i l l e r , and Borton 1972. 5 , 6 , 7 , F o r r e s t e r 1964b, c , 1969b. 8. F o r r e s t e r and Thompson 1969. 9. Hart 1973. 10. Haw and Buckley 1971. 11. Ketchen, Pe t e r son , and F o r r e s t e r 1951. 12. Larkin and Ricker 1964. 13. Levings 1965. 14. M i l l e r and Lea 1972. 15. Mineva 1964. 16. Nishishiamamoto 1958. 17. Roedel 1948. 18. Shubnikov and Lisovenko 1964. 19. Smith 1936. 20. Wilimovsky, Peden, and Peppar 1967.
Figure 38 Known distribution of rock sole, Lepidopsetta bilineata, in Puget Sound.
English s o l e , Parophrys v e t u l u s
Local Names: Lemon s o l e , common s o l e , C a l i f o r n i a s o l e , pointed nose s o l e , sharp nose so le .
Recognition: Pointed head with a small mouth. The l a t e r a l l i n e does not have an arch. A r a t h e r s lender f i s h . Uniformly brown on eyed s ide . Blind s i d e i s pa l e yellow t o white.
D i s t r ibu t ion : Baja C a l i f o r n i a t o Alaska. 17
Washington Dis t r ibu t ion : Extremely abundant i n a l l marine waters of t h e s t a t e .
Habi ta t : P r e f e r s s o f t o r sandy bottoms. Most of t hese f i s h a r e found i n deep water but dur ing t h e summer,many move i n t o shallow water.
U t i l i z a t i o n : A t a r g e t spec ie s of a s u b s t a n t i a l commerc:ial f i s h e r y along t h e Washington coas t (500,000 l b per yea r ) and i n Puget Sound (1.5 m i l l i o n l b per yea r ) .
Size: Length t o 22 inches.17 Specimens t o 17-112 inches (44.2 cm) from Puget Sound.
B a i t s and Lures: Herr ing, pileworms, clam necks, rubber worms.
Addit ional References:
1. Alderdice and F o r r e s t e r 1968. 2. Alverson 1960. 3. Angell 1972. 4. Ba t t s 1964. 5. Budd 1940. 6. C lo th ie r 1950. 7. DeLacy, M i l l e r , and Borton 1972. 8. Demory 1971. 9 , 10. English 1966*, 1967, 11. El-Sayed 1959. 12. F i s h e r i e s Research Board Canada 1967. 13, 14 , 15, 16. F o r r e s t e r 1956, 1967, 1969b, c. 17. Hart 1973. 18. Harry 1959. 19. Haw and Buckley 1971. 20. Holland 1969. 21. Jow 1969. 22, 23, 24, 25, 26. Ketchen 1945, 1947a, b , 1950b. 27. Ketchen and F o r r e s t e r 1951. 28. Ketchen, Pe terson , and F o r r e s t e r 1955. 29. Larkin and Ricker 1964. 30, 31. Manzer 1946b, 1951. 32. Manzer and Taylor 1946. 33. Menasveta 1958. 34. Mi l l e r and Lea 1972. 35. O r s i 1968. 36. P a c i f i c Marine F i s h e r i e s Commission 1961. 37. Palmer 1954. 38. P a t t i e 1969. 39. P r u t e r and VanCleve 1954. 40, 41. Taylor 1946, 1947. 42. Westrheim 1955b.
+:See f o o t n o t e 1 page 40.
Figure 39 Known distribution of English sole, Parophrys vetulus, in Puget Sound.
Starry flounder, Platichthys stellatus
Local Narnes: Grindstone flounder, great flounder, rough jacket.
Recognition: Fins along side of body have alternating bands of orange and black. Scales are very rough. Individuals with eyes on right side common. Eyed side is dark brown to nearly black with dark blotches. Blind side is white and may have black blotches.
Distribution: Southern California to Alaska. 9,17
Washington Distribution: Very common in all marine waters of Washington.
Habitat: Prefers shallow water over muddy or sandy bottoms. Often found in the lower reaches of streams.
Utilization: There is a commercial fishery for this species in the Gulf of Georgia and Saratoga Passage.
9 Size: Length to 36 inches. Generally smaller in Puget Sound.
Baits and Lures: Herring, jigs, spinners.
Eating Qualities: Fair.
Additional References:
1. Alverson 1960. 2. Alverson, Pruter, and Ronholt 1964. 3. Andriashev 1937. 4. Barraclough 1967c. 5. Batts 1964. 6. Clothier 1950. 7. DeLacy, Miller, and Borton 1972. 8. Forrester 1969a. 9. Hart 1973. 10. Haw and Buckley 1971. 11. C. Hubbs 1947. 12. Hubbs and Hubbs 1945. 13. Manzer 1952. 14. Matsubara 1955. 15, 16. Miller 1965, 1967. 17. Miller and Lea 1972. 18. Nikol'skii, 1961. 19. Okada 1955. 20. Orcutt 1950. 21. Popov 1933. 22,23. Robinson, Barraclough, and Fulton 1968a, b. 24. Roedel 1948. 25. Smith 1936. 26. Town- send 1936. 27. Westrheim 1955. 28. Yusa 1957.
Figure 40 Known distribution of starry flounder, Platichthys stellatus, in Puget Sound.
C - 0 s o l e , P leuronich thys coenosus.
Local Names: Popeye s o l e , C - 0 t u r b o t , mo t t l ed t u r b o t .
Recognition: Very round f i s h wi th a dark spot i n t h e middle of t h e body. Dark brown t o b lack on eyed s ide . F i n s a r e very dark.
D i s t r i b u t i o n : Baja C a l i f o r n i a t o sou theas t ~ l a s k a . ~ y 8
Washington D i s t r i b u t i o n : Uncommon i n S t r a i t of Juan de Fuca and San Juans. More abundant i n Puget Sound south of Admiralty I n l e t . 5
Hab i t a t : P r e f e r s deep water (over 100 f t ) bu t found i n e e l g r a s s beds commonly i n Puget Sound.
U t i l i z a t i o n : The commercial market f o r t h e s e f i s h i s l i m i t e d because t h e i r small s i z e and tough sk in makes f i l l e t i n g d i f f i c u l t .
S ize : Length t o 1 4 inches (35.6 cm). 6
B a i t s and Lures: Small b a i t and lures--worms, shrimp, he r r ing .
Ea t ing Q u a l i t i e s : Good.
Addi t iona l References:
1. B a t t s 1964. 2. Budd 1940. 3 . Clemens and Wilby 1961. 4. C l o t h i e r 1950. 5. DeLacy, M i l l e r , and Borton 1972. 6. Hart 1973. 7. Haw and Buckley 1971. 8. Miller and Lea 1972.
Figure 41 Known distribution of C-0 sole, Pleuronichthys coenosus,, in Puget Sound.
Sand s o l e , P s e t t i c h t h y s me lanos t i c tu s
Local Names: Sand dabs, spo t t ed f lounde r , f r i n g e so le .
Recognition: Rather l a r g e mouth. The f i n r a y s near t h e head a r e f r e e from t h e f i n membrane. Var iab le co lo r . Eyed s i d e may be l i g h t green t o brown, speckled throughout wi th small b lack spots .
D i s t r i b u t i o n : Southern C a l i f o r n i a t o Bering Sea. 7,13
Washington D i s t r i b u t i o n : Very common i n a l l marine waters of Washington.
Habi ta t : P r e f e r s water l e s s t han 100 f e e t deep over sandy bottoms.
U t i l i z a t i o n : A r a t h e r small commercial f i s h e r y e x i s t s f o r t h i s spec ies .
Size: To 25 inchese7 Genera l ly smal le r i n Puget Sound.
B a i t s and Lures: Herr ing , j i g s , rubber worms.
Ea t ing Q u a l i t i e s : Very good,much i n demand.
Addi t iona l References:
1. Alverson, P r u t e r , and Ronholt 1964. 2. B a t t s 1964. 3. Clemens and Wilby 1961. 4. DeLacy, M i l l e r , and Borton 1972. *5. English 1966. 6 , 7. Hart 1944, 1973. 8. Haw and Buckley 1971. 9. Hickman 1959. 10. Manzer 1947. 11, 12. M i l l e r 1965, 1967. 13. M i l l e r and Lea 1972. 14. Roedel 1948. 15. Smith 1936. 16. Soman i 1969. 17. Townsend 1936.
*See foo tno te 1 page 40.
Figure 42 Known distribution of sand sole, Psettichthys melanostictus, in Puget Sound.
ACKNOWLEDGMENTS
I am e s p e c i a l l y g r a t e f u l t o Gregory G. Bargmann, a F ishery Bio logis t
wi th t h e Washington Department of F i s h e r i e s , f o r h i s he lp i n compiling d a t a
and h i s sugges t ions on organiz ing t h e m a t e r i a l i n t h i s paper. I a l s o wish
t o express my g r a t i t u d e t o Raymond Buckley, Mark Pederson, and James Beam
of t h e Washington Department of F i s h e r i e s , t o D r . Bruce Mi l l e r of t h e Uni-
v e r s i t y of Washington, and t o Ronnie Gowan, Lawrence Moulton, and Freder ick
VanderWerff of my s t a f f f o r t h e i r h e l p f u l reviews of e a r l y d r a f t s of t h i s
paper. I am a l s o very a p p r e c i a t i v e of t y p i s t Cheryl Mor isaki ' s e f f o r t s and
pa t ience .
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100, 20 p.
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1955. Pre l iminary r e s u l t s of gray cod tagging i n Georgia S t r a i t i n
t h e winter of 1954-55. Fish. Res. Board Can., Prog. Rep. Pac. Coast
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F o r r e s t e r , C. R., and M. S. Smith.
1971. Ambiocoloration i n a p e t r a l e s o l e (Eopse t ta jordani) . J. Fish.
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1969. Populat ion s t u d i e s on t h e rock s o l e (Lepidopset ta b i l i n e a t a ) of
no r the rn Hecate S t r a i t , B r i t i s h Columbia. Fish. Res. Board Can.,
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1912. F i s h e s from t h e Bering Sea and Kamchatka. Bull . U.S. Bur. Fish.
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1965. Aspects of t h e age and growth of Cymatogaster aggregata Gibbons,
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nor thern Ca l i fo rn i a . Ca l i f . F ish Game 55(1): 75-82.
Gray, G. W.
1964. Hal ibut preying on l a r g e c rus tacea . Copeia 1964 (3) : 590.
Gr ino l s , R. B.
1965. Check-l is t of t h e o f f s h o r e marine f i s h e s occurr ing i n t h e nor th-
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217 p.
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1968. Feeding behavior of t h r e e oceanic f i s h e s (Oncorhynchus k i s u t c h ,
Trachurus symmetricus and Anoplopoma f imbr i a ) from t h e n o r t h e a s t Paci-
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1965. Sampling t h e commercial ca t ch and use of ca: lculated l eng ths i n
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Har l ing , W. R., D. Davenport, M. S. Smith, and R. M. Wowchuk.
1970. - G. 2. Reed groundfish c r u i s e no. 70-3 off southwest Vancouver
I s l a n d , September 9 t o 25, 1970. Fish. Res. Board Can., Tech. Rep.
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Harry, G. Y.
1959. Time of spawning, l eng th a t m a t u r i t y , and f ecund i ty of t h e English,
p e t r a l e and Dover s o l e s (parophrys v e t u l u s , Eopset ta jordani , and Micro-
stomus p a c i f i c u s , r e s p e c t i v e l y ) . Fish. Corn. Oreg., R.es. Br i e f s 7(1) : 5-
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Har t , J. L.
1942. News item. Red snapper fecundi ty . Fish. Res. Board Can., Prog.
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1943b. Migrat ion of l ingcod. Fish. Res. Board Can., Prog. Rep. Pac.
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1944. F l a t f i s h e s i n Skidegate I n l e t . Fish. Res. Board Can., Prog. Rep.
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1948. Age and growth r a t e i n t h e b u t t e r s o l e ( I s o p s e t t a i s o l e p i s ) .
Trans. Roy. Soc. Can., Ser. 3 , 42(5): 65-72.
1949. Food of f i s h of t h e cod family. Fish. Res. Board Can., Prog.
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1967. Fecundi ty and length-weight r e l a t i o n s h i p i n l ingcod. J. Fish.
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1973. P a c i f i c f i s h e s of Canada. Fish. R e s Board Can., Bul l . 180, 740 p.
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1971. Sa l twa te r f i s h i n g i n Washington. S t an l ey N. Jones Pub. CO.,
S e a t t l e , Wash. 192 p.
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1941. F i r s t r eco rd of t h e hybr id f l ounde r , I n o p s e t t a i s c h r y a , from
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Heyarnoto, H.
1962. Age of young s a b l e f i s h , Anoplopoma f imbr i a ( p a l l a s ) 1811. J.
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Heyamoto, H., and M. S. Alton.
1964. S a b l e f i s h , a major r e sou rce of t h e e a s t e r n P a c i f i c ? Pac. Fisherman
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Heyamoto, H., and M. S. Alton.
1964. Sablef i s h , a major r e sou rce of t h e e a s t e r n P a c i f i c ? Pac. F isher -
man 62(10): 25-27.
1965. D i s t r i b u t i o n , abundance, and s i z e of s a b l e f i s h (~noplopoma f imbr i a )
found i n deep water o f f t h e mouth s f t h e Columbia River. C m e r .
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1959. The l a r v a e development of t h e sand s o l e ( ~ s e t t i c h t h y s melanos t ic -
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1962. Seasons of b i r t h of r o c k f i s h (Sebastodes spp.) i n Oregon c o a s t a l
waters . Trans. Am. Fish. Soc. 91(2): 231-233.
1965. F i e l d i d e n t i f i c a t i o n of t h e n o r t h e a s t e r n P a c i f i c r o c k f i s h (Sebas-
t odes ) . U.S. F ish . Wildl. Serv . , Ci rc . 203, 58 p.
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1960. Reproduction and f e c u n d i t y of r o c k f i s h e s ( ~ e b a s t o d e s ) . Univ. Wash.,
Col l . F i sh . Res. F i sh . 1960, Contr ib . 77: 24.
1965. C lea r ing of t h e yo lk i n eggs of t h e r o c k f i s h e s , (Sebastodes cau r i -
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1961. Bottom t r awl ing e x p l o r a t i o n s o f f t h e Washington and B r i t i s h Colum-
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1969. Age, growth and m o r t a l i t y of r a c e s of Engl i sh s o l e ( ~ a r o p h r ~ s
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1954. R e s u l t s of s a b l e f i s h tagging experiments i n Washington, Oregon,
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1917. The breeding h a b i t s of t h e v iv ipa rous perch , Cymatogaster. Copeia
1917 (47): 72-74.
Hubbs, C. L., and L. C. Hubbs.
1954. Data on the life history variation, ecology, and relationships
of the kelp perch, Brachvistius frenatus, an embiotocid fish of the
Californias. Calif. Fish Game 40(2): 183-198.
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1933. Descriptions of two new American species referable to the rockfish
genus Sebastodes, with notes on related species. Univ. Wash., Publ.
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1951. Toxicity of the roe of cabezon Scorpaenichthys mannoratus. Calif.
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1965. Regulation and investigation of the Pacific halibut fishery in
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1969. Results of English sole tagging off California. Pac. Mar. Fish.
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1936. Studies in migration, fishing mortality, and growth in length
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Kennedy, W. A.
1968. Effect of the FP-66 tag on sablefish growth. J. Fish. Res. Board
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Kennedy, W. A.
1970. Reading scales to age Pacific cod adus us macroce~halus) from
Hecate Strait. J. Fish. Res. Board Can. 27: 915-922.
1970. Mor ta l i t y of dar t - tagged cap t ive sablef i s h ( ~ n o ~ l o p o m a f imbria).
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1968. The 1964-65 s a b l e f i s h study. Fish. Res. Board Can., Tech. Rep.
74, 24 p.
Ketchen, K. S.
1945. Prel iminary r e p o r t on age and growth of lemon s o l e s from B r i t i s h
Columbia f i s h i n g grounds. Fish. Res. Board Can., Prog. Rep. Pac.
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1947a. The age, growth and m o r t a l i t y of t h e lemon s o l e Parophrys ve tu lus
Girard on t h e B r i t i s h Columbia f i s h i n g grounds. Masters Thes is , Univ.
B r i t i s h Columbia, Vancouver, 54 p., 33 f i g .
1947b. S tud ie s on lemon s o l e development and egg production. Fish.
Res. Board Can., Prog. Rep. Pac. Coast Stn. 73: 68-70.
1950a. A study of t h e winter t rawl f i s h e r i e s a t Cape Lago and Nanoose
Bay i n t h e S t r a i t of Georgia with spec ia l r e fe rence t o t h e cap tu re
of l ingcod. Fish. Res. Board Can., Prog. Rep. Pac. Coast Stn. 84:
64-67.
1950b. The migra t ion of lemon s o l e s i n no r the rn Hecate S t r a i t . Fish.
Res. Board Can., Prog. Rep. Pac. Coast Stn. 85: 75-79.
1956a. Cl imat ic t r e n d s and f l u c t u a t i o n s i n ~ i e l d of marine f i s h e r i e s
of t h e no r theas t Pac i f i c . J. Fish. Res. Board Can. 13: 357-374.
1956b. Fac to r s inf luencing t h e su rv iva l of lemon s o l e ( ~ a r o p h r y s v e t u l u s )
i n Hecate S t r a i t , B r i t i s h Columbia. J. Fish. R.es. Board Can. 13:
647-694.
1961. Observat ions on t h e ecology of t h e P a c i f i c cod. adus us macrocepha-
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Ketchen, K. S.
1964. Pre l iminary r e s u l t s of s t u d i e s on growth and m o r t a l i t y of P a c i f i c
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1970, An examination of c r i t e r i a f o r determining t h e age of P a c i f i c
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1954, The s a b l e f i s h f i s h e r y of B r i t i s h Columbia. Pac. Mar, F ish , Corn.,
Bull . 3: 57-72.
1955. Migra t ions of lemon s o l e (Parophrys v e t u l u s ) i n t h e S t r a i t of
Georgia. Fish. Res. Board Can., Prog. Rep. Pac. Coast Stn. 87: 27-31,
1966. Populat ion dynamics of t h e p e t r a l e s o l e (Eopset ta i o rdan i ) i n
waters o f f western Canada, Fish. Res. Board Can., Bull . 153, 195
P
Ketchen, K. S., R. I. Pe terson , and C. R. F o r r e s t e r ,
1951. F l u c t u a t i o n s i n t h e l eng th and age compositions of lemon s o l e s
and rock s o l e s i n no r the rn Hecate S t r a i t . Fish. Res, Board Can.,
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Kutty, M. K.
1963. An eco log ica l s tudy and t h e o r e t i c a l cons ide ra t ion of a b u t t e r
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Univ. B r i t i s h Colubmia, Vancouver, 167 p.
Levings, C. D.
1967. A comparison of t h e growth r a t e s of t h e rock s o l e , Lepidopset ta
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1968. F e r t i l i z e d eggs of t h e b u t t e r s o l e , I s o p s e t t a i s o l e p i s , i n Skide-
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MacGregor, J. S.
1966. Fecundity of t h e P a c i f i c hake, Merluccius productus ( ~ ~ r e s ) . Ca l i f .
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1971. Addi t ional d a t a on t h e spawning of t h e hake. Fish. Bull. , U.S.,
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Manzer, J. I.
1946. F i r s t year r e t u r n s of lemon s o l e t a g s used o f f t h e west coas t
of Vancouver I s l and . Fish. Res. Board Can., Prog. Rep. Pac. Coast
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1947. A J u l y spawning popula t ion of sand s o l e s i n Sidney I n l e t . Fish.
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1949. The a v a i l a b i l i t y , e x p l o i t a t i o n , abundance and movement of b u t t e r
s o l e ( I s o p s e t t a i s o l e p i s Lockington) i n Skidegate I n l e t , Queen Char-
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Vancouver.
1951. Growth i n lemon s o l e i n no r the rn Hecate S t r a i t . Fish. Res. Board
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1952. Notes on d i spe r s ion and growth of some B r i t i s h Columbia bottom-
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1947. Rate of growth i n lemon s o l e i n t h e S t r a i t of Georgia. F ish .
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1943. S tud ie s on scorpaenid f i s h e s of Japan: Anatomy, phylogeny, and
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1958. Migration and fishing mortality of English sole (Parophrys vetulus)
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Miller, B. S.
1965. Food and feeding studies on adults of two species of pleuronectids
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Orcas Island, Washington. Masters Thesis, Univ. Washington, Seattle,
131 p.
1967. Stomach contents of adult starry flounder and sand sole in East
Sound, Orcas Island, Washington. J. Fish. Res. Board Can. 24: 2515-2526.
1969. Life history observations on normal and tumor-bearing flathead
sole in East Sound, Orcas Island, Washington. Ph.D. Thesis, Univ.
Washington, Seattle, 131 p.
1970. Food of flathead sole (~i~~oglossoides elassodon) in East Sound,
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1972. Guide to the coastal marine fishes of California. Calif. Dep.
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1970. Hermaphroditism in a Pacific hake, Merluccius productus, from
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1964. (On the biology of some flatfishes in the eastern Bering sea).
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1953. (Cod and f l o u n d e r s of f a r e a s t e r n s ea s ) . Izv . Ilikhookean. Nauch.-
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1961. S i l e t z Bay sur f perch tagging. Fish. Comm. Oreg., Res. B r i e f s
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1967. Reproduct ion and development of Sebastodes p a u c i s p i n i s and com-
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1966. Loss of t a g s from P a c i f i c h a l i b u t a s determined by double- tag
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1968. The embryology of t h e English s o l e , Parophrys ve tu lus . Cal i f .
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1961. Migrat ions of English s o l e ( ~ a r o p h r ~ s ve tu lus ) on t h e P a c i f i c
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1954. A comparison of o t o l i t h s and in t e rope rcu la r bones a s age i.ndi-
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1973. Bio logica l information on copper rockf i sh i n Puget Sound, Washing-
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P h i l l i p s , J. B.
1942. Walleye pol lock caught i n Monterey Bay. C a l i f . F i sh Game 28(3):
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1943. Another walleye pol lock a t Monterey. C a l i f . F i sh Game 29(1):
83.
1951. P a c i f i c cod of f c e n t r a l C a l i f o r n i a . C a l i f . F i sh Game, 37(2):
351.
1952. Yellow s a b l e f i s h (blackcod) taken i n Monterey Bay. C a l i f . F i sh
Game 38(3): 437-438.
1953. Addi t iona l P a c i f i c cod taken of f c e n t r a l Ca l i fo rn i a . C a l i f . F i sh
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1957. A review of t h e r o c k f i s h e s of C a l i f o r n i a ( fami ly Scorpaenidae).
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Popov, A. M.
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1959. Tagging exper iments on s a b l e f i s h a t Holmes Harbor , Washington.
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1954. Eng l i sh s o l e i n Holmes Harbor , Washington. Wash. Dep. F i sh . ,
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1964. Recent occurrences of intergeneric hybrid flounders, Inopsetta
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Reeves, J. E.
1966. An estimate of survival, mortality, and the number of ling cod
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1953. Common ac.ean fishes of the California coast. Calif. Dep. Fish
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1936. Keys to the fishes of Washington, Oregon and closely adjoining
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1936. Fishes of the American Northwest: A catalogue of the fishes of
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1936b. Fishes of the American Northwest: A catalogue of the fishes
of Washington and Oregon with distributional records and a bibliogra-
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1936~. Fishes of the American Northwest: A catalogue of the fishes
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1936d. Fishes of the American Northwest: A catalogue. of the fishes
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1962. A method of calculating body lengths from otolith measurements
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1952. An accessory r e s p i r a t o r y device i n embryos of t h e embioi-ocid
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1927. The rediscovery of Inopse t t a i schyra , , a r a r e spec ie s of f lounder
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$ -
Waldron, K. D.
1968. Ea r ly l a r v a e of t h e canary r o c k f i s h , Sebastodes p inniger . J.
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Wales, J. H.
1929. A n o t e on t h e breeding of t h e v iv ipa rous perch Damalichthys.
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1966a. Northern range extens ion records f o r two rockf i sh spec ies (Sebas-
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1456.
1966b. Northern range extens ions f o r t h r e e species of rockf i sh ( ~ e b a s -
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1781-1809.
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1968. Prel iminary r e p o r t on m a t u r i t y , spawning and l a r v a l i d e n t i f i c a t i o n
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Wiebe, J. P.
1968. The e f f e c t s of temperature and day length on t h e physiology of
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1937. The l ingcod, Ophiodon e lonpatus Girard. Biol. Board Can., ~ u l l .
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W i l imovsky, N. J.
1954. L i s t of t h e f i s h e s of Alaska. Stanford ~ c h t h y o l . Bull. 4(5):
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1964. Inshore f i s h fauna of t h e Aleut ian archipelago. Science i n Alaska,
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Assoc. Advance Sci .
Wilimovsky, N. J., A. Peden, and J. Peppar.
1967. Systematics of six demersal fishes of the North Pacific Ocean.
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1969. Relationships of female age and size to embryo number and size
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1957. Eggs and larvae of flatfishes in the coastal waters of Hokkaido.
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