Case study IIIThe continental scale: home ranges

Species differ in home rangeHome ranges may be continuous or scattered.Scattered distributions might not allow for any gene flow. Such regional subpopulation quickly evolve into new species

Many species have seasonal migrations. Then summer and winter home ranges differ.

The UTM grid of Poland used for species distribution atlases

Philaeus chrysops

Regression analysis with environmental factors allows for the assessment of habitat preferences.

No gene flow

Relict speciesIntroduced speciesHabitat specialists

S AlpsC BalkansE France

LanguedocW FranceMarche

VenetianRomagnaTuscany

CalabriaP. liolepisP. lsiculusP. melidellensis

Miocene Pliocene Pleistocene H

Case study IIIThe continental scale: home ranges

How did Podarcis muralis colonise Europe aft the glacial period?

How will the species spread in the face of climate change?

Podarcis muralis

olocene

Substitutions

Species distribution modelling (MaxEnt) under five different climatic scenarios

Distance from Athens 9.03 9.87 17.20 17.54 18.18 18.68 18.69 18.91 18.92 19.06 19.10 19.11 19.12 19.21 19.25 20.35 20.50 21.95 22.29 22.53S 103 56 55 63 95 49 64 32 51 30 46 44 48 58 49 33 69 42 43 61

S Locus 32 33 17 16 19 12 13 15 11 10 29 4 1 5 2 28 22 24 23 2519 120 0.083 0.056 0.5 0.184 0.125 0.364 0.367 0.611 0.25 0.25 0.1 0.5 0.339 0.167 0.417 0.028 0.038 0.063 0.083 0

18 127 0.417 0 0.175 0.211 0.063 0.409 0.75 0.5 1 1 0.6 0.833 1 0.733 0.917 0 0.885 0.75 1 0.333

18 132 0.167 0.222 0.65 0.684 0.563 0.818 0.65 1 1 0.6 0.45 0.833 0.143 0.833 0.958 1 0 0.125 0 0.722

18 118 0.042 0.167 0.125 0.158 0.063 0.227 0.367 0.389 0.111 0.1 0.25 0 0.482 0.2 0 0.667 0.077 0.313 0.333 0.111

17 153 0.042 0.111 0.125 0.184 0.188 0.273 0.1 0 0.056 0.05 0.05 0.056 0.089 0 0.167 0 0.692 0.063 0.167 0.333

16 154 0.125 0 0.875 0.5 0.438 0.409 0.783 0.111 0.806 0.7 0.35 0.722 0.982 0.7 0.958 0.972 0 0 0 0.111

16 151 0.125 0.056 0.1 0.132 0.125 0.045 0.35 0 0.111 0.1 0 0 0 0.233 0.042 0.444 0.231 0.5 0.5 0.389

16 155 0.083 0.056 0.5 0.211 0.281 0.136 0.067 0 0.167 0.7 0.3 0.333 0.357 0.4 0.375 0 0 0.125 0 0.056

15 160 0.083 0 0.125 0.368 0.188 0.591 0.183 0.889 0.083 0.3 0.55 0.278 0.018 0.3 0.042 0 0 0 0 0.5

14 205 0 0 0.375 0.184 0.281 0.364 0.4 0.222 0.139 0 0 0.111 0.482 0.5 0.125 0 0.115 0 0.333 0.111

14 208 0 0 0.175 0.184 0 0.227 0.133 0.056 0.694 0 0 0.389 0.482 0.1 0.333 0.056 0.077 0.125 0.083 0

14 190 0 0 0.6 0.263 0.188 0.364 0.333 0 0.139 0 0 0.556 0.286 0.467 0.292 0 0.077 0.25 0.333 0.167

14 192 0 0 0.075 0.263 0.063 0.045 0.183 0.167 0.333 0.4 0 0.167 0.036 0.033 0.083 0 0.115 0 0 0.222

13 219 0.083 0.111 0 0 0.094 0.227 0.2 0 0.306 0 0.5 0.222 0.482 0.2 0.5 0.278 0 0 0 0.056

13 221 0.125 0 0.225 0.079 0.125 0.091 0.1 0.778 0.25 0 0.1 0.389 0 0.167 0.083 0 0 0 0 0.167

13 188 0.167 0 0.05 0.105 0.313 0.182 0.033 0.444 0 0 0.7 0 0.036 0 0.417 0.833 0.038 0 0.083 0

13 135 0.042 0 0.35 0.316 0.188 0.182 0.35 0 0 0.4 0.55 0.167 0.839 0.167 0.042 0 0 0 0 0.111

13 157 0 0.056 0.15 0.237 0.125 0.045 0.267 0.444 0.333 0 0.45 0.556 0.036 0.233 0.042 0 0 0 0 0

12 133 0 0.056 0 0 0.063 0 0.017 0 0.028 0.05 0 0.056 0.054 0.067 0.083 0.278 0.038 0 0.083 0

12 197 0 0 0 0 0.031 0.273 0.067 0.278 0 1 0 0.333 0 0.133 0.083 0 0.346 0.875 0.5 0.611

12 116 0.042 0.167 0.225 0.184 0.25 0 0 0 0.028 0 0.05 0 0.018 0.033 0 0 0.269 0.188 0 0.167

11 131 0 0 0.7 0.263 0.344 0.409 0.25 0.5 0 0 0 0.167 0 0.233 0.083 0 0 0.063 0 0.167

11 109 0 0 0.25 0.079 0.156 0.273 0.3 0.222 0.333 0 0 0.056 0 0.1 0 0 0.154 0 0 0.056

11 137 0 0 0 0 0.094 0 0.083 0.056 0.139 0 0 0.222 0.018 0.167 0.083 0.389 0 0 0.25 0.056

11 193 0.083 0 0 0 0.25 0 0.017 0.056 0 0 0.05 0.056 0 0.033 0 0.056 0.077 0 0.083 0.111

11 194 0 0.056 0.075 0.184 0.125 0.045 0.017 0 0 0 0.1 0 0 0 0 0.111 0.115 0.063 0 0.167

11 149 0.333 0.389 0 0.184 0.156 0.091 0 0.056 0 0 0.1 0 0 0 0 0.083 0.077 0.25 0 0.111

11 122 0 0 0.15 0.263 0.25 0.273 0.083 0 0.444 0.6 0.1 0.278 0 0.2 0 0 0 0 0 0.056

10 153 0 0 0 0 0.033 0 0 0.444 0.028 0.1 0.15 0.167 0.054 0.2 0.417 0 0.038 0 0 0

10 215 0.083 0 0.025 0.026 0 0.045 0.15 0 0 0.4 0.05 0.111 0 0.1 0.125 0 0 0 0 0

10 225 0.083 0 0 0.053 0.031 0.136 0.2 0 0.139 0.3 0 0 0 0.233 0.208 0.194 0 0 0 0

10 125 0.083 0.667 0 0.237 0.063 0.136 0 0 0 0 0.4 0 0 0.033 0 1 0.115 0.125 0 0

10 196 0.083 0.833 0.025 0 0.063 0 0.267 0.056 0.167 0 0.25 0 0 0.033 0 0 0.231 0 0 0

10 186 0.042 0 0.15 0 0.031 0.182 0.35 0.389 0.306 0 0.15 0 0 0 0.042 0 0 0 0.083 0

9 223 0.042 0 0.25 0.053 0.094 0 0 0 0.028 0.1 0 0.111 0 0.033 0 0 0 0 0 0.111

9 117 0 0.167 0.1 0.211 0.063 0.455 0.4 0.389 0.139 0 0 0 0 0 0 0 0 0 0 0.056

9 131 0.167 0 0.025 0 0.125 0 0 0 0.167 0.75 0 0 0.393 0.233 0.208 0 0 0 0.083 0

9 135 0.125 0 0.025 0 0.188 0 0.05 0 0.111 0 0.1 0.111 0 0.067 0 0 0 0 0 0.056

9 196 0 0 0 0.132 0.063 0.045 0.017 0 0 0.5 0 0 0.089 0 0.167 0 0.115 0 0 0.278

9 198 0 0 0.05 0 0 0.136 0 0 0.111 0.05 0.05 0.056 0.018 0.2 0 0 0 0 0 0.056

9 159 0.208 0 0 0.053 0 0 0.017 0.222 0 0 0.05 0 0.232 0.1 0.125 0.306 0 0 0 0

9 110 0.292 0.167 0 0 0.063 0 0.067 0 0.167 0 0 0 0 0.133 0 0 0.038 0 0.083 0.111

9 114 0.042 0.278 0 0.079 0.094 0 0 0 0 0 0.25 0.056 0 0 0.083 0 0.192 0 0.25 0

9 124 0.167 0 0 0 0.063 0.091 0.017 0 0 0.05 0 0.056 0.054 0.033 0.208 0 0 0 0 0

8 151 0 0 0.025 0 0.3 0.045 0 0 0.028 0 0.15 0.167 0.25 0.067 0 0 0 0 0 0

8 159 0.042 0 0 0 0.033 0 0.033 0 0.111 0 0 0.167 0 0.2 0 0.194 0.077 0 0 0

8 217 0 0 0.15 0.184 0.156 0 0 0 0 0 0.05 0.056 0.089 0 0 0.028 0 0 0 0.333

8 127 0.167 0 0 0 0.063 0 0.017 0 0 0.15 0 0.111 0 0 0.042 0.167 0 0 0 0.056

8 139 0.042 0.111 0.025 0 0 0 0 0 0.028 0 0.15 0 0.089 0.233 0.125 0 0 0 0 0

8 184 0 0 0 0.053 0.031 0 0.067 0 0.083 0.05 0 0 0.107 0 0 0.056 0 0 0.083 0

7 141 0.042 0 0 0.026 0.033 0.455 0.067 0 0.056 0 0 0 0 0 0 0 0 0.125 0 0

7 154 0.125 0 0 0 0 0 0.033 0 0.167 0 0 0 0.161 0 0.25 0 0 0.063 0 0.278

7 155 0.042 0 0 0 0 0.182 0 0 0.194 0.8 0.45 0.167 0 0 0 0.583 0 0 0 0

7 157 0 0 0 0 0.067 0.045 0 0 0.028 0 0 0.056 0.036 0.033 0 0.222 0 0 0 0

7 209 0 0.389 0.2 0.342 0 0 0.017 0 0.25 0.05 0 0 0.071 0 0 0 0 0 0 0

7 213 0 0 0.05 0.053 0 0 0.017 0 0 0 0.1 0 0 0.233 0 0.5 0 0 0 0.111

7 115 0.083 0 0.525 0.395 0 0.182 0 0 0.056 0 0.55 0 0 0 0 0 0.038 0 0 0

7 141 0 0.111 0 0 0 0.091 0 0 0 0 0.05 0 0.25 0.1 0.167 0 0 0 0 0.056

7 159 0.083 0.111 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0.154 0.25 0.5 0.056

7 161 0 0.056 0.05 0 0 0.182 0 0 0.139 0 0 0 0.286 0.033 0 0 0 0 0 0.111

6 149 0 0 0.025 0 0.067 0.091 0 0.056 0.222 0 0 0 0 0 0.083 0 0 0 0 0

6 156 0.042 0 0 0 0 0.136 0.067 0.222 0 0 0 0.278 0.018 0 0 0 0 0 0 0

6 201 0.125 0 0.075 0.158 0 0 0.05 0 0 0 0 0 0.161 0.033 0 0 0 0 0 0

6 211 0 0.111 0.025 0.053 0 0 0.15 0 0 0 0.1 0 0 0 0.083 0 0 0 0 0

6 111 0.042 0.278 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.346 1 0.583 0.556

6 129 0 0 0 0 0 0 0.05 0 0 0.05 0 0.222 0.071 0.033 0 0 0 0 0 0.056

6 194 0.25 0 0 0 0 0 0.117 0.333 0 0 0 0.056 0 0.067 0.375 0 0 0 0 0

6 166 0.083 0 0 0 0.156 0 0.017 0 0 0 0 0 0 0 0 0 0.038 0 0.5 0.111

6 175 0.042 0.111 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.115 0.438 0.083 0.167

6 174 0.042 0 0 0 0.094 0 0 0 0 0 0 0 0 0 0 0 0.231 0.563 0.083 0.056

6 147 0 0 0.075 0 0 0.091 0.1 0 0 0 0.05 0 0 0 0 0.028 0 0.063 0 0

6 126 0 0 0 0 0 0 0.083 0 0 0 0.05 0 0.018 0.167 0 0.306 0 0 0 0.111

5 145 0.042 0 0.025 0.263 0 0 0.05 0 0 0 0 0 0 0 0 0 0 0.063 0 0

5 148 0 0 0.025 0 0 0 0.383 0 0 0 0.25 0 0.071 0 0 0 0 0 0 0.111

5 152 0 0 0 0 0 0.045 0 0 0.028 0 0 0 0.196 0.033 0 0 0 0.125 0 0

5 158 0 0 0 0 0.033 0 0 0 0.083 0 0 0 0.214 0.067 0 0 0 0 0 0.111

5 164 0.125 0.167 0 0.026 0 0 0.033 0 0 0 0 0 0 0.167 0 0 0 0 0 0

5 165 0 0 0.15 0 0.133 0 0.233 0.278 0 0 0 0 0 0 0 0 0 0 0 0.222

5 203 0 0 0 0 0.156 0.455 0 0 0 0 0.1 0 0.196 0 0 0 0 0.5 0 0

5 129 0.167 0 0 0.211 0.156 0.045 0 0 0 0 0 0 0 0 0 0 0 0.063 0 0

5 107 0.167 0 0 0.132 0.156 0 0 0 0 0 0 0.222 0 0 0 0.167 0 0 0 0

5 121 0 0 0 0.079 0 0 0 0.167 0 0 0 0 0 0 0.042 0 0.038 0 0 0.056

5 200 0 0 0 0 0.031 0 0 0 0.028 0 0 0.222 0 0.3 0 0 0 0.125 0 0

5 156 0.125 0.167 0 0 0.094 0 0 0 0 0 0 0 0 0 0 0 0.115 0.25 0 0

5 163 0.125 0.056 0 0 0.063 0 0 0 0.194 0 0 0.056 0 0 0 0 0 0 0 0

5 112 0.083 0.056 0 0 0.063 0 0 0 0 0 0.2 0 0 0 0 0 0 0 0 0.056

4 150 0 0 0 0 0 0 0.083 0 0 0.1 0 0 0 0.067 0 0 0 0 0 0.222

4 167 0 0.333 0 0 0.1 0 0 0 0 0 0 0 0 0 0 0 0.077 0.25 0 0

4 173 0.042 0 0 0 0.033 0 0 0 0 0 0 0 0 0 0 0 0.154 0.063 0 0

4 177 0.083 0.111 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.038 0 0.083 0

4 227 0 0.111 0 0 0 0 0.117 0 0.028 0 0 0.111 0 0 0 0 0 0 0 0

4 211 0 0 0 0.053 0.063 0 0 0 0 0 0 0 0 0.1 0.042 0 0 0 0 0

4 130 0.542 0.778 0 0 0.031 0 0 0 0 0 0.05 0 0 0 0 0 0 0 0 0

4 163 0 0 0 0.026 0 0 0 0 0.056 0 0 0 0 0 0 0 0 0 0.167 0.056

4 169 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.077 0.063 0 0.056

4 172 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.115 0.125 0.167 0

4 178 0 0 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0.231 0.25 0.083 0

4 162 0 0.056 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0.192 0 0.333 0

4 143 0 0 0 0 0.031 0 0.017 0 0 0 0 0 0 0 0.25 0 0 0 0.083 0

4 100 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.154 0.25 0.167 0.389

4 128 0 0 0 0 0 0 0 0 0 0 0 0.111 0.089 0 0.25 0 0 0 0.083 0

4 130 0 0 0 0 0.031 0.045 0 0 0 0 0 0 0 0.033 0.042 0 0 0 0 0

3 142 0 0 0.3 0.158 0.033 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

3 143 0 0 0.025 0.132 0.033 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

3 144 0 0 0.1 0.132 0 0 0 0 0 0 0 0 0 0 0.167 0 0 0 0 0

3 162 0 0 0 0 0 0 0 0 0.056 0 0 0 0 0 0.083 0 0.038 0 0 0

3 174 0.125 0.056 0 0 0.033 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

3 207 0 0 0 0 0.281 0 0 0.111 0 0.15 0 0 0 0 0 0 0 0 0 0

3 261 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.038 0 0 0.222

3 267 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.115 0 0.25 0

3 143 0 0 0.025 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.167

3 145 0 0 0.025 0 0.094 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.278

3 113 0 0.278 0.05 0 0 0 0 0 0 0 0 0 0 0 0 0 0.385 0 0 0

3 119 0 0 0 0.053 0.063 0 0 0.167 0 0 0 0 0 0 0 0 0 0 0 0

3 143 0 0 0 0 0 0 0.067 0 0 0 0 0 0.125 0 0.042 0 0 0 0 0

3 191 0.083 0 0 0 0.125 0 0 0 0 0 0 0 0.036 0 0 0 0 0 0 0

3 199 0 0 0.425 0.447 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.111

3 204 0.125 0 0 0 0 0 0 0 0 0 0.6 0 0 0 0 0.389 0 0 0 0

3 176 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.231 0 0.25 0

3 138 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.038 0 0 0.056

3 147 0.042 0 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0 0.063 0 0

3 145 0 0 0 0 0 0 0.083 0.278 0 0.15 0 0 0 0 0 0 0 0 0 0

3 102 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.231 0.188 0 0

2 139 0 0 0 0.053 0.033 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 146 0 0 0 0 0 0 0.017 0 0 0 0 0 0 0.033 0 0 0 0 0 0

2 147 0 0 0 0.132 0 0 0 0 0 0 0 0 0 0.133 0 0 0 0 0 0

2 163 0 0 0 0.026 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.333 0

2 170 0 0.111 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.269 0 0 0

2 171 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.077 0 0.083 0

2 172 0.042 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 175 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.077 0 0.333 0

2 176 0.083 0.111 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 229 0.083 0 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 263 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.462 0 0.333 0

2 265 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.269 0 0.333 0

2 133 0 0.333 0 0.079 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 141 0 0 0.075 0 0.063 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 123 0 0 0 0 0 0 0 0 0 0 0.05 0 0 0 0.125 0 0 0 0 0

2 125 0 0 0 0 0.031 0 0 0 0 0 0.1 0 0 0 0 0 0 0 0 0

2 145 0 0 0 0 0 0 0.017 0 0 0 0 0 0 0 0.042 0 0 0 0 0

2 190 0 0 0 0 0.188 0 0 0 0 0 0 0 0 0 0 0.139 0 0 0 0

2 198 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0.042 0 0 0 0 0

2 200 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.038 0 0 0.056

2 201 0.083 0 0 0 0 0 0 0 0 0 0.1 0 0 0 0 0 0 0 0 0

2 202 0 0 0 0 0 0.136 0 0 0 0 0 0 0 0.033 0 0 0 0 0 0

2 210 0 0 0 0.026 0 0 0 0 0 0 0 0 0 0 0 0.333 0 0 0 0

2 148 0.042 0 0 0.105 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 151 0 0 0 0 0.063 0 0.017 0 0 0 0 0 0 0 0 0 0 0 0 0

2 157 0.042 0 0 0 0 0 0 0 0.056 0 0 0 0 0 0 0 0 0 0 0

2 181 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.077 0 0 0

2 184 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.192 0.125 0 0

2 170 0 0 0 0 0.063 0 0 0 0 0 0 0 0 0 0 0 0.038 0 0 0

2 178 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.083 0

2 180 0 0 0 0 0 0 0 0 0 0 0 0 0.143 0 0 0 0.038 0 0 0

2 400 0.042 0.778 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 406 0.083 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 120 0.083 0 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 123 0.292 0.389 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 141 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.077 0 0 0

2 144 0 0 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0.346 0 0 0

2 150 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.038 0.063 0 0

2 153 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.167 0

2 171 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.038 0.188 0 0

2 141 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.111 0 0 0 0

2 165 0 0 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0.028 0 0 0 0

2 169 0 0.056 0 0 0 0.136 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 108 0.083 0 0 0.132 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 132 0 0 0 0 0 0 0.017 0 0 0 0 0 0 0.033 0 0 0 0 0 0

1 132 0 0 0.075 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 135 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.056

1 136 0 0 0.025 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 138 0 0 0 0.026 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 140 0 0 0.225 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 160 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.313 0 0

1 161 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.167 0

1 166 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 168 0 0 0 0.026 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 179 0 0 0 0 0.033 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 180 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 181 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 182 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 183 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 185 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.038 0 0 0

1 188 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.038 0 0 0

1 192 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.038 0 0 0

1 196 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.038 0 0 0

1 195 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.5 0 0

1 199 0 0 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 205 0 0 0 0 0 0 0 0.111 0 0 0 0 0 0 0 0 0 0 0 0

1 231 0 0 0 0 0 0.045 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 235 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 237 0 0.111 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 239 0 0.167 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 241 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 243 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 257 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.115 0 0 0

1 271 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.083 0

1 277 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 119 0.167 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 121 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 135 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 137 0 0 0 0 0.063 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 147 0 0 0 0 0.063 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 149 0 0 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 151 0 0 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 153 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.056

1 99 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 101 0.125 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 147 0 0 0 0.053 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 151 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.042 0 0 0 0 0

1 171 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 192 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.028 0 0 0 0

1 195 0 0 0 0 0 0 0 0 0 0 0 0.056 0 0 0 0 0 0 0 0

1 207 0 0.167 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 209 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.115 0 0 0

1 214 0 0 0 0.053 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 227 0 0 0 0.053 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 298 0.125 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 127 0 0 0 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 145 0 0 0 0 0.063 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 152 0 0 0 0 0 0 0 0 0 0 0.05 0 0 0 0 0 0 0 0 0

1 161 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.028 0 0 0 0

1 187 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.115 0 0 0

1 190 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.038 0 0 0

1 168 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 182 0 0 0 0 0 0 0 0 0 0 0 0 0.286 0 0 0 0 0 0 0

1 202 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.056

1 390 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 392 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 394 0.125 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 398 0.167 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 404 0 0.111 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 410 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 126 0 0 0 0 0 0 0 0 0 0 0 0 0.018 0 0 0 0 0 0 0

1 165 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 168 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 172 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.056

1 174 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.063 0 0

1 135 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.25 0

1 167 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 171 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 173 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 106 0 0.111 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 144 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 146 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

Distance from Athens 9.03 18.18 20.50 18.69 17.54 22.53 19.21 9.87 17.20 18.92 18.68 19.25 19.12 19.10 19.11 22.29 21.95 20.35 18.91 19.06S 103 95 69 64 63 61 58 56 55 51 49 49 48 46 44 43 42 33 32 30

S Locus 32 19 22 13 16 25 5 33 17 11 12 2 1 29 4 23 24 28 15 1019 120 0.083 0.125 0.038 0.367 0.184 0 0.167 0.056 0.5 0.25 0.364 0.417 0.339 0.1 0.5 0.083 0.063 0.028 0.611 0.25

18 127 0.417 0.063 0.885 0.75 0.211 0.333 0.733 0 0.175 1 0.409 0.917 1 0.6 0.833 1 0.75 0 0.5 1

18 132 0.167 0.563 0 0.65 0.684 0.722 0.833 0.222 0.65 1 0.818 0.958 0.143 0.45 0.833 0 0.125 1 1 0.6

18 118 0.042 0.063 0.077 0.367 0.158 0.111 0.2 0.167 0.125 0.111 0.227 0 0.482 0.25 0 0.333 0.313 0.667 0.389 0.1

17 153 0.042 0.188 0.692 0.1 0.184 0.333 0 0.111 0.125 0.056 0.273 0.167 0.089 0.05 0.056 0.167 0.063 0 0 0.05

16 154 0.125 0.438 0 0.783 0.5 0.111 0.7 0 0.875 0.806 0.409 0.958 0.982 0.35 0.722 0 0 0.972 0.111 0.7

16 151 0.125 0.125 0.231 0.35 0.132 0.389 0.233 0.056 0.1 0.111 0.045 0.042 0 0 0 0.5 0.5 0.444 0 0.1

16 155 0.083 0.281 0 0.067 0.211 0.056 0.4 0.056 0.5 0.167 0.136 0.375 0.357 0.3 0.333 0 0.125 0 0 0.7

15 160 0.083 0.188 0 0.183 0.368 0.5 0.3 0 0.125 0.083 0.591 0.042 0.018 0.55 0.278 0 0 0 0.889 0.3

14 205 0 0.281 0.115 0.4 0.184 0.111 0.5 0 0.375 0.139 0.364 0.125 0.482 0 0.111 0.333 0 0 0.222 0

14 208 0 0 0.077 0.133 0.184 0 0.1 0 0.175 0.694 0.227 0.333 0.482 0 0.389 0.083 0.125 0.056 0.056 0

14 190 0 0.188 0.077 0.333 0.263 0.167 0.467 0 0.6 0.139 0.364 0.292 0.286 0 0.556 0.333 0.25 0 0 0

14 192 0 0.063 0.115 0.183 0.263 0.222 0.033 0 0.075 0.333 0.045 0.083 0.036 0 0.167 0 0 0 0.167 0.4

13 219 0.083 0.094 0 0.2 0 0.056 0.2 0.111 0 0.306 0.227 0.5 0.482 0.5 0.222 0 0 0.278 0 0

13 221 0.125 0.125 0 0.1 0.079 0.167 0.167 0 0.225 0.25 0.091 0.083 0 0.1 0.389 0 0 0 0.778 0

13 188 0.167 0.313 0.038 0.033 0.105 0 0 0 0.05 0 0.182 0.417 0.036 0.7 0 0.083 0 0.833 0.444 0

13 135 0.042 0.188 0 0.35 0.316 0.111 0.167 0 0.35 0 0.182 0.042 0.839 0.55 0.167 0 0 0 0 0.4

13 157 0 0.125 0 0.267 0.237 0 0.233 0.056 0.15 0.333 0.045 0.042 0.036 0.45 0.556 0 0 0 0.444 0

12 133 0 0.063 0.038 0.017 0 0 0.067 0.056 0 0.028 0 0.083 0.054 0 0.056 0.083 0 0.278 0 0.05

12 197 0 0.031 0.346 0.067 0 0.611 0.133 0 0 0 0.273 0.083 0 0 0.333 0.5 0.875 0 0.278 1

12 116 0.042 0.25 0.269 0 0.184 0.167 0.033 0.167 0.225 0.028 0 0 0.018 0.05 0 0 0.188 0 0 0

11 131 0 0.344 0 0.25 0.263 0.167 0.233 0 0.7 0 0.409 0.083 0 0 0.167 0 0.063 0 0.5 0

11 109 0 0.156 0.154 0.3 0.079 0.056 0.1 0 0.25 0.333 0.273 0 0 0 0.056 0 0 0 0.222 0

11 137 0 0.094 0 0.083 0 0.056 0.167 0 0 0.139 0 0.083 0.018 0 0.222 0.25 0 0.389 0.056 0

11 193 0.083 0.25 0.077 0.017 0 0.111 0.033 0 0 0 0 0 0 0.05 0.056 0.083 0 0.056 0.056 0

11 194 0 0.125 0.115 0.017 0.184 0.167 0 0.056 0.075 0 0.045 0 0 0.1 0 0 0.063 0.111 0 0

11 149 0.333 0.156 0.077 0 0.184 0.111 0 0.389 0 0 0.091 0 0 0.1 0 0 0.25 0.083 0.056 0

11 122 0 0.25 0 0.083 0.263 0.056 0.2 0 0.15 0.444 0.273 0 0 0.1 0.278 0 0 0 0 0.6

10 153 0 0.033 0.038 0 0 0 0.2 0 0 0.028 0 0.417 0.054 0.15 0.167 0 0 0 0.444 0.1

10 215 0.083 0 0 0.15 0.026 0 0.1 0 0.025 0 0.045 0.125 0 0.05 0.111 0 0 0 0 0.4

10 225 0.083 0.031 0 0.2 0.053 0 0.233 0 0 0.139 0.136 0.208 0 0 0 0 0 0.194 0 0.3

10 125 0.083 0.063 0.115 0 0.237 0 0.033 0.667 0 0 0.136 0 0 0.4 0 0 0.125 1 0 0

10 196 0.083 0.063 0.231 0.267 0 0 0.033 0.833 0.025 0.167 0 0 0 0.25 0 0 0 0 0.056 0

10 186 0.042 0.031 0 0.35 0 0 0 0 0.15 0.306 0.182 0.042 0 0.15 0 0.083 0 0 0.389 0

9 223 0.042 0.094 0 0 0.053 0.111 0.033 0 0.25 0.028 0 0 0 0 0.111 0 0 0 0 0.1

9 117 0 0.063 0 0.4 0.211 0.056 0 0.167 0.1 0.139 0.455 0 0 0 0 0 0 0 0.389 0

9 131 0.167 0.125 0 0 0 0 0.233 0 0.025 0.167 0 0.208 0.393 0 0 0.083 0 0 0 0.75

9 135 0.125 0.188 0 0.05 0 0.056 0.067 0 0.025 0.111 0 0 0 0.1 0.111 0 0 0 0 0

9 196 0 0.063 0.115 0.017 0.132 0.278 0 0 0 0 0.045 0.167 0.089 0 0 0 0 0 0 0.5

9 198 0 0 0 0 0 0.056 0.2 0 0.05 0.111 0.136 0 0.018 0.05 0.056 0 0 0 0 0.05

9 159 0.208 0 0 0.017 0.053 0 0.1 0 0 0 0 0.125 0.232 0.05 0 0 0 0.306 0.222 0

9 110 0.292 0.063 0.038 0.067 0 0.111 0.133 0.167 0 0.167 0 0 0 0 0 0.083 0 0 0 0

9 114 0.042 0.094 0.192 0 0.079 0 0 0.278 0 0 0 0.083 0 0.25 0.056 0.25 0 0 0 0

9 124 0.167 0.063 0 0.017 0 0 0.033 0 0 0 0.091 0.208 0.054 0 0.056 0 0 0 0 0.05

8 151 0 0.3 0 0 0 0 0.067 0 0.025 0.028 0.045 0 0.25 0.15 0.167 0 0 0 0 0

8 159 0.042 0.033 0.077 0.033 0 0 0.2 0 0 0.111 0 0 0 0 0.167 0 0 0.194 0 0

8 217 0 0.156 0 0 0.184 0.333 0 0 0.15 0 0 0 0.089 0.05 0.056 0 0 0.028 0 0

8 127 0.167 0.063 0 0.017 0 0.056 0 0 0 0 0 0.042 0 0 0.111 0 0 0.167 0 0.15

8 139 0.042 0 0 0 0 0 0.233 0.111 0.025 0.028 0 0.125 0.089 0.15 0 0 0 0 0 0

8 184 0 0.031 0 0.067 0.053 0 0 0 0 0.083 0 0 0.107 0 0 0.083 0 0.056 0 0.05

7 141 0.042 0.033 0 0.067 0.026 0 0 0 0 0.056 0.455 0 0 0 0 0 0.125 0 0 0

7 154 0.125 0 0 0.033 0 0.278 0 0 0 0.167 0 0.25 0.161 0 0 0 0.063 0 0 0

7 155 0.042 0 0 0 0 0 0 0 0 0.194 0.182 0 0 0.45 0.167 0 0 0.583 0 0.8

7 157 0 0.067 0 0 0 0 0.033 0 0 0.028 0.045 0 0.036 0 0.056 0 0 0.222 0 0

7 209 0 0 0 0.017 0.342 0 0 0.389 0.2 0.25 0 0 0.071 0 0 0 0 0 0 0.05

7 213 0 0 0 0.017 0.053 0.111 0.233 0 0.05 0 0 0 0 0.1 0 0 0 0.5 0 0

7 115 0.083 0 0.038 0 0.395 0 0 0 0.525 0.056 0.182 0 0 0.55 0 0 0 0 0 0

7 141 0 0 0 0 0 0.056 0.1 0.111 0 0 0.091 0.167 0.25 0.05 0 0 0 0 0 0

7 159 0.083 0.031 0.154 0 0 0.056 0 0.111 0 0 0 0 0 0 0 0.5 0.25 0 0 0

7 161 0 0 0 0 0 0.111 0.033 0.056 0.05 0.139 0.182 0 0.286 0 0 0 0 0 0 0

6 149 0 0.067 0 0 0 0 0 0 0.025 0.222 0.091 0.083 0 0 0 0 0 0 0.056 0

6 156 0.042 0 0 0.067 0 0 0 0 0 0 0.136 0 0.018 0 0.278 0 0 0 0.222 0

6 201 0.125 0 0 0.05 0.158 0 0.033 0 0.075 0 0 0 0.161 0 0 0 0 0 0 0

6 211 0 0 0 0.15 0.053 0 0 0.111 0.025 0 0 0.083 0 0.1 0 0 0 0 0 0

6 111 0.042 0 0.346 0 0 0.556 0 0.278 0 0 0 0 0 0 0 0.583 1 0 0 0

6 129 0 0 0 0.05 0 0.056 0.033 0 0 0 0 0 0.071 0 0.222 0 0 0 0 0.05

6 194 0.25 0 0 0.117 0 0 0.067 0 0 0 0 0.375 0 0 0.056 0 0 0 0.333 0

6 166 0.083 0.156 0.038 0.017 0 0.111 0 0 0 0 0 0 0 0 0 0.5 0 0 0 0

6 175 0.042 0 0.115 0 0 0.167 0 0.111 0 0 0 0 0 0 0 0.083 0.438 0 0 0

6 174 0.042 0.094 0.231 0 0 0.056 0 0 0 0 0 0 0 0 0 0.083 0.563 0 0 0

6 147 0 0 0 0.1 0 0 0 0 0.075 0 0.091 0 0 0.05 0 0 0.063 0.028 0 0

6 126 0 0 0 0.083 0 0.111 0.167 0 0 0 0 0 0.018 0.05 0 0 0 0.306 0 0

5 145 0.042 0 0 0.05 0.263 0 0 0 0.025 0 0 0 0 0 0 0 0.063 0 0 0

5 148 0 0 0 0.383 0 0.111 0 0 0.025 0 0 0 0.071 0.25 0 0 0 0 0 0

5 152 0 0 0 0 0 0 0.033 0 0 0.028 0.045 0 0.196 0 0 0 0.125 0 0 0

5 158 0 0.033 0 0 0 0.111 0.067 0 0 0.083 0 0 0.214 0 0 0 0 0 0 0

5 164 0.125 0 0 0.033 0.026 0 0.167 0.167 0 0 0 0 0 0 0 0 0 0 0 0

5 165 0 0.133 0 0.233 0 0.222 0 0 0.15 0 0 0 0 0 0 0 0 0 0.278 0

5 203 0 0.156 0 0 0 0 0 0 0 0 0.455 0 0.196 0.1 0 0 0.5 0 0 0

5 129 0.167 0.156 0 0 0.211 0 0 0 0 0 0.045 0 0 0 0 0 0.063 0 0 0

5 107 0.167 0.156 0 0 0.132 0 0 0 0 0 0 0 0 0 0.222 0 0 0.167 0 0

5 121 0 0 0.038 0 0.079 0.056 0 0 0 0 0 0.042 0 0 0 0 0 0 0.167 0

5 200 0 0.031 0 0 0 0 0.3 0 0 0.028 0 0 0 0 0.222 0 0.125 0 0 0

5 156 0.125 0.094 0.115 0 0 0 0 0.167 0 0 0 0 0 0 0 0 0.25 0 0 0

5 163 0.125 0.063 0 0 0 0 0 0.056 0 0.194 0 0 0 0 0.056 0 0 0 0 0

5 112 0.083 0.063 0 0 0 0.056 0 0.056 0 0 0 0 0 0.2 0 0 0 0 0 0

4 150 0 0 0 0.083 0 0.222 0.067 0 0 0 0 0 0 0 0 0 0 0 0 0.1

4 167 0 0.1 0.077 0 0 0 0 0.333 0 0 0 0 0 0 0 0 0.25 0 0 0

4 173 0.042 0.033 0.154 0 0 0 0 0 0 0 0 0 0 0 0 0 0.063 0 0 0

4 177 0.083 0 0.038 0 0 0 0 0.111 0 0 0 0 0 0 0 0.083 0 0 0 0

4 227 0 0 0 0.117 0 0 0 0.111 0 0.028 0 0 0 0 0.111 0 0 0 0 0

4 211 0 0.063 0 0 0.053 0 0.1 0 0 0 0 0.042 0 0 0 0 0 0 0 0

4 130 0.542 0.031 0 0 0 0 0 0.778 0 0 0 0 0 0.05 0 0 0 0 0 0

4 163 0 0 0 0 0.026 0.056 0 0 0 0.056 0 0 0 0 0 0.167 0 0 0 0

4 169 0 0 0.077 0 0 0.056 0 0.056 0 0 0 0 0 0 0 0 0.063 0 0 0

4 172 0.042 0 0.115 0 0 0 0 0 0 0 0 0 0 0 0 0.167 0.125 0 0 0

4 178 0 0.031 0.231 0 0 0 0 0 0 0 0 0 0 0 0 0.083 0.25 0 0 0

4 162 0 0.031 0.192 0 0 0 0 0.056 0 0 0 0 0 0 0 0.333 0 0 0 0

4 143 0 0.031 0 0.017 0 0 0 0 0 0 0 0.25 0 0 0 0.083 0 0 0 0

4 100 0 0 0.154 0 0 0.389 0 0 0 0 0 0 0 0 0 0.167 0.25 0 0 0

4 128 0 0 0 0 0 0 0 0 0 0 0 0.25 0.089 0 0.111 0.083 0 0 0 0

4 130 0 0.031 0 0 0 0 0.033 0 0 0 0.045 0.042 0 0 0 0 0 0 0 0

3 142 0 0.033 0 0 0.158 0 0 0 0.3 0 0 0 0 0 0 0 0 0 0 0

3 143 0 0.033 0 0 0.132 0 0 0 0.025 0 0 0 0 0 0 0 0 0 0 0

3 144 0 0 0 0 0.132 0 0 0 0.1 0 0 0.167 0 0 0 0 0 0 0 0

3 162 0 0 0.038 0 0 0 0 0 0 0.056 0 0.083 0 0 0 0 0 0 0 0

3 174 0.125 0.033 0 0 0 0 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0

3 207 0 0.281 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.111 0.15

3 261 0.042 0 0.038 0 0 0.222 0 0 0 0 0 0 0 0 0 0 0 0 0 0

3 267 0.083 0 0.115 0 0 0 0 0 0 0 0 0 0 0 0 0.25 0 0 0 0

3 143 0 0.031 0 0 0 0.167 0 0 0.025 0 0 0 0 0 0 0 0 0 0 0

3 145 0 0.094 0 0 0 0.278 0 0 0.025 0 0 0 0 0 0 0 0 0 0 0

3 113 0 0 0.385 0 0 0 0 0.278 0.05 0 0 0 0 0 0 0 0 0 0 0

3 119 0 0.063 0 0 0.053 0 0 0 0 0 0 0 0 0 0 0 0 0 0.167 0

3 143 0 0 0 0.067 0 0 0 0 0 0 0 0.042 0.125 0 0 0 0 0 0 0

3 191 0.083 0.125 0 0 0 0 0 0 0 0 0 0 0.036 0 0 0 0 0 0 0

3 199 0 0 0 0 0.447 0.111 0 0 0.425 0 0 0 0 0 0 0 0 0 0 0

3 204 0.125 0 0 0 0 0 0 0 0 0 0 0 0 0.6 0 0 0 0.389 0 0

3 176 0.083 0 0.231 0 0 0 0 0 0 0 0 0 0 0 0 0.25 0 0 0 0

3 138 0.042 0 0.038 0 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0

3 147 0.042 0.031 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.063 0 0 0

3 145 0 0 0 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.278 0.15

3 102 0.042 0 0.231 0 0 0 0 0 0 0 0 0 0 0 0 0 0.188 0 0 0

2 139 0 0.033 0 0 0.053 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 146 0 0 0 0.017 0 0 0.033 0 0 0 0 0 0 0 0 0 0 0 0 0

2 147 0 0 0 0 0.132 0 0.133 0 0 0 0 0 0 0 0 0 0 0 0 0

2 163 0 0 0 0 0.026 0 0 0 0 0 0 0 0 0 0 0.333 0 0 0 0

2 170 0 0 0.269 0 0 0 0 0.111 0 0 0 0 0 0 0 0 0 0 0 0

2 171 0 0 0.077 0 0 0 0 0 0 0 0 0 0 0 0 0.083 0 0 0 0

2 172 0.042 0 0 0 0 0 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0

2 175 0 0 0.077 0 0 0 0 0 0 0 0 0 0 0 0 0.333 0 0 0 0

2 176 0.083 0 0 0 0 0 0 0.111 0 0 0 0 0 0 0 0 0 0 0 0

2 229 0.083 0.031 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 263 0 0 0.462 0 0 0 0 0 0 0 0 0 0 0 0 0.333 0 0 0 0

2 265 0 0 0.269 0 0 0 0 0 0 0 0 0 0 0 0 0.333 0 0 0 0

2 133 0 0 0 0 0.079 0 0 0.333 0 0 0 0 0 0 0 0 0 0 0 0

2 141 0 0.063 0 0 0 0 0 0 0.075 0 0 0 0 0 0 0 0 0 0 0

2 123 0 0 0 0 0 0 0 0 0 0 0 0.125 0 0.05 0 0 0 0 0 0

2 125 0 0.031 0 0 0 0 0 0 0 0 0 0 0 0.1 0 0 0 0 0 0

2 145 0 0 0 0.017 0 0 0 0 0 0 0 0.042 0 0 0 0 0 0 0 0

2 190 0 0.188 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.139 0 0

2 198 0.083 0 0 0 0 0 0 0 0 0 0 0.042 0 0 0 0 0 0 0 0

2 200 0 0 0.038 0 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 201 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0.1 0 0 0 0 0 0

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2 151 0 0.063 0 0.017 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

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1 126 0 0 0 0 0 0 0 0 0 0 0 0 0.018 0 0 0 0 0 0 0

1 165 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 168 0 0 0 0 0 0 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0

1 172 0 0 0 0 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 174 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.063 0 0 0

1 135 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0.25 0 0 0 0

1 167 0 0 0 0 0 0 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0

1 171 0 0 0 0 0 0 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0

1 173 0 0 0 0 0 0 0 0.056 0 0 0 0 0 0 0 0 0 0 0 0

1 106 0 0 0 0 0 0 0 0.111 0 0 0 0 0 0 0 0 0 0 0 0

1 144 0.083 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 146 0.042 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

The map of 144 alleles from 10 loci spread across 20 sites from southern and western Europe

Sites sorted

according to allele richness

Sites sorted according to distance from Athens

The mean number of elleles

per locus decreases with distance from

Greece

Colonisation gradient

Founder effectsOrdered genetic loss

Relict populationsVicariant (scattered) genetic loss

Popu-lations

SiteAlleles 1 2 3 4 5 6 7 8 9 10

1 1 1 1 1 1 1 1 1 1 12 1 1 1 1 1 1 1 1 0 03 1 1 1 1 1 1 0 0 0 04 1 1 1 1 1 0 0 0 0 05 1 1 1 1 0 0 0 0 0 06 1 1 1 0 0 0 0 0 0 07 1 0 0 0 0 0 0 0 0 08 1 0 0 0 0 0 0 0 0 0

A perfectly nested pattern of allele loss

SiteAlleles 1 2 3 4 5 6 7 8 9 10

1 1 1 1 1 1 1 1 1 1 12 1 1 0 1 1 1 1 1 0 03 1 1 1 1 1 1 0 0 0 04 0 1 1 1 1 0 0 0 0 05 1 1 1 1 0 0 0 0 0 06 1 1 1 0 0 0 0 1 0 07 1 0 0 0 0 0 0 0 1 08 1 0 0 0 0 0 0 0 0 0

Unexpected absences

Unexpected presences

Rows and columns are sorted according to species richness

SiteAlleles 1 2 3 4 5 6 7 8 9 10

1 1 1 1 0 1 0 0 0 0 02 1 1 1 1 0 0 0 0 0 03 0 1 1 1 1 1 0 0 0 04 0 0 1 1 1 1 0 0 0 05 0 0 0 1 1 1 1 1 0 06 0 0 0 0 1 1 1 1 1 07 0 0 0 0 0 0 1 1 1 18 0 0 0 0 0 0 0 1 1 1

A matrix sorted to maximise the number of occurrences along the diagonale reveals the degree of allele turnover in the population

SiteAlleles 1 2 3 4 5 6 7 8 9 10

1 1 0 1 0 1 0 0 1 0 12 1 1 0 1 0 0 0 0 1 03 0 1 0 0 1 1 0 0 0 04 0 0 1 1 1 1 0 0 1 05 0 0 1 0 1 0 1 1 0 06 0 1 0 0 1 1 0 1 0 07 1 0 0 1 0 0 1 1 0 18 0 1 0 0 0 0 0 1 1 1

A random matrix

A nested pattern of allele loss with unexpected absences and presences

Italy W-Balkans E France S Luxemburg C Luxemburg N-Germany N Luxemburg

S-Belgiu

mW France

Distance

from Athens

9.03 9.87 17.20 17.54 18.18 18.68 18.69 18.91 18.92 19.06 19.10 19.11 19.12 19.21 19.25 20.35 20.50 21.95 22.29 22.53

N Alleles 103 56 55 63 95 49 64 32 51 30 46 44 48 58 49 33 69 42 43 61

Alleles Locus 32 33 17 16 19 12 13 15 11 10 29 4 1 5 2 28 22 24 23 25

19 120 0.083

0.056 0.5 0.184 0.125 0.364 0.367 0.611 0.25 0.25 0.1 0.5 0.339 0.167 0.417 0.028 0.038 0.063 0.083 0

18 127 0.417 0 0.175 0.211 0.063 0.409 0.75 0.5 1 1 0.6 0.833 1 0.733 0.917 0 0.885 0.75 1 0.333

18 132 0.167

0.222 0.65 0.684 0.563 0.818 0.65 1 1 0.6 0.45 0.833 0.143 0.833 0.958 1 0 0.125 0 0.722

18 118 0.042

0.167 0.125 0.158 0.063 0.227 0.367 0.389 0.111 0.1 0.25 0 0.482 0.2 0 0.667 0.077 0.313 0.333 0.111

17 153 0.042

0.111 0.125 0.184 0.188 0.273 0.1 0 0.056 0.05 0.05 0.056 0.089 0 0.167 0 0.692 0.063 0.167 0.333

Eastern limeage with ordered loss of alleles along the colonisation gradient

Western limeage with ordered loss

of alleles along the colonisation

gradient

The ordered (nested) loss of alleles along putative cpolonisation gradients

Intermediate situation

S 32 33 16 28 29 17 19 12 15 11 13 4 5 10 2 1 25 24 22 23181 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0298 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

171 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

99 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

121 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

277 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

392 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

144 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

135 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

235 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

119 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

183 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

182 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

165 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

390 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

243 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

180 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

101 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

394 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

241 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

168 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

398 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

146 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

410 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

400 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

406 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

123 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

172 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

176 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

166 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

108 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

173 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

404 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

106 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

167 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

168 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

148 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

239 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

237 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

207 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

171 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

141 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

133 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

147 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

201 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

227 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

214 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

168 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

138 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

204 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

174 1 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

210 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

130 1 1 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

120 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

229 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

192 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

161 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

139 0 0 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

169 0 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0

152 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

143 0 0 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0

142 0 0 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0

107 1 0 1 1 0 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0

165 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

190 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

157 1 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0

132 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0

140 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0

136 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0

164 1 1 1 0 0 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0

125 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

119 0 0 1 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0

112 1 1 0 0 1 0 1 0 0 0 0 0 0 0 0 0 1 0 0 0

163 1 1 0 0 0 0 1 0 0 1 0 1 0 0 0 0 0 0 0 0

141 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0

151 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

127 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

147 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

199 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

145 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

149 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

179 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

137 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0

211 0 1 1 0 1 1 0 0 0 0 1 0 0 0 1 0 0 0 0 0

129 1 0 1 0 0 0 1 1 0 0 0 0 0 0 0 0 0 1 0 0

115 1 0 1 0 1 1 0 1 0 1 0 0 0 0 0 0 0 0 1 0

155 1 0 0 1 1 0 0 1 0 1 0 1 0 1 0 0 0 0 0 0

145 1 0 1 0 0 1 0 0 0 0 1 0 0 0 0 0 0 1 0 0

125 1 1 1 1 1 0 1 1 0 0 0 0 1 0 0 0 0 1 1 0

191 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0 0 0 0

198 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0

147 0 0 1 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0

231 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0

144 0 0 1 0 0 1 0 0 0 0 0 0 0 0 1 0 0 0 0 0

117 0 1 1 0 0 1 1 1 1 1 1 0 0 0 0 0 1 0 0 0

141 1 0 1 0 0 0 1 1 0 1 1 0 0 0 0 0 0 1 0 0

196 1 1 0 0 1 1 1 0 1 1 1 0 1 0 0 0 0 0 1 0

201 1 0 1 0 0 1 0 0 0 0 1 0 1 0 0 1 0 0 0 0

213 0 0 1 1 1 1 0 0 0 0 1 0 1 0 0 0 1 0 0 0

149 1 1 1 1 1 0 1 1 1 0 0 0 0 0 0 0 1 1 1 0

139 1 1 0 0 1 1 0 0 0 1 0 0 1 0 1 1 0 0 0 0

159 1 0 1 1 1 0 0 0 1 0 1 0 1 0 1 1 0 0 0 0

225 1 0 1 1 0 0 1 1 0 1 1 0 1 1 1 0 0 0 0 0

147 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0

147 0 0 0 1 1 1 0 1 0 0 1 0 0 0 0 0 0 1 0 0

199 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0

217 0 0 1 1 1 1 1 0 0 0 0 1 0 0 0 1 1 0 0 0

227 0 1 0 0 0 0 0 0 0 1 1 1 0 0 0 0 0 0 0 0

215 1 0 1 0 1 1 0 1 0 0 1 1 1 1 1 0 0 0 0 0

209 0 1 1 0 0 1 0 0 0 1 1 0 0 1 0 1 0 0 0 0

151 0 0 0 0 0 0 1 0 0 0 1 0 0 0 0 0 0 0 0 0

205 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0

221 1 0 1 0 1 1 1 1 1 1 1 1 1 0 1 0 1 0 0 0

157 0 1 1 0 1 1 1 1 1 1 1 1 1 0 1 1 0 0 0 0

113 0 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0

194 0 1 1 1 1 1 1 1 0 0 1 0 0 0 0 0 1 1 1 0

135 1 0 0 0 1 1 1 0 0 1 1 1 1 0 0 0 1 0 0 0

149 0 0 0 0 0 1 1 1 1 1 0 0 0 0 1 0 0 0 0 0

186 1 0 0 0 1 1 1 1 1 1 1 0 0 0 1 0 0 0 0 1

223 1 0 1 0 0 1 1 0 0 1 0 1 1 1 0 0 1 0 0 0

219 1 1 0 1 1 0 1 1 0 1 1 1 1 0 1 1 1 0 0 0

114 1 1 1 0 1 0 1 0 0 0 0 1 0 0 1 0 0 0 1 1

156 1 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 1 0

154 1 0 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 0 0 0

156 1 0 0 0 0 0 0 1 1 0 1 1 0 0 0 1 0 0 0 0

132 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 0 0

211 0 0 1 0 0 0 1 0 0 0 0 0 1 0 1 0 0 0 0 0

188 1 0 1 1 1 1 1 1 1 0 1 0 0 0 1 1 0 0 1 1

151 0 0 0 0 1 1 1 1 0 1 0 1 1 0 0 1 0 0 0 0

159 1 0 0 1 0 0 1 0 0 1 1 1 1 0 0 0 0 0 1 0

122 0 0 1 0 1 1 1 1 0 1 1 1 1 1 0 0 1 0 0 0

116 1 1 1 0 1 1 1 0 0 1 0 0 1 0 0 1 1 1 1 0

160 1 0 1 0 1 1 1 1 1 1 1 1 1 1 1 1 1 0 0 0

135 1 0 1 0 1 1 1 1 0 0 1 1 1 1 1 1 1 0 0 0

155 1 1 1 0 1 1 1 1 0 1 1 1 1 1 1 1 1 1 0 0

143 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 1 0 0 0

145 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 1 0 0 0

157 0 0 0 1 0 0 1 1 0 1 0 1 1 0 0 1 0 0 0 0

207 0 0 0 0 0 0 1 0 1 0 0 0 0 1 0 0 0 0 0 0

123 0 0 0 0 1 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0

141 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0

165 0 0 0 0 0 1 1 0 1 0 1 0 0 0 0 0 1 0 0 0

127 1 0 0 1 0 0 1 0 0 0 1 1 0 1 1 0 1 0 0 0

120 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 0 1 1 1

118 1 1 1 1 1 1 1 1 1 1 1 0 1 1 0 1 1 1 1 1

194 1 0 0 0 0 0 0 0 1 0 1 1 1 0 1 0 0 0 0 0

161 0 1 0 0 0 1 0 1 0 1 0 0 1 0 0 1 1 0 0 0

109 0 0 1 0 0 1 1 1 1 1 1 1 1 0 0 0 1 0 1 0

151 1 1 1 1 0 1 1 1 0 1 1 0 1 1 1 0 1 1 1 1

170 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0

181 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0

178 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1

153 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1

177 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1

202 0 0 0 0 0 0 0 1 0 0 0 0 1 0 0 0 0 0 0 0

184 0 0 1 1 0 0 1 0 0 1 1 0 0 1 0 1 0 0 0 1

193 1 0 0 1 1 0 1 0 1 0 1 1 1 0 0 0 1 0 1 1

130 0 0 0 0 0 0 1 1 0 0 0 0 1 0 1 0 0 0 0 0

124 1 0 0 0 0 0 1 1 0 0 1 1 1 1 1 1 0 0 0 0

203 0 0 0 0 1 0 1 1 0 0 0 0 0 0 0 1 0 1 0 0

131 0 0 1 0 0 1 1 1 1 0 1 1 1 0 1 0 1 1 0 0

153 1 1 1 0 1 1 1 1 0 1 1 1 0 1 1 1 1 1 1 1

141 0 1 0 0 1 0 0 1 0 0 0 0 1 0 1 1 1 0 0 0

126 0 0 0 1 1 0 0 0 0 0 1 0 1 0 0 1 1 0 0 0

148 0 0 0 0 1 1 0 0 0 0 1 0 0 0 0 1 1 0 0 0

110 1 1 0 0 0 0 1 0 0 1 1 0 1 0 0 0 1 0 1 1

198 0 0 0 0 1 1 0 1 0 1 0 1 1 1 0 1 1 0 0 0

173 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 1 0

145 0 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 0 0 0 0

127 1 0 1 0 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1

131 1 0 0 0 0 1 1 0 0 1 0 0 1 1 1 1 0 0 0 1

192 0 0 1 0 0 1 1 1 1 1 1 1 1 1 1 1 1 0 1 0

167 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 1 0

163 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1

208 0 0 1 1 0 1 0 1 1 1 1 1 1 0 1 1 0 1 1 1

205 0 0 1 0 0 1 1 1 1 1 1 1 1 0 1 1 1 0 1 1

133 0 1 0 1 0 0 1 0 0 1 1 1 1 1 1 1 0 0 1 1

162 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1

200 0 0 0 0 0 0 1 0 0 1 0 1 1 0 0 0 0 1 0 0

159 1 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 1 1 1 1

153 0 0 0 0 1 0 1 0 1 1 0 1 1 1 1 1 0 0 1 0

195 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0

132 0 0 0 0 0 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0

146 0 0 0 0 0 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0

137 0 0 0 1 0 0 1 0 1 1 1 1 1 0 1 1 1 0 0 1

196 0 0 1 0 0 0 1 1 0 0 1 0 0 1 1 1 1 0 1 0

261 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0

138 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0

166 1 0 0 0 0 0 1 0 0 0 1 0 0 0 0 0 1 0 1 1

163 0 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 0 0 1

190 0 0 1 0 0 1 1 1 0 1 1 1 1 0 1 1 1 1 1 1

154 1 0 0 0 0 0 0 0 0 1 1 0 0 0 1 1 1 1 0 0

158 0 0 0 0 0 0 1 0 0 1 0 0 1 0 0 1 1 0 0 0

121 0 0 1 0 0 0 0 0 1 0 0 0 0 0 1 0 1 0 1 0

102 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0

111 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 1 1

175 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 1 1

170 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0

144 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0

145 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 0 0 0 0 0

152 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 1 0 1 0 0

143 0 0 0 0 0 0 1 0 0 0 1 0 0 0 1 0 0 0 0 1

267 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1

176 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1

197 0 0 0 0 0 0 1 1 1 0 1 1 1 1 1 0 1 1 1 1

174 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 1 1 1 1

150 0 0 0 0 0 0 0 0 0 0 1 0 1 1 0 0 1 0 0 0

129 0 0 0 0 0 0 0 0 0 0 1 1 1 1 0 1 1 0 0 0

143 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1 0 0 0 0

169 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 1 0

172 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 1

162 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 0 0 0 1 0

151 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0

128 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1 0 0 0 1

178 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 1 1

182 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0

126 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0

135 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0

153 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0

202 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0

172 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0

180 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 0

200 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0

160 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0

195 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0

174 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0

100 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 1 1

184 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0

150 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0

171 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0

185 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0

188 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0

192 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0

196 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0

257 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0

209 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0

187 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0

190 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0

171 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1

175 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1

263 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1

265 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1

161 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1

271 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1

135 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1

Balkans and Italy

Western France

Eastern and western clades do not completely overlap in allele distribution.

Apparently we have two postglacial colonisation trajectories: from Spain and from the Balkans.

Seriation : sorting the matrix according to reciprocal averaging.Occurrences are maximised along the matrix diagonal.

S AlpsC BalkansE France

W FranceMarche

VenetianRomagnaTuscany

Pleistocene Holocene

1.2 my

Languedoc

0.4 my

Calabria

Eastern Clade

Italian Clade

Western Clade