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389Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Manis palaeojavanica +Manis javanica + +Ursus malayanus +Paradoxurus hermaphroditus + +Arctogalidia sp. + +Panthera sp. +?Panthera tigris subsp. + + + + + +Prionailurus (=Felis) bengalensis + + +Aonix (=Amblonix) cinerea + +Lutrogale palaeoleptonyx +Martes flavigula + +Hyaena brevirostris +Mececyon trinilensis +Cuon? javanicus +Cuon javanicus (=alpinus) +Sinomastodon bumiajuensis +Stegodon cf. elephantoides +Stegodon trigonocephalus + + + +Stegodon? hypsilophus +?Elephas hysudrindicus + +Elephas maximus + +Rhinoceros sondaicus + + + + +Rhinoceros unicornis kendengindicus +Tapirus indicus + + + +Hexaprotodon simplex +Hexaprotodon sivalensis + + +Muntiacus muntjak + + + + + +Cervids + + + +Tragulus javanicus + +Axis lydekkeri + + +Axis kuhli +Rusa sp. + +Rusa (=Cervus) timorensis + +Duboisia santeng + +Capricornis sumatraensis +Epileptobos groeneveldtii +Bubalus palaeokerabau + + +Bubalus bubalus (= arnee) + +Bibos palaesondaicus + + +Bibos sp. +
Sati
r
Buk
uran
sit
e,be
low
tuff
9
Ci S
aat
Tri
nil
Ked
ung
Bru
bus
Nga
ndon
g
Punu
ng
Hol
ocen
e ca
ves
com
posi
te
Rec
ent
13. Tables 1-5, 38-42 (other tables on disc)
Table 1. Faunal lists of the fossil vertebrate localities from Java discussed in this paper. The localitiesare placed in chronological order from old (left) to young (right) (based on de Vos et al., 1982, Son-daar, 1984, Cranbook, 1987, and own data).
pp 391-421 15-01-2007 10:57 Pagina 389
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)390
Fossil species lacking stratigraphical information: Hystrix giganteaHemimachairodus zwierzyckiiMeganthereon sp.Homotherium ultimumPanthera pardusLutrogale robustaMegacyon sp.Nestoritherium cf. sivalenseMerycopotamus dissimilisLeptobos? problematicus+“Elephas” indonesicus
Bos javanicus (=Bibos sondaicus) + +Bovidae +Sus brachygnathus + +?Sus macrognathus + +Sus scrofa vittatus + + +Sus sp. +Sus verrucosus + +Nycticebus coucang + +Presbytis (=Trachypithecus) cristatus + + +Presbytis comata +Macaca fascicularis + + + +Macaca nemestrina +Hylobates syndactylus +Hylobates moloch +Pongo pygmaeus +Homo erectus +? + + +Homo sapiens + + +Hystrix (=Acanthion) brachyurus + + +Hystrix javanica + +Panthera pardus +Felis viverrina +Herpestes javanicus +Viverricula indica +Mustela lutreolina +Melogale personata +Mydaus javanensis +Lutra lutra +Lutra sumatrana +Arctitis binturong +Prionodon linsang +
Sati
r
Buk
uran
sit
e,be
low
tuff
9
Ci S
aat
Tri
nil
Ked
ung
Bru
bus
Nga
ndon
g
Punu
ng
Hol
ocen
e ca
ves
com
posi
te
Rec
ent
pp 391-421 15-01-2007 10:57 Pagina 390
391Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)T
able
2. P
oint
coun
t res
ults
of 2
2 sa
ndst
one
sam
ples
of t
he W
alan
ae F
m. A
nd T
anru
ng F
m. O
f peb
bly
sand
ston
e sa
mpl
es o
nly
grai
ns s
mal
ler
than
2 m
m h
ave
been
cou
nted
. “-”
ind
icat
es “
not p
rese
nt; “
+”
ind
icat
es “
trac
es p
rese
nt”.
Fel
dsp
ar in
clud
es a
ll K
-Fel
dsp
ar a
nd p
lagi
ocla
se w
itho
ut a
lbit
e tw
inni
ng.
sam
ple
sect
ion/
stra
tigr
aphi
cce
men
t%
gr
ain
sort
ing
volc
anic
pyro
-pl
agio
clas
efe
ldsp
arm
agne
-m
ono
und
ulou
s +
cher
tbi
otit
ebi
ocla
stsi
licic
last
.ca
lcar
eous
gran
ite/
wea
ther
edot
hers
loca
lity
unit
type
cem
ent
size
xene
(alb
ite)
tite
quar
tzpo
lyqu
artz
sed
imen
t.se
dim
ent.
pert
hite
S-49
III
Bur
ecin
g M
b.ca
lcit
e29
coar
sepo
or34
32
11-
-1
++
232
8-
412
S-29
BPu
ncak
oro
Bur
. Mb?
calc
ite
30co
arse
good
392
19
--
11
130
48
-2
1S-
43II
ISa
mao
ling
Mb.
bas
esi
der
ite
30m
ediu
mgo
od36
22
15-
58
34
-17
-2
6-
S-37
XI
Sam
aolin
g M
b.si
der
ite
35ve
ry fi
nego
od20
19
91
2714
13
-5
-7
-3
S-39
XI
Sam
aolin
g M
b. m
idd
lesi
der
ite
31fi
ne-m
ed.
good
273
58
-18
71
15-
2-
85
1S-
46II
ISa
mao
ling
Mb.
top
sid
erit
e35
fine
-med
.go
od57
113
62
44
21
--
-3
43
S-34
ISa
mao
ling
Mb.
top
mic
rite
36
coar
sepo
or53
233
46
11
22
-1
1-
3-
mat
rix
+ c
alci
teS-
30I
Ber
u M
b. b
ase
sid
erit
e30
coar
sego
od68
15+
51
71
3+
--
-+
--
S-41
IVB
eru
Mb.
bas
esi
der
ite
25co
arse
good
2143
18
19-
11
--
--
-5
1S-
42A
IIB
eru
Mb,
Sub
unit
Asi
der
ite
43ve
ry c
oars
epo
or45
84
12
86
4-
-1
16
113
S-42
BII
Ber
u M
b, S
ubun
it B
calc
ite
26ve
ry c
oars
epo
or45
323
-2
--
2-
-3
--
121
S-50
VII
IB
ase?
Ber
u M
b.si
der
ite
49ve
ry c
oars
epo
or48
303
31
6-
--
-4
--
41
+ li
mon
ite
S-70
IXB
eru
Mb.
Sub
unit
B,
sid
erit
e30
pebb
ly s
st.
poor
4433
-3
+1
-1
--
-12
32
1ba
se+
lim
onit
eS-
71IX
idem
sid
erit
e28
pebb
ly s
st.
poor
3842
--
3-
-1
--
-12
31
-+
lim
onit
eS-
17IX
idem
sid
erit
e51
very
coa
rse
poor
5415
211
4-
-2
3-
2-
1-
6+
lim
onit
eS-
23V
Ber
u M
b. S
ubun
it B
,ca
lcit
e38
very
coa
rse
good
4324
12
310
+-
--
2-
210
3M
idd
leS-
24V
idem
sid
erit
e24
very
coa
rse
good
5421
+-
16
-3
1-
1-
39
1S-
6L
enra
ngB
eru
Mb.
sid
erit
e40
fine
-med
ium
good
704
27
+1
22
1-
1-
16
2(F
VL
-8)
S-57
Paro
toB
eru
Mb.
si
der
ite
37fi
nego
od47
201
95
61
2-
--
-2
43
(FV
L-1
4)Su
buni
t B, t
opS-
27B
ulu
Cep
oB
eru
Mb.
si
der
ite
48fi
nego
od28
162
715
129
21
-1
--
61
(FV
L-1
3)Su
buni
t BS-
54T
anru
ngT
anru
ng F
m.
calc
ite
23pe
bbly
sst
poor
4-
25
-10
11-
-17
243
-3
3R
iver
(gra
nule
s)S-
64T
anru
ngT
anru
ng F
m.
calc
ite
25pe
bbly
sst
.po
or16
-7
6-
389
6-
35
8-
+2
Riv
er
pp 391-421 15-01-2007 10:57 Pagina 391
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)392T
able
3.S
hort
des
crip
tion
of m
icro
scop
ic s
ampl
es n
ot in
clud
ed in
Tab
le 2
.
Sam
ple
Sect
ion/
Loc
alit
ySt
rati
grap
hic
Uni
tT
ype
of S
ampl
eSh
ort D
escr
ipti
on
S-55
Tan
rung
Riv
erT
anru
ng F
mV
ery
coar
se c
alca
reou
s sa
ndst
one
Sim
ilar
as s
ampl
e s-
54S-
65T
anru
ng R
iver
Tan
rung
Fm
Cal
care
ous
cong
lom
erat
eSi
mila
r as
sam
ple
s-64
S-12
Lak
ibon
g FV
L-6
Ber
u M
b., S
ubun
it B
Ver
y co
arse
san
dst
one
Con
tain
ing
volc
anic
roc
k fr
agm
ents
, pyr
oxen
e, m
agne
tite
,pl
agio
clas
e, fe
ldsp
ar; s
ider
ite
cem
ent
S-21
Vi
Ber
u M
b, S
ubun
it B
Fine
san
dst
one
Con
tain
ing
volc
. Roc
kfra
gm.,
Pyro
xene
, qua
rtz,
feld
spar
; wit
hsi
der
ite
cem
ent
S-36
IB
eru
Mb,
Sub
unit
AC
lays
tone
Cal
care
ous
clay
ey m
icri
te w
ith
clia
chit
e lu
mps
(bau
xite
)S-
33I
Sam
aolin
g M
b, T
opC
lays
tone
Cal
care
ous
clay
ey m
icri
teS-
60C
iang
kang
e (F
VL
-21)
Col
luvi
umC
alic
he c
oncr
etio
n in
foss
il bo
neB
one:
col
orle
ss c
ollo
phan
e bu
ndle
s; c
alic
he: m
icri
te a
nd
mic
rosp
arit
e lu
mps
. Silt
-siz
ed s
ilici
clas
tic
grai
ns a
nd c
lay
lum
ps w
ith
spar
itic
rim
. Tub
ulou
s ca
viti
es w
ith
spar
ite-
lath
rim
s (r
ootl
ets)
. Ir
regu
lar
limon
ite
crus
ts a
roun
d b
one,
mic
rite
lum
ps a
nd tu
bule
s.Sp
arit
e in
cra
cks
in b
one
and
mat
rix
S-25
Lak
ibon
g (F
VL
-5)
Ber
u M
b., S
ubun
it B
Cal
iche
con
cret
ion
from
foss
ilife
rous
cla
yM
icri
tic
lum
ps w
ith
mic
rosp
arit
e pa
tche
s an
d o
paqu
e he
mat
ite
patc
hes.
Som
e m
icri
te lu
mps
wit
h ad
mix
ture
of c
lay,
oth
ers
part
ly
repl
aced
by
spar
ite
wit
h m
icri
tic
isla
nds.
Cle
ar b
lock
y sp
arit
e in
fill
ofca
viti
es, c
rack
s an
d tu
bule
s, a
nd a
roun
d d
etri
tal s
ilt-s
ized
gra
ins
S-61
A &
BB
ulub
arer
e (F
VL
-28)
Ber
u M
b., B
ase
Foss
ilife
rous
gra
vel w
ith
clay
ey m
icri
teG
rave
l: vo
lcan
ic r
ock
frag
m.,
Che
rt, b
one,
she
lls. M
atri
x: b
row
nish
mat
rix
clay
ey m
icri
te w
ith
sand
and
silt
-siz
ed d
etri
tal g
rain
s. B
iocl
asts
st
rong
ly a
ltera
ted
to m
icri
te. B
one:
col
orle
ss a
pati
te a
ggre
gate
s w
ith
brow
n ad
sorp
tion
rim
sS-
62M
aral
e (F
VL
-19)
Ber
u M
b., S
ubun
it B
Foss
ilize
d W
ood
Cal
cifi
ed w
ood
; ori
gina
l cel
l str
uctu
re p
oorl
y pr
eser
ved
. Cel
l wal
ls
visi
ble
as b
row
nish
ferr
ogen
ous
incl
usio
ns in
larg
e ir
regu
lar
calc
ite
crys
tals
. Eac
h ca
lcit
e cr
ysta
l cov
ers
man
y ce
llsS-
63B
ulu
Cep
o (F
VL
-11)
Ber
u M
b., S
ubun
it A
Foss
ilize
d W
ood
Silic
ifie
d w
ood
; ori
gina
l cel
l str
uctu
re p
erfe
ctly
pre
serv
ed. C
ells
re
plac
ed b
y m
icro
-cry
stal
line
cher
t, ce
ll w
all v
isib
le a
s co
lore
d b
and
sS-
64)
Paro
to (F
VL
-14)
Ber
u M
b., S
ubun
it B
Foss
ilize
d W
ood
Silic
ifie
d w
ood
; ori
gina
l cel
l str
uctu
re p
erfe
ctly
pre
serv
ed. Q
uart
z cr
ysta
l siz
e va
riab
le fr
om m
icro
-cry
stal
line
cher
t to
sing
le c
ryst
als
filli
ng c
ells
. Opa
que
crys
tal a
t cel
l bou
ndar
ies.
Bla
ded
qua
rtz
crys
tals
in c
avit
ies
pp 391-421 15-01-2007 10:57 Pagina 392
393Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Table 4A. Microfossil analysis of samples from marine layers of the Walanae Formation, SengkangAnticline area, South Sulawesi (sections I and III). Stratigraphic position of the samples is indicated inFig. 40. Foraminifera were determined by Sudijono, nannofossils by A. Priadi.
A: Foraminifera (Biozones according to Berggren, 1995)Sample Stratigraphic Unit Contents Biozonation/Age Environment
Planktonoic foraminifera:F-9B: Section III, Globigerina bulloides, PL1-PL2 open sea,
Burecing Member; Ga. Decoraperta, (latest late outernear the base of Ga. Venezuelana, Miocene to early sublittoralexposed sequence Globoquadrina altispira, Pliocene) to upper at the core of the Globorotalia (Turborotalia) acostalensis, bathyalSengkang Anticline Globorotalia (Globorotalia) menardii,
Gt. (Gt.) margaritae,Gt. (Gt.) tumida,Gt. (Gt.) plesiotumida,Globigerinoides extremus, Gds. Conglobatus, Gds. Obliquus, Gds. Immaturus,Gds. Trilobus,Hastigerina aequilateralis,Orbulina universa,Sphaeroidinellopsis seminulina,Sp. sphaeroides,Pulleniatina primalis,Globorotalia flexuosa.
Benthonic foraminifera:Planulina wuellerstorfi,Lenticulina sp.,Gyroidina sp.,Oridorsalis umbonatus,Pleurostomella alternans,Pleurostomella sp.,Bulimina striata,Uvigerina hispida,Uvigerina peregrina,Uvigerina proboscidea,Sphaeroidina sp.,Textularia flinti,Melonis sp.,Stilostomella sp.
F-13 sandstone reworked assemblage with early Pliocene underlying Tanrung amongst others: or youngerFormation (Walanae Sphaeroidinella dehiscensFormation?), 3 kmwest of Mancirivillage.
pp 391-421 15-01-2007 10:57 Pagina 393
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)394
A: Foraminifera (continued)Sample Stratigraphic Unit Contents Biozonation/Age Environment
F-1 Section I, ostracods, ? lagoonalSamaoling Member, echinoderm fragments3 m below basal fluvialsandstone of Benthonic foraminifera:the Beru Member Elphidium advenum,
Ammonia spp.,Ammonia gaimardi,Lenticulina spp.,Florilus spp.,Rectobolivina sp.,Operculina ammonoides
F-6 Section I, similar as sample F-1 ? lagoonalBeru Member,35 m above basalfluvial sandstonelayer of Beru Member
F-4B Puncakoro, marine Planktonic foraminifera: PL1-PL2 open seainterval of clays similar as sample F-9B.and sandstones(Burecing Member?),Walanae Formation. Benthonic foraminifera:
Lenticulina spp.,Heterolepa praecinctus,H. subhaidingeri,Cyclammina sp.,Quinqueloculina sp.,Martinottiella sp.,Textularia sp.,Dentalia sp.,Cibicides ungerianus.
pp 391-421 15-01-2007 10:57 Pagina 394
395Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Table 4B: Calcareous nannoplankton (Biozones according to Okada & Bukry, 1980).
Sample Stratigraphic Unit Contents Biozonation/Age
SF1/211091 Section I, Samaoling Member, Reticulofenestra minuta ?390 m below the basal fluvialsandstone layer of theBeru Member.
SF3/211091 Section I, Samaoling Member, Helicosphaera selli, ?415 m below the basal fluvial Pontosphaera japonica,sandstone layer of the Reticulofenestra minutaBeru Member.
SF11/211091 Section I, Burecing Member, Poatosphaera japonica, CN12 (late Pliocene)850 m below the basal fluvial Calcidiscus leptoporus, sandstone layer of the Discoaster brouweri, Beru Member. Discoaster decorus,
Discoaster sp., Helicosphaera carteri,Helicosphaera selli,Rhabdosphaera procera,Sphenolithus moriformis.
SF1/221091 Section I, Burecing Member, Discoaster brouweri, CN12 (late Pliocene)1200 m below the basal fluvial Helicosphaera carteri, sandstone layer of the Helicosphaera wallichi, Beru Member. Reticulofenestra minuta,
Sphenolithus abies.
SF3/221091 Section I, Burecing Member, Calcidiscus macintyrei, CN12 (late Pliocene)1430 m below the basal fluvial Discoaster brouweri, sandstone layer of the Helicosphaera carteri, Beru Member Helicosphaera selli,
Pontosphaera sp. Reticulofenestra minuta,Sphenolithus abies.
SF5/221091 Section I, Burecing Member, Discoaster brouweri, CN12 (late Pliocene)1650 m below the basal fluvial Discoaster decorus, sandstone layer of the Discoaster tristellifera, Beru Member Helicosphaera carteri,
Helicosphaera wallichi, Rhabdolithus sp.,Reticulofenestra minuta.
pp 391-421 15-01-2007 10:57 Pagina 395
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)396T
able
5. N
umbe
r of
id
enti
fiab
le f
ossi
l sp
ecim
ens,
att
ribu
tabl
e to
one
of
the
taxa
dis
ting
uish
ed, f
or t
he m
ost
impo
rtan
t fo
ssil
vert
ebra
te l
ocal
itie
s (F
VL
). T
heFV
L a
re o
rder
ed p
er s
trat
igra
phic
uni
t, ro
ughl
y in
chr
onol
ogic
al o
rder
fro
m o
ld (
left
) to
you
ng (
righ
t). N
umbe
rs w
itho
ut b
rack
ets
refe
r to
the
am
ount
of
insi
tu c
olle
cted
foss
ils a
ttri
buta
ble
to a
cer
tain
taxo
n, w
here
as th
e nu
mbe
r of
sur
face
col
lect
ed fo
ssils
for
a ce
rtai
n ta
xon
are
ind
icat
ed b
etw
een
brac
kets
. Exc
ept
othe
rwis
e in
dic
ated
, the
fos
sil s
peci
men
s w
ere
colle
cted
in th
e co
urse
of
this
stu
dy
betw
een
1990
and
199
4. F
ossi
ls c
olle
cted
ear
lier,
but
of
whi
ch th
e or
igin
alst
rati
grap
hic
leve
l (=
loc
.) co
uld
be
asce
rtai
ned
are
des
igna
ted
wit
h th
e fo
llow
ing
sym
bols
: *
= f
ossi
ls c
olle
cted
by
Azi
z in
198
5/86
; #
= f
ossi
ls w
hich
are
rolle
d (o
nly
ind
icat
ed fo
r FV
L-2
1 an
d F
VL
-29)
; + =
foss
ils fr
om th
e M
PC.
Stra
tigr
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Tot
al fo
ssils
in s
itu
14
35
01
810
184
00
10
114
01
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02
01
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00
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otal
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itu
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905
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215
113
36
629
23
pp 391-421 15-01-2007 10:57 Pagina 396
397Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Table 38. Ridge-crest formulas of selected Stegodon species. Incompletely developed halfridges are notincluded in the formulas. Notes: 1) S. ganesa (Falconer & Cautley, 1846) and S. insignis (Falconer &Cautley, 1846) are distinct species based on differences in skull morphology, but their molars cannotbe distinguished (Saegusa, 1987). 2) S. shinshuensis from Japan is by some authors considered to be asynonym of S. zdanskyi from China (Saegusa, 1996). The M3 of S. shinshuensis bears 7 to 9 ridges,whereas the M3 bears 8 to 9 ridges (Taruno, 1991). Its plateformula is thus slightly more advancedthan the Chinese species. 3) The M3s of S. orientalis from Japan bear 10 ridges, both the lowers anduppers (Taruno, 1991). This is less than in the mainland variety listed in the table.
Species dP2 dP3 dP4 M1 M2 M3 Source
S. elephantoides (Clift, 1828) upper - - - 6 6 - Osborn, 1942lower - - - - - 10
S. zdanskyi Hopwood, 1935 upper - 4 5 5 5-6 6-7 De Chardin &lower - 4 6 5 6 7-8 & Trassaert, 1937
S. aurorae (Matsumoto, 1918) upper - - - - - 11-13 Taruno, 1991lower - - - 8 10 12-13
S. orientalis Owen, 1887 upper 3 5-6 6-7 6-7 6-8 11-12 Colbert & Hooijer, 1953;lower 2 5-6 7 7-8 7-9 12 Hopwood, 1935; this thesis
S. bombifrons (Falc. & Cautley, upper - 4 6 6 7 9 Osborn, 19421846) lower - 4 6-7 7 8 9
S. insignis (Falc. & Cautley, upper 2 5-6 7 7-8 7-8 10-11 Osborn, 1942; this thesis1845) lower 2 6 7-9 7-10 9 11-13
S. trigonocephalus Martin, 1887subspec. S. t. trigonocephalus upper 3 6 7-8 7 9 11 this thesis
lower 2-3 6-7 8 8-9 10 13subspec. S. t. ngandongensis upper - - - 8 - - this thesis
lower - 7 9 9+ - -
S. florensis Hooijer, 1957 upper - - - 7 - 12 this thesislower - - 7+ 9 10 14
S. sondaari nov. sp. upper - 6-7 6 6-7 - - this thesislower - 6 - - 8 8
S. sompoensis Hooijer, 1964 upper - - 7 - 8 8-9 this thesislower - - - - 8 9-10
S. timorensis Sartono, 1969 upper - - - - 7 - Hooijer, 1969, 1972alower - - - - 9 10
pp 391-421 15-01-2007 10:57 Pagina 397
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)398
Table 39. Mean directions of the Natural Remanent Magnetization of the samples from the two sec-tions shown in Fig. 65. n = number of samples; D = mean declination; I = mean inclination; k = Fish-er’s precision estimate; α95 = semiangle of cone of 95% confidence; N = Normal magnetization; R =Reversed magnetization.
Sample Lithology n D I k α95 Polarity
01 very fine sandstone 5 147 14 13.1 17.1 R02 silty sandstone 5 238 -20 10.1 14.1 R03 silty sandstone 4 189 -10 10.2 13.0 R04 very fine sandstone 4 14 -5 9.3 23.0 N05 tuff 6 160 -20 9.6 18.1 R06 siltstone 6 234 -39 12.1 16.1 R07 tuff 5 34 -13 5.1 23.0 N08 silty sandstone 5 40 -24 4.3 20.0 N09 siltstone 5 59 -80 173.8 4.7 N10 siltstone 4 7 -19 4.0 35.0 N11 siltstone 4 213 -23 15.7 17.6 R12 siltstone 5 188 -14 4.7 22.0 R13 sandstone 4 253 -7 104.1 6.9 R/N14 siltstone 5 322 -65 6.1 25.3 N15 very fine sandstone 4 36 -27 6.8 25.7 N16 silty sandstone 4 2 -33 38.3 11.3 N17 silty sandstone 5 350 -24 318.5 4.7 N18 limestone 6 - - - - -19 very fine sandstone 6 341 -30 66.9 7.0 N
pp 391-421 15-01-2007 10:57 Pagina 398
399Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Table 40. Body mass estimations of adult individuals of various Indonesian Stegodon species, based ontotal length of femur or humerus, or based on midshaft circumference of femur or humerus. FemurCD-4315 bears a label indicating that it originates from Trinil, but not from the ‘Haupt-Knochen-schicht’. The label states that it comes from a layer at an elevation 4 to 5 m above the lowest waterlev-el of the Soloriver, whereas the fossils from the ‘Haupt-Knochenschicht’ were excavated from belowthe lowest waterlevel (de Vos & Sondaar, 1982). Unlike the brown-coloured fossils from the ‘Haupt-Knochenschicht’, femur CD-4315 has a reddish colour and a red sandstone matrix attached to it. Forfurther explanation of the mass estimation methods see text. Mass estimates based on shaft circumfer-ence are more than twice as large as estimates based on length. These discrepancies are caused by therelatively sturdy built limbbones of Stegodon. The length-based bodymass estimates are thought toapproach the true bodymass closer. In addition, the circumference-based bodymass estimate for ahumerus of “Elephas” celebensis is given.
species element specimen length circumference mass estimate mass estimate(mm) (mm) based on length based on cir-
(kg) cumf. (kg)
S. sondaari sp. nov. femur TT-4083 460e ± 50 151.5 207 +65/-54 553diaphysis
S. florensis humerus CV-72 630 - 852 -(Hooijer,1972a: 23)
S. florensis humerus (Hooijer, 530+ 273e-280e - 2169-23171957a: 125)
S. t. trigonocephalus femur CD-2890 922 307 1310 4072
S. t. trigonocephalus femur CD-4315 838 296 1017 3673
S. t. trigonocephalus femur CD-2889 1020 358 1713 6287
S. sompoensis humerus LR-3707 452e ± 8 199 350 +17/-12 950
"Elephas" celebensis humerus PL-3736 - 187 - 807
pp 391-421 15-01-2007 10:57 Pagina 399
Table 41. Successive stages which can be distinguished in a single, progressing elephantoid molar(after Beden, 1979). 11 stages were defined, designated with the combination of a letter (A-D) and anumber (except for stage C). To the right the alveolar states, corresponding with the successive molarstages, are indicated. For further explanation, see chapter 8.
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)400
opening of thealveole
(0)
new
Sta
te o
f th
e m
olar
Sta
te o
f th
e al
veol
e
posterior ridges not yetconsolidated
entire molar consolidated
cementum worn
a few ridges worn
half of the ridges worn
all ridges worn (except insome cases the posteriorhalfridge)
most of the ridges worn
half of the ridges gone
anterior border of the firstridge partially brokenaway
first ridges gone
remnant of a few ridgesleft
pro
gres
sive
wea
r of
mol
ar
increasingnumber ofridges in function
all ridges infunction
progressivedestructionof ridges
molar crownpartly visible
molar crownvisible
rootmass stillvisible
1
A
B
C
2
1
2
3
4
1
2
3
4
(A)
(A)
(B1)
(B2)
(B3)
B4 (B4)
C (C)
(D1)
D1
D2
D3
D4
D
0
pp 391-421 15-01-2007 10:57 Pagina 400
401Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Table 42. Five age groups that are here distinguished for reconstructing mortality profiles of elephan-toid dental assemblages. Each age group represents a lifespan of approximately 12 AEY. The dentalwear age classes of Beden (1979) were used for aging individual elephantoid dentitions and certainclasses define the boundaries of the age groups (see also Figs. 75-76). The corresponding dental wearage classes of Laws (1966) are also given.
age social status range of dental wear classesgroup after Beden (1979) after Laws (1966)
1 juveniles 0 - M1A I - XI2 young adults M1A’ - M2A XII - XVII3 prime adults M2B - M2/M3B’ XVIII - XXI4 senior adults M2/M3C - M3B XXII - XXIV5 senile M3C - M3E’ XXV - XXX
pp 391-421 15-01-2007 10:57 Pagina 401
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)402
Plate 1
Figs. 1-4. Stegodon elephantoides (Clift, 1828) MPS-358, Bukuran, Central Java; Lower Pleistocene; mandible with both M3s; 1: oblique occlusalview; 2: lateral view; 3: frontal view; 4: occlusal view.
1
3
2
4
50 cm
50 cm
50 c
m
20 c
m
pp 391-421 15-01-2007 10:57 Pagina 402
403Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Plate 2
Figs. 1-2. Stegodon trigonocephalus ngandongensis subsp. nov.1: GRDC/K-330B, Ngandong excavation; Upper Pleistocene; dex. dP3 occlusal view.2: GRDC/K-330A: dex. dP3, Watualang excavation, occlusal view.
Figs. 3-4. Elephas maximus Linnaeus, 1758GRDC CPD90-1, sin. M2, Cipeundeuy sand quarry, West Java; Upper Pleistocene; 3: occlusal view; 4:medial view.
12
3
4
5 cm 5
cm
5 cm
5 cm
pp 391-421 15-01-2007 10:57 Pagina 403
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)404
Plate 3
Figs. 1-7. “Elephas” celebensis (Hooijer, 1949)1-2: GRDC S-3949; loc.: FVL-25 (Sompe), South Sulawesi; Upper Pliocene; mandible fragment withdP3 and anterior dP4 fragment; 1: occlusal view; 2: lateral (buccal) view. 3: GRDC Lp-3192; loc.: FVL-1 (Lepangeng), South Sulawesi; Upper Pliocene; dextral mandibularramus with roots of dP4 and P3 and the M1 in the alveole. 4-5: GRDC 1307; loc.: FVL-5 (Lakibong), South Sulawesi; Lower Pleistocene; sinistral mandibularramus with P4; 4: occlusal view; 5: lateral view; 6-7: GRDC 1344; dextral maxilla fragment with P3; 6:occlusal view; 7: lateral view.
1
2
3
4
5
6
7
5 cm
5 cm
5 cm
10 c
m
pp 391-421 15-01-2007 10:57 Pagina 404
405Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Plate 4
Figs. 1-4. “Elephas” celebensis (Hooijer, 1949)1-2: GRDC 1342; loc.: FVL-5 (Lakibong); dextral dP4; 1: occlusal view; 2: lateral view.
Figs. 3-4. Stegodon sompoensis Hooijer, 1964 GRDC MUTL/171186-1; loc.: c. 1 km east of FVL-17 (Alupang), South Sulawesi; dextral mandibularramus fragment with M3 fragment; Upper Pliocene or Lower Pleistocene; 3: occlusal view; 4: lateralview.
1
2
3
4
5 cm
10 cm
10 cm
pp 391-421 15-01-2007 10:57 Pagina 405
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)406
Plate 5
Figs. 1-2. “Elephas” celebensis (Hooijer, 1949) GRDC LWTL/151186-1; loc.: FVL-7 (Lakibong), South Sulawesi; Lower Pleistocene; skull with M3; 1:frontal view; 2: lateral view.
1
2
20 cm
20 c
m
pp 391-421 15-01-2007 10:57 Pagina 406
407Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Plate 6
Figs. 1-5. Stegodon sompoensis Hooijer, 19641-2: PMC C3/2/79; loc.: FVL-17 (Marale), South Sulawesi; Lower Pleistocene; sin. mandible with pos-terior M2 fragment and M3 in alveole; 1: lingual view; 2: occlusal view. 3-4: GRDC PA-3730; loc.: FVL-9 (Paroto) South Sulawesi; Upper Pliocene or Lower Pleistocene; dex-tral dP4; 3: occlusal view; 4: buccal view. 5: GRDC L/III-3036; loc.: FVL-2 (Palangiseng); Lower Pleistocene; premaxilla, dorsal view.
13
4
2
5
5 cm
10 c
m
20 c
m
10 c
m
pp 391-421 15-01-2007 10:57 Pagina 407
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)408
Plate 7
Figs. 1-4. Stegodon sompoensis Hooijer, 1964 GRDC BC-3050; loc.: FVL-10 (Pajalela), South Sulawesi; adult skull lacking dentition; 1: ventral view;2: lateral view; 3: occipital view; 4a, b: frontal view.
3
2
1
4a 4b
20 cm
20 cm20
cm
pp 391-421 15-01-2007 10:57 Pagina 408
409Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Plate 8
Figs. 1-2. Stegodon sp. B Loc.: FVL-29 (Tanrung River), South Sulawesi; Middle Pleistocene; 1: GRDC TA-3711; M1 sin; occlusalview. 2: GRDC TA-3712; dP4 remnant and M1; 2a: occlusal view; 2b: buccal view.
Figs. 3-4. large-sized Stegodon sp. GRDC L/III-3040; loc.: FVL-2 (Palangiseng), South Sulawesi; Lower Pleistocene?; sinistral lowermolar fragment; 3: occlusal view (lingual side partly embedded in matrix; 4: buccal view.
1
2a
2b
34
5 cm
5 cm
5 cm 5 cm
5 cm
pp 391-421 15-01-2007 10:57 Pagina 409
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)410
Plate 9
Figs. 1-3. Stegodon sompoensis Hooijer, 1964 Loc.: FVL-24 (Lonrong), South Sulawesi; Upper Pliocene; 1: GRDC LR-3707; dextral humerus; anteriorview; 2-3: GRDC LR-3546; dextral radius; 2: posterolateral view; 3: anteromedial view.
Plate 10
Figs. 1-2. pygmy Elephantoidea GRDC LR-3552; loc.: FVL-24 (Lonrong), South Sulawesi; Upper Pliocene; dextral metatarsus III; 1:anterior view; 2: lateral view.
1
2 3
20 c
m
20 c
m
pp 391-421 15-01-2007 10:57 Pagina 410
411Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Figs. 3-6. Elephas sp. Loc.: FVL-29 (Tanrung River), South Sulawesi; Middle or Late Pleistocene; 3-4: GRDC TA-3061; dex-tral metacarpus III; 3: posterior view; 4: lateral view; 5-6: GRDC TA-3062: dextral metacarpus V; 5:posterior view; 6: lateral view.
1
2
3
56
4
5 cm
5 cm
5 cm
5 cm
5 cm
5 cm
pp 391-421 15-01-2007 10:57 Pagina 411
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)412
Plate 11
Figs. 1-6. Stegodon sondaari sp. nov. Loc.: Tangi Talo, west Central Flores; Lower Pleistocene (c. 0.9 Ma); 1-2: GRDC TT-4044; dP3 sin.; 1:occlusal view; 2: lingual view; 3-4: GRDC TT-3835; dP4 sin.; 3: occlusal view; 4: lingual view; 5a:GRDC TT-4033; dP4 dex, and 5b: GRDC TT-4034; dP4 sin.; both dP4 constitute a pair; occlusal view; 6:GRDC TT-4034; dP4 sin., lingual view.
12
3 4
5a 5b
6
2 cm
2 cm
5 cm
5 cm
pp 391-421 15-01-2007 10:57 Pagina 412
413Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Plate 12
Figs. 1-5. Stegodon sondaari sp. nov. Loc.: Tangi Talo, west Central Flores; Lower Pleistocene (c. 0.9 Ma); 1-2: GRDC TT-3836; dex. maxil-lary fragment with dP4; 1: occlusal view; 2: lateral (buccal) view; 3: GRDC TT-4032; dP4 sin., mesialview; 4a: GRDC TT-3814; M2 sin.; 4b: GRDC TT-3818; M2 dextral, both M2 constitute a pair; occlusalview; 5: TT-3818: M2 dextral, buccal view.
1
2
3
4a 4b
5
5 cm
5 cm
5 cm
5 cm
pp 391-421 15-01-2007 10:57 Pagina 413
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)414
Plate 13
Figs.1-3. Stegodon sondaari sp. nov. Loc.: Tangi Talo, west Central Flores; Lower Pleistocene (c. 0.9 Ma); holotype; GRDC TT-3837;mandible with both worn M1, slightly worn M2 and the uncompleted sinistral M3 still in the alveolarcavity; 1: occlusal view; 2: lateral view; 3: medial view of the sinistral ramus.
1
2
3
10 cm
10 cm
10 c
m
pp 391-421 15-01-2007 10:57 Pagina 414
415Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Plate 14
Figs.1-6. Stegodon sondaari sp. nov. Loc.: Tangi Talo, west Central Flores; Lower Pleistocene (c. 0.9 Ma).1: GRDC TT-4035; M1 dex., and GRDC TT-4031: M2 dex.; occlusal view; 2: GRDC TT-4037; M1 sin.,and GRDC TT-4030: M2 sin.; buccal view; (teeth figured in figs. 1-2 belong to the same individual asfigured in Pl. 13); 3-4: GRDC TT-3816; M2 dex.; 3: occlusal view; 4: lingual view; 5-6: GRDC TT-4029;M2 sin.; 5: occlusal view; 6: lingual view.
1
2
3
4
6
5
5 cm
5 cm
5 cm
5 cm
5 cm
pp 391-421 15-01-2007 10:57 Pagina 415
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)416
Plate 15
Figs.1-4. Stegodon sondaari sp. nov. Loc.: Tangi Talo, west Central Flores; Lower Pleistocene (c. 0.9 Ma); 1: GRDC TT-3856; M3 sin. frag-ment, occlusal view; 2: GRDC TT-3887/88; sin. tusk; dorso-medial view; 3-4: GRDC TT-3819; sin. tusk;3: dorso-medial view; 4: ventro-medial view.
1
2
3
4
5 cm
5 cm
5 cm
pp 391-421 15-01-2007 10:57 Pagina 416
417Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Plate 16
Figs.1-5. Stegodon sondaari sp. nov. Loc.: Tangi Talo, west Central Flores; Lower Pleistocene (c. 0.9 Ma); 1-2: F.BS 3.1; atlas; 1: caudal view;2: dorsal view; 3-4: GRDC TT-4086; vertebra thoracale IV or V; 3: lateral view; 4: caudal view; 5: innerenamel layer (IEL) and outer enamel layer (OEL). enamel/dentine junction (EDJ) to the left; polariz-ing microscope.
Fig. 6. Stegodon sompoensis Hooijer, 1964 IEL and OEL. EDJ to the left; polarizing microscope.
1 2
3 4
5 6
5 cm
5 cm
5 cm
1 mm1 mm
pp 391-421 15-01-2007 10:57 Pagina 417
Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)418
Plate 17
Figs. 1-6. Stegodon sondaari sp. nov. Loc.: Tangi Talo, west Central Flores; Lower Pleistocene (c. 0.9 Ma); 1-2: GRDC TT-4082; vertebra tho-racale (XVI - XIX ?); 1: lateral view; 2: caudal view; 3-4: GRDC TT-4083; sinistral femur diaphysis; 3:anterior view, 4: posterior view; 5-6: GRDC TT-4065; phalanx I; 5: lateral view; 6: palmar view.
1
2
3
4
5
6
10 c
m
5 cm
2 cm
pp 391-421 15-01-2007 10:57 Pagina 418
419Van den Bergh. The Late Neogene elephantoid-bearing faunas of Indonesia. Scripta Geol., 117 (1999)
Plate 18
Figs. 1-3. Stegodon florensis Hooijer, 1957; west Central Flores.1-2: GRDC DD-4160; loc.: Dozo Dhalu; mandible with M1 remnants and both M2s; 1: lateral view; 2:occlusal view. 3: GRDC MM-4118; loc.: Mata Menge; dextral M3; 3: occlusal view.
1
2
3
20 cm
10 c
m
20 c
m
pp 391-421 15-01-2007 10:57 Pagina 419