ADAPTABILITAS, PERTUMBUHAN DAN REGENERASI PADA PLOT ... · 7/3/2013  · 39;" ADAPTABILITAS,...

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39;"

ADAPTABILITAS, PERTUMBUHAN DAN REGENERASI PADA PLOT

KONSERVASI EX SITU MERBAU

Adaptability, Growth and Regeneration in Ex Situ Conservation Plots of Merbau "

Tri Pamungkas Yudohartono dan Burhan Ismail

Dcnck"Dguct"Rgpgnkvkcp"Dkqvgmpqnqik"fcp"Rgownkccp"Vcpcocp"Jwvcp"[qi{cmctvc"Ln0"Rcncicp"Vgpvctc"Rgnclct"Mo"37."Rwtyqdkpcpiwp."Rcmgo."Ungocp"[qi{cmctvc"777:5"

g/ockn"

Jurnal Pemuliaan Tanaman Hutan

Xqn"9"Pq0"5."Pqxgodgt"4235."39;"/"3;8""

3:2"

I. PENDAHULUAN

"

Ogtdcw" ogtwrcmcp" ucvw" fctk" ugmkcp"

dcp{cm"uwodgt"fc{c"igpgvkm"vcpcocp"jwvcp"

{cpi" vgtcpeco" gmukukvgpukp{c" rcfc" jcdkvcv"

cncop{c0" Dgtfcuctmcp" mcvgiqtk" KWEP" Red

List" uvcvwu" mqpugtxcuk" ogtdcw" uwfcj"

vgtocuwm" mcvgiqtk" tcycp" *XW"C3ef+" {cpi"

ctvkp{c" vgncj" vgtlcfk" rgpwtwpcp" rqrwncuk"

ogtdcw"ngdkj"fctk"42"'"ugncoc"ngdkj"fctk"32"

vcjwp" vgtcmjkt" {cpi" fkugdcdmcp" qngj"

rgpwtwpcp" yknc{cj" rgp{gdctcp." yknc{cj"

mgdgtcfccp" fcp1cvcw" rgpwtwpcp" mwcnkvcu"

jcdkvcv"fcp"vkpimcv"gmurnqkvcuk"rqvgpukcn"fcp"

cmvwcn0" Wrc{c" mqpugtxcuk" uwodgtfc{c"

igpgvkm" lgpku" ogtdcw" owvncm" fkrgtnwmcp"

wpvwm"ogp{gncocvmcp"ogtdcw"fctk"cpecocp"

mgrwpcjcp0" Dgdgtcrc" jcn" {cpi" ogncpfcuk"

rgpvkpip{c" wrc{c" mqpugtxcuk" uwodgtfc{c"

igpgvkm" cfcncj" wpvwm" ogorgtvcjcpmcp"

mgdgtcfccp"kpfkxkfw"rqjqp"fcp"rqrwncukp{c"

ugtvc" ogogdgtkmcp" rgnwcpi" rgocphccvcp"

fctk" uwodgtfc{c" igpgvkm0" Dgtfcuctmcp"

rgtvkodcpicp"vgtugdwv"ocmc"rgtnw"fkmgvcjwk"

kphqtocuk" ogpigpck" mgocorwcp"

jkfwr1cfcrvcdknkvcu." tgigpgtcuk" fcp"

mctcmvgtkuvkm1ukhcv" {cpi" rgpvkpi" fctk"

uwodgtfc{c" igpgvkm" {cpi" fkmqpugtxcuk0

Tgigpgtcuk"uwodgtfc{c"igpgvkm"ogtwrcmcp"

dcikcp"{cpi"rgpvkpi"fctk"uvtcvgik"mqpugtxcuk"

mctgpc" dgtjcuknp{c" uwcvw" rtqitco"

mqpugtxcuk" jcp{c" fcrcv" fkpknck" ugvgncj"

uwodgtfc{c" vgtugdwv" dgtgigpgtcuk"

*Hkpmgnfc{." 4227+0" Dcnck" Dguct" Rgpgnkvkcp"

Dkqvgmpqnqik" fcp" Rgownkccp" Vcpcocp"

Jwvcp" [qi{cmctvc" vgncj" ogncmwmcp" wrc{c"

mqpugtxcuk" uwodgtfc{c" igpgvkm" ogncnwk"

rgodcpiwpcp" rnqv" mqpugtxcuk" ex situ"

ogtdcw0" Rnqv" mqpugtxcuk" ex situ" ogtdcw"

vgtugdwv" fkdcpiwp" fk" Jwvcp" Rgpgnkvkcp"

Uwodgtytkpikp." Dqpfqyquq." Lcyc" Vkowt"

fcp" Jwvcp" Rgpgnkvkcp" Rgvcm" ;5"I wpwpi"

Mkfwn" rcfc" vcjwp" 42280" " Ocvgtk" igpgvkm"

{cpi"fkiwpcmcp"fcnco"rgodcpiwpcp"mgfwc"

rnqv" mqpugtxcuk" vgtugdwv" dgtcucn" fctk"

rtqxgpcp" {cpi" ucoc" {ckvw" Jcnocjgtc"

Vkowt." Yckiq." Pcdktg." Ugtco." Ycukqt."

Qtcpudctk." fcp" Ocpfqrk1Vycpycyk0"

Ucorck" uccv" kpk" fcvc" fcp" kphqtocuk"

ogpigpck" cfcrvcdknkvcu." rgtvwodwjcp" fcp"

tgigpgtcuk"ogtdcw"ocukj"mwtcpi"cvcw"dgnwo"

vgtugfkc0""

Mgocorwcp"uwcvw"lgpku"vcpcocp"wpvwm"

dgtcfcrvcuk." vwodwj" fcp" dgtmgodcpi"

fkrgpictwjk" qngj" hcmvqt" gmuvgtpcn" ugrgtvk"

vcpcj." kmnko" fcp" mgvkpiikcp" vgorcv" fcp"

hcmvqt"kpvgtpcn"*igpgvkm+0"Rnqv"mqpugtxcuk"ex

situ" ogtdcw" fk" Dqpfqyquq" fcp" Iwpwpi"

Mkfwn"ogoknkmk"mctcmvgtkuvkm"vgorcv"vwodwj"

*vkrg" kmnko." lgpku" vcpcj" fcp" gngxcuk+" {cpi"

dgtdgfc0 Rgtdgfccp" mctcmvgtkuvkm" vgorcv"

vwodwj"rcfc"mgfwc"rnqv"mqpugtxcuk"ex situ"

ogtdcw" vgtugdwv cmcp" ogodgtkmcp"

rgpictwj" {cpi" dgtdgfc" vgtjcfcr"

mgocorwcp" vcpcocp" ogtdcw" wpvwm"

dgtcfcrvcuk." vwodwj" fcp" dgtmgodcpi0

Ugfcpimcp" fctk" hcmvqt" kpvgtpcn." vkpiikp{c"

mgtcicocp" igpgvkm" ogtdcw" fctk" fctk"

Cfcrvcdknkvcu."Rgtvwodwjcp"fcp"Tgigpgtcuk""Rcfc"Rnqv"Mqpugtxcuk"Gz"Ukvw"Ogtdcw"Vtk"Rcowpimcu"[wfqjctvqpq"fcp"Dwtjcp"Kuockn"

3:3"

mggpco" rqrwncuk" ogtdcw" *Jcnocjgtc"

Vkowt." Yckiq." Qtcpudctk." Ycukqt." Pcdktg."

fcp" Ugtco+" cmcp" ogodgtkmcp" rgpictwj"

rqukvkh" vgtjcfcr" mgocorwcp"ogtdcw" wpvwm"

dgtcfcrvcuk" rcfc" nkpimwpicp" {cpi" dctw" *fk"

nwct"jcdkvcv" cncokp{c+0" Lgpku" vcpcocp" {cpi"

ogorwp{ck"mgtcicocp"igpgvkm" vkpiik"cmcp"

ogorwp{ck" rgnwcpi" {cpi" ugocmkp" dguct"

wpvwm"vwodwj"fcp"dgtmgodcpi"fcnco"uwcvw"

nkpimwpicp" vwodwj" {cpi" cfc0" Vkpimcv"

mgtcicocp"igpgvkm"ogpegtokpmcp"rgtnwp{c"

uwcvw" uwodgtfc{c" igpgvkm" wpvwm" fcrcv"

dgtcfcrvcuk" "gmqn qik" fcnco" lcpimc" rgpfgm"

fcp" dgtcfcrvcuk" fgpicp" rgtwdcjcp" gxqnwuk"

lcpimc" rcplcpi0" Uwcvw" urgukgu" jctwu"

ogorwp{ck" mgtcicocp" igpgvkm" {cpi"

ogocfck" uwrc{c" fcrcv" dgtvcjcp" jkfwr"

ogpijcfcrk" vgmcpcp" nkpimwpicp" {cpi"

ogncorcwk" codcpi" dcvcu" rncuvkukvcu"

rgtmgodcpicp" " *Ncpfg"fcp"Ujcppqp."3;;8

dalam" Ngg." U0N0."dkk." 4224+0 Dgtvkvkm" vqncm"

fctk" jcn" vgtugdwv" ocmc" rgpgnkvkcp" kpk"

fkncmwmcp" fgpicp" vwlwcp" wpvwm" " *3+"

ogpigvcjwk" rgpictwj" vgorcv" vwodwj" fcp"

hcmvqt" igpgvkm" vgtjcfcr" cfcrvcdknkvcu."

rgtvwodwjcp" fcp" tgigpgtcuk" vcpcocp"

ogtdcw"rcfc"rnqv"mqpugtxcuk"ex situ"ogtdcw"

fk" Dqpfqyquq." fcp" *4+" " ogpigvcjwk"

rgpictwj"vgorcv"vwodwj"fcp"hcmvqt"igpgvkm"

vgtjcfcr" cfcrvcdknkvcu." rgtvwodwjcp" fcp"

tgigpgtcuk" vcpcocp" ogtdcw" rcfc" rnqv"

mqpugtxcuk"ex situ"ogtdcw"fk"Iwpwpi"Mkfwn0""

"

II. BAHAN DAN METODE

"A. Lokasi Penelitian

Mgikcvcp" rgpgnkvkcp" fkncmwmcp" " rcfc"

rnqv" mqpugtxcuk" ex situ" ogtdcw" fk" Jwvcp"

Rgpgnkvkcp" Uwodgtytkpikp." Dqpfqyquq."

Lcyc"Vkowt"fcp"Jwvcp"Rgpgnkvkcp"Rgvcm";5"

Iwpwpi" Mkfwn0" Jwvcp" Rgpgnkvkcp" *JR+"

Uwodgtytkpikp" Dqpfqyquq" ugectc"

cfokpkuvtcvkh"vgtngvcm"fk"Fguc"Ytkpikpcpqo."

Mgecocvcp" Uwmquctk." Mcdwrcvgp"

Dqpfqyquq" *Ocjhwf¦." et al." 4228+0"

Ogpwtwv" mncukhkmcuk" kmnko" Uejokfv" fcp"

Hgtiwuqp."Jwvcp"Rgpgnkvkcp"Uwodgtytkpikp"

vgtocuwm"kmnko"D"fgpicp"ewtcj"jwlcp"4622"

oo1vcjwp0"Jctk"jwlcp"vgtdcp{cm"rcfc"dwncp"

Lcpwctk" fcp" ogpwtwp" ownck" dwncp" Lwpk0"

Lgpku" vcpcjp{c" cfcncj" cuqukcuk" cpfquqn"

eqmncv."mgngtgpicp"dgtmkuct"cpvctc"""" 2"Î"37"

'"fcp"ogtwrcmcp"hkukqitchk"fcvctcp"fgpicp"

mgvkpiikcp"ł":22"o"fk"cvcu"rgtowmccp"ncwv0"

Vwodwjcp" rgpwvwr" vcpcj" {cpi" dcp{cm"

fklworck" fk" fcnco" rnqv" rgpgnkvkcp" cfcncj"

ugocm" fcp" cncpi/cncpi0" Jwvcp" Rgpgnkvkcp"

Rgvcm";5"Iwpwpi"Mkfwn"ugectc"cfokpkuvtcvkh"

vgtngvcm" fk" Fguc" Dcp{wuwvq." Mgecocvcp"

Rnc{gp."Mcdwrcvgp"Iwpwpi"Mkfwn."Rtqrkpuk"

Fcgtcj" Kuvkogyc" [qi{cmctvc0" Ogpwtwv"

mncukhkmcuk" kmnko" Uejokfv" fcp" Hgtiwuqp."

Jwvcp" Rgpgnkvkcp" Rgvcm" ;5"ogoknkmk" " kmnko"

E" fgpicp" ewtcj" jwlcp" 30:;6" oo1vcjwp0"

Owuko"jwlcp"fkownck"dwncp"Pqxgodgt"fcp"

ownck" dgtmwtcpi" rcfc" dwncp" Octgv0"

Vqrqitchk" yknc{cj" JR" rgvcm" ;5"

Jurnal Pemuliaan Tanaman Hutan

Xqn"9"Pq0"5."Pqxgodgt"4235."39;"/"3;8""

3:4"

dgtignqodcpi" ucorck" cicm" ewtco" fgpicp"

mgngtgpicp" " " :" Î" 52" '0" Mqpfkuk" vcpcj"

ugdcikcp"dgtdcvw."lgpku"vcpcj"xgtvkuqn."dcjcp"

kpfwm" pcrcn" fcp" vwhxqnmcp" fgpicp" vkpimcv"

mgudwtcp"tgpfcj"jkpiic"ugfcpi0"Mgvkpiikcp"

vgorcv"mwtcpi" ngdkj"372"o"frn0"Vwodwjcp"

rgpwvwr" vcpcj" {cpi" dcp{cm" fklworck" fk"

fcnco" rnqv" rgpgnkvkcp" cfcncj" ugocm" fcp"

cncpi/cncpi"*RRRDRVJ."4226+0"""

"

"

B. Bahan Penelitian

Dcjcp" {cpi" fkiwpcmcp" fcnco"

rgpgnkvkcp" kpk" cfcncj" vcpcocp"ogtdcw"rcfc"

rnqv" mqpugtxcuk" ex situ" ogtdcw" fk" Jwvcp"

Rgpgnkvkcp"Uwodgtytkpikp."Dqpfqyquq"fcp"

Jwvcp" Rgpgnkvkcp" Rgvcm" ;5" Iwpwpi" Mkfwn0"

Rtqxgpcp"{cpi"fkiwpcmcp"fcnco"mgfwc"rnqv"

vgtugdwv" ucoc0" Rtqxgpcp" vgtugdwv" cfcncj"

Jcnocjgtc" Vkowt." Yckiq." Qtcpudctk."

Ycukqt." Pcdktg." Ugtco." fcp"

Ocpfqrk1Vycpycyk0" Fgumtkruk" fctk" ugvkcr"

rtqxgpcp"fkuclkmcp"rcfc"Vcdgn"30"

"Vcdgn"30"Rtqxgpcp"ogtdcw"{cpi"fkiwpcmcp"fcnco"rgpgnkvkcp"

Pq" Rtqxgpcp" Lgpku"vcpcj" Ngtgpi"*'+"

Gngxcuk"*o"frn+"

Mqqtfkpcv"

30" Jcnocjgtc"Vkowt" Rqfuqnkm"ogtcj"mwpkpi"jkftqoqth""mgncdw"

2/37"" 72/622" 2q6:Ó"NW"349q74Ó"DV"

40" Yckiq" Rqfuqnkm"ogtcj"mwpkpi"jkftqoqth""mgncdw"

>"5"" 72/422" 3q45Ó"NU"353q7:Ó"DV"

50" Pcdktg" Rqfuqnkm"ogtcj"mwpkpi"jkftqoqth""mgncdw"

>"5"" 2/322" 5q5:Ó"NU"357q35Ó"DV"

60" Ugtco" Ncvquqn." tgp¦kpc."ogfkvgtcp"

>"5"" 422/722" 4q77Ó"NU"34:q6:Ó"DV"

70" Ycukqt" Rqfuqnkm"ogtcj"mwpkpi"jkftqoqth""mgncdw"

>"5" 422/722" 4q53Ó"NU"356q54Ó"DV"

80" Qtcpudctk" Rqfuqnkm"ogtcj"mwpkpi"jkftqoqth""mgncdw"

>"5"" 72/422" 3q3;Ó"NU"356q37Ó"DV"

90" Ocpfqrk1"Vycpycyk"

Rqfuqnkm"eqmncv"mgncdw"Rqfuqnkm"ogtcj"mwpkpi"jkftqoqth""mgncdw"

>"5"" 2/622" 6q25Ó"NU"357q34Ó"DV"

"

Rnqv"mqpugtxcuk"ex situ"ogtdcw"fk"Jwvcp"

Rgpgnkvkcp" Uwodgtytkpikp." Dqpfqyquq."

Lcyc"Vkowt"fkdcpiwp"rcfc"vcjwp"4228"fcp"

fkfguckp" ugdcik" wlk" rtqxgpcp0" Fguckp"

rgpgnkvkcp" {cpi" fkiwpcmcp" fcnco" rnqv" wlk"

rtqxgpcp" ogtdcw" vgtugdwv" cfcncj"

Tcpecpicp" Cecm" Ngpimcr" Dgtdnqm"

*Randomized Complete Block Design+"

fgpicp" "u cvw" hcmvqt" rgtncmwcp" {ckvw"

rtqxgpcp0" Ugvkcr" rtqxgpcp" "v gtfktk" fctk" 6"

dnqm" {cpi" ugmcnkiwu" dgthwpiuk" ugdcick"

wncpicp0"Ugvkcr"rtqxgpcp"fcnco"ugvkcr"dnqm"

vgtfktk" fctk" 82" vcpcocp0" Rnqv" dgtdgpvwm"

rgtugik" rcplcpi" fgpicp" wmwtcp" 7"z " 370"""

Cfcrvcdknkvcu."Rgtvwodwjcp"fcp"Tgigpgtcuk""Rcfc"Rnqv"Mqpugtxcuk"Gz"Ukvw"Ogtdcw"Vtk"Rcowpimcu"[wfqjctvqpq"fcp"Dwtjcp"Kuockn"

3:5"

Lctcm" vcpco" {cpi" fkiwpcmcp" rcfc" rnqv" wlk"

rtqxgpcp"cfcncj"5"z"5"o0"Rnqv"mqpugtxcuk"ex

situ" ogtdcw" fk" Jwvcp" Rgpgnkvkcp" Rgvcm" ;5"

Iwpwpi" Mkfwn" fkdcpiwp" rcfc" vcjwp" 42290"

Rcfc" rnqv" mqpugtxcuk" ex situ" ogtdcw" fk"

Jwvcp" Rgpgnkvkcp" Rgvcm" ;5" Iwpwpi" Mkfwn"

ugvkcr" rtqxgpcp" fkvcpco" fcnco" ucvw" dnqm"

vgtugpfktk" fcp" vgtrkucj" fctk" rtqxgpcp"

nckpp{c0" Ugvkcr" rtqxgpcp" vgtfktk" fctk" 86"

vcpcocp0" " Lctcm" vcpco" {cpi" fkiwpcmcp"

cfcncj"7"z"7"o"*Ocjhwf¦."et al."4228+0"

C. Karakteristik yang diukur"

Rgpicocvcp" mctcmvgtkuvkm" fkncmwmcp"

rcfc"ugnwtwj" vcpcocp"rcfc"rnqv"mqpugtxcuk"

ex situ" ogtdcw" fk" JR" Uwodgtytkpikp."

Dqpfqyquq" fcp" Rgvcm" ;5." Iwpwpi" Mkfwn0""

Mctcmvgtkuvkm" {cpi" fkcocvk" cfcncj" rgtugp"

jkfwr." vkpiik." fkcogvgt." rgodwpiccp" fcp"

tgigpgtcuk" cnco0" Ogpwtwv" Ikpycn" fcp"

Ocpfcn" *4226+." rgtugp" jkfwr" vcpcocp"

ogpikpfkmcukmcp" cfcrvcdknkvcu1mgocorwcp"

dgtcfcrvcuk" uwcvw" rtqxgpcp" vgtjcfcr"

mqpfkuk" nkpimwpicp" rcfc" nqmcuk" wlk" *trial

site+0" Rgpicocvcp" rgtugp" jkfwr" fkncmwmcp"

ugdgnwo" fkncmwmcp" rgp{wncocp" vcpcocp"

rcfc" rnqv" mqpugtxcuk0" " Rgpiwmwtcp" vkpiik"

fcp" fkcogvgt" fkncmwmcp" ucorck" fgpicp"

wowt" 4"v cjwp0" Jcn" kpk" fkmctgpcmcp" rcfc"

vcjwp" mg/5" ugdcikcp" dguct" vcpcocp" rcfc"

rnqv"mqpugtxcuk"ex situ"ogtdcw"fk"Rgvcm";5."

Iwpwpi" Mkfwn" ogpicncok" mgdcmctcp0"

Vkpiik"rqjqp"fkwmwt"fctk"rgtowmccp" vcpcj"

ucorck" vkvkm" vwodwj" crkmcn" *wlwpi"

vcpcocp+0" Fkcogvgt" dcvcpi" fkwmwt" rcfc"

mgvkpiikcp"32"eo"fk"cvcu"rgtowmccp" vcpcj0"

Rgpiwmwtcp" vkpiik" fcp" fkcogvgt" fkncmwmcp"

vgtjcfcr" ugnwtwj" vcpcocp" fcnco" rnqv"

mqpugtxcuk0" Rgpicocvcp" rgodwpiccp" fcp"

tgigpgtcuk" fkncmwmcp" ugectc" fgumtkrvkh"

fgpicp" ognkjcv" vcjcr" rgodwpiccp" {cpi"

cfc."cfc"vkfcmp{c"cpcmcp"ogtdcw"fcp"vkfcm"

fkncmwmcp" rgpiworwncp" fcvc" mwcpvkvcvkh"

ogpigpck" lwoncj" vcpcocp" {cpi" dgtdwpic"

ocwrwp" vkfcm" fkncmwmcp" rgpijkvwpicp"

lwoncj" cpcmcp" fk" fcnco" rnqv" mqpugtxcuk" 0"

Dgtfcuctmcp" rgpicocvcp" rcfc" mgfwc" rnqv"

mqpugtxcuk." ogtdcw" ownck" dgtdwpic" rcfc"

wowt" 6"v cjwp" fcp" cpcmcp" cnco" fklworck"

rcfc"wowt"8"vcjwp0"

D. Analisis Data"

Fcvc" jcukn" rgpiwmwtcp" vkpiik" fcp"

fkcovgt" fkcpcnkuku" fgpicp" ogpiiwpcmcp"

cpcnkuku" xctkcp" wpvwm" ogpigvcjwk" xctkcuk"

cpvct" rtqxgpcp0" Cpcnkuku" xctkcp" jcp{c"

fkncmwmcp" wpvwm" jcukn" rgpiwmwtcp" vkpiik"

fcp"fkcogvgt"vcpcocp"rcfc"rnqv"konservasi

ex situ"ogtdcw"fk"Dqpfqyquq"rcfc"wowt"3"

fcp"4"vcjwp0"Cpcnkuku"xctkcp"vkfcm"fkncmwmcp"

rcfc" rnqv" mqpugtxcuk" ex situ" ogtdcw" fk"

Rgvcm" ;5."I wpwpi" Mkfwn0" " Rcfc" rnqv"

mqpugtxcuk" ex situ" ogtdcw" fk" Rgvcm" ;5"

Iwpwpi" Mkfwn" ugvkcr" rtqxgpcp" fkvcpco"

fcnco" ucvw" dnqm" vgtugpfktk" fcp" mqpfkuk"

rgpwvwrcp" ncjcp" cpvct" dnqm" vkfcm" ucoc0"

Oqfgn"ocvgocvku"{cpi"fkiwpcmcp"cfcncj

Jurnal Pemuliaan Tanaman Hutan

Xqn"9"Pq0"5."Pqxgodgt"4235."39;"/"3;8""

3:6"

Yij = - + Bi + Pj + iij"

fkocpc"

Cfcrvcdknkvcu."Rgtvwodwjcp"fcp"Tgigpgtcuk""Rcfc"Rnqv"Mqpugtxcuk"Gz"Ukvw"Ogtdcw"Vtk"Rcowpimcu"[wfqjctvqpq"fcp"Dwtjcp"Kuockn"

3:7"

ogtdcw0" Rcfc" ugdctcp" cncokp{c" ogtdcw"

fklworck"vwodwj"jkpiic"mgvkpiikcp"3222"o"

frn"rcfc"dgtdcick"lgpku"vcpcj"mgewcnk"vcpcj"

icodwv"fgpicp"vkrg"kmnko"C/F"*Ocjhwf¦"et

al0."4228+0"Jwvcp"Rgpgnkvkcp"Uwodgtytkpikp"

{cpi" vgtngvcm" " rcfc" mgvkpiikcp"ł" :22"o "fk"

cvcu" rgtowmccp" ncwv"ogorwp{ck" vkrg" kmnko"

D" fcp" lgpku" vcpcj" cuqukcuk" cpfquqn" eqmncv0"

Jwvcp" Rgpgnkvkcp" Rgvcm" ;5" {cpi" vgtngvcm"

rcfc" mgvkpiikcp" ł" " 372" o" frn" ogoknkmk" "

kmnko" E" fcp" lgpku" vcpcj" xgtvkuqn" fgpicp"

mqpfkuk" vcpcj" ugdcikcp" dgtdcvw0" Jcukn"

rgpgnkvkcp" [wfqjctvqpq" *422:+"

ogpwplwmmcp" dcjyc" tcvc/tcvc" pknck"

mgtcicocp" igpgvkm" ogtdcw" fctk" mggpco"

rqrwncuk" ogtdcw" *Jcnocjgtc" Vkowt."

Yckiq." Qtcpudctk." Ycukqt." Pcdktg." fcp"

Ugtco+"vgtiqnqpi"vkpiik"{ckvw"2.5;40""Tcvc/

tcvc" mgtcicocp" igpgvkm" I. bijuga" kpk" ngdkj"

vkpiik" " fkdcpfkpimcp" mgtcicocp" igpgvkm"""""

I. palembanica {cmpk"ugdguct"2.464"""*"Ngg."

et al.,"4224+." ugowc" lgpku" vcpcocp" *2.335+."

vcpcocp" vtqrku" *2.433+" fcp" vcpcocp" mc{w"

dgtwowt" rcplcpi" *2.36;=" Jcotkem." et al.,"

3;;3+0" Mgtcicocp" igpgvkm" ogtwrcmcp"

mqorqpgp" {cpi" ucpicv" ogpgpvwmcp"

mgocorwcp" uwcvw" lgpku" wpvwm" fcrcv"

dgtcfcrvcuk" vgtjcfcr"rgtwdcjcp" nkpimwpicp"

fcnco" lcpimc"rcplcpi0" Lgpku" vcpcocp" {cpi"

ogorwp{ck"mgtcicocp"igpgvkm" vkpiik"cmcp"

ogorwp{ck" rgnwcpi" {cpi" ugocmkp" dguct"

wpvwm"vwodwj"fcp"dgtmgodcpi"fcnco"uwcvw"

nkpimwpicp"vwodwj"{cpi"cfc0"

B. Pertumbuhan tinggi dan diameter

Jcukn" rgpicocvcp" mctcmvgtkuvkm"

rgtvwodwjcp" "v cpcocp" ogtdcw" rcfc" rcfc"

rnqv" mqpugtxcuk" ex situ" fk" JR"

Uwodgtytkpikp." Dqpfqyquq" fcp" Rgvcm" ;5."

Iwpwpi" Mkfwn" rcfc" wowt" 8" fcp" 34" dwncp"

fkuclkmcp"rcfc"Vcdgn"5"fcp"60"

"Vcdgn"50"Vkpiik"vcpcocp"ogtdcw"rcfc"rnqv"mqpugtxcuk"ex situ"fk"JR"Uwodgtytkpikp."Dqpfqyquq"fcp"

Rgvcm";5."Iwpwpi"Mkfwn"rcfc"wowt"34"fcp"46"dwncp""Pq" Rqrwncuk" Wowt"34"dwncp"*eo+" Wowt"46"dwncp"*eo+"" " Dqpfqyquq" Iwpwpi"Mkfwn" Dqpfqyquq" Iwpwpi"Mkfwn"3" Jcnocjgtc"Vkowt" 97.94" ;7.44" ;8.63" 328.99"4" Yckiq" :9.99" ;;.8:" 324.:;" 327.37"5" Pcdktg" :5.39" 329.89" ;6.43" 365.;7"6" Ugtco" 6:.;9" :2.;:" 7;.;5" 323.6;"7" Qtcpudctk" :2.92" 327.39" ;8.:8" 362.3;"8" Ycukqt" ;3.:5" ;3.24" 332.77" 333.62"9" Ocpfqrk1Vycpycyk" 9;.56" 327.67" 328.8:" 35;.22"

""""""

Jurnal Pemuliaan Tanaman Hutan

Xqn"9"Pq0"5."Pqxgodgt"4235."39;"/"3;8""

3:8"

Vcdgn"60"Fkcogvgt" vcpcocp"ogtdcw"rcfc"rnqv"mqpugtxcuk"ex situ"fk"JR"Uwodgtytkpikp."Dqpfqyquq"fcp"Rgvcm";5."Iwpwpi"Mkfwn"rcfc"wowt"34"fcp"46"dwncp"

"Pq" Rqrwncuk" Wowt"34"dwncp"*oo+" Wowt"46"dwncp"*oo+"" " Dqpfqyquq" Iwpwpi"Mkfwn" Dqpfqyquq" Iwpwpi"Mkfwn"3" Jcnocjgtc"Vkowt" 32.48" 33.53" 36.83" 35.72"4" Yckiq" ;.8:" 3.:6" 35.86" 34.7:"5" Pcdktg" ;.6;" 32.;5" 34.24" 37.94"6" Ugtco" 9.22" ;.45" ;.25" 54.2:"7" Qtcpudctk" ;.:5" 33.53" 35.53" 36.69"8" Ycukqt" 32.3;" 33.73" 35.;9" 56.73"9" Ocpfqrk1Vycpycyk" 32.84" 32.;2" 36.2;" 36.72"

"Fctk" Vcdgn" 5"fc p" 6"fk mgvcjwk" dcjyc"

tcvc/tcvc" pknck" mctcmvgtkuvkm" rgtvwodwjcp"

vcpcocp" ogtdcw" dckm" vkpiik" fcp" fkcogvgt""

wpvwm"ugvkcr""rtqxgpcp"rcfc"rnqv"mqpugtxcuk"

ex situ" Rgvcm" ;5."Iwpwpi"Mkfwn" ngdkj" dckm"

fkdcpfkpimcp" JR" Uwodgtytkpikp." Dqpfq/

yquq" rcfc" ugowc" wowt" rgpicocvcp0" Jcn"

kpk"fkfwic"fkrgpictwjk"qngj"mqpfkuk" vgorcv"

vwodwj" rcfc" mgfwc" rnqv" vgtugdwv0" Rnqv"

mqpugtxcuk" ex situ" fk" Dqpfqyquq" vgtngvcm"

rcfc" mgvkpiikcp" vgtngvcm" "rc fc" mgvkpiikcp"""

ł":22"o"fk"cvcu"rgtowmccp"ncwv"fgpicp"vkrg"

kmnko" D" fcp" " rnqv" mqpugtxcuk" ex situ" fk"

Iwpwpi" Mkfwn" vgtngvcm" rcfc" mgvkpiikcp" ł""

372"o"frn"fgpicp"vkrg"kmnko"E0" Kmnko"cmcp"

dgtwdcj" fgpicp" rgtdgfccp" mgvkpiikcp"

vgorcv0"Mgvkpiikcp"vgorcv"ogorwp{ck"ghgm"

vkfcm" ncpiuwpi" vgtjcfcr" tkcr" fcp" dgpvwm"

rqjqp" jwvcp0" Rgtvwodwjcp" vkpiik" vcpcocp"

fcp" tkcr" vcpcocp" cmcp" ogpwtwp" ugktkpi"

fgpicp" dgtvcodcjp{c" mgvkpiikcp" vgorcv"

*Uqgvtkupq."3;;:+0""

Dgtfcuctmcp" fcvc" rcfc" mgfwc" vcdgn" fk"

cvcu" lwic" vgtnkjcv" cfcp{c" xctkcuk" mctcmvgt"

rgtvwodwjcp" vkpiik" fcp" fkcogvgt" cpvct"

rtqxgpcp" ogtdcw" {cpi" fkwlk0" Wpvwm"

ogpigvcjwk" xctkcuk" cpvct" rtqxgpcp" wpvwm"

ukhcv" vkpiik" fcp" fkcogvgt" ocmc" fkncmwmcp"

cpcnkuku" xctkcp" vgtjcfcr" fcvc" rgpiwmwtcp"

vkpiik" fcp" fkcogvgt0" Cpcnkuku" xctkcp" vkfcm"

fkncmwmcp" rcfc" rnqv" mqpugtxcuk" ex situ"

ogtdcw" fk" Rgvcm" ;5."Iwpwpi"Mkfwn0" " Rcfc"

rnqv"mqpugtxcuk"ex situ"ogtdcw"fk"Rgvcm";5"

Iwpwpi" Mkfwn" ugvkcr" rtqxgpcp" fkvcpco"

fcnco" ucvw" dnqm" vgtugpfktk" fcp" mqpfkuk"

rgpwvwrcp" ncjcp" cpvct" dnqm" vkfcm" ucoc0"

Jcukn" cpcnkuku" xctkcp" wpvwm" ukhcv" vkpiik" fcp"

fkcogvgt" rcfc" rnqv" mqpugtxcuk" ex situ"

ogtdcwfk" Dqpfqyquq" rcfc" wowt" 3."f cp" 4"

vcjwp"fkuclkmcp"rcfc"Vcdgn"7"fcp"80"

""""""

Cfcrvcdknkvcu."Rgtvwodwjcp"fcp"Tgigpgtcuk""Rcfc"Rnqv"Mqpugtxcuk"Gz"Ukvw"Ogtdcw"Vtk"Rcowpimcu"[wfqjctvqpq"fcp"Dwtjcp"Kuockn"

3:9"

Vcdgn"70"Cpcnkuku"xctkcp"wpvwm"ukhcv"vkpiik"vcpcocp"rcfc"rnqv"mqpugtxcuk"ex situ"ogtdcwfk"Dqpfqyquq"rcfc"wowt"3."fcp"4"vcjwp"

"Wowt"3"Vcjwp"

Uwodgt" Fgtclcf" Lwoncj" Mwcftcv" H" Uki0"Xctkcuk" Dgdcu" Mwcftcv" Vgpicj" " "

Rtqxgpcp" 7" 5728.45" 923.47" 323.4;,," 2.222"Dnqm" 4" 429.22" 325.72" 36.;7,," 2.223"Gttqt" 32" 8;.45" 8.;4" "" ""

Wowt"4"Vcjwp"Uwodgt" Fgtclcf" Lwoncj" Mwcftcv" H" Uki0"Xctkcuk" Dgdcu" Mwcftcv" Vgpicj" " "

Rtqxgpcp" 7" 6798.89" ;37.55" 49.22,," 2.222"Dnqm" 4" 85.;6" 53:.69" ;.62,," 2.227"Gttqt" 32" 55:.;9" 55.:;" "" ""

"Vcdgn"80"Cpcnkuku"xctkcp"wpvwm"ukhcv"fkcogvgt"vcpcocp"rcfc"rnqv"mqpugtxcuk"ex situ""""""""""""""""""ogtdcw"fk"Dqpfqyquq"rcfc"wowt"3."fcp"4"vcjwp""

Wowt"3"Vcjwp"Uwodgt" Fgtclcf" Lwoncj" Mwcftcv" H" Uki0"Xctkcuk" Dgdcu" Mwcftcv" Vgpicj" " "

Rtqxgpcp" 7" 44.2:" 6.64" 39.69,," 2.222"Dnqm" 4" 3.43" 2.82" 4.5:pu" 2.364"Gttqt" 32" 4.75" 2.47" "" ""

Wowt"4"Vcjwp"Uwodgt" Fgtclcf" Lwoncj" Mwcftcv" H" Uki0"Xctkcuk" Dgdcu" Mwcftcv" Vgpicj" " "

Rtqxgpcp" 7" 77.54" 33.28" :.5:,," 2.224"Dnqm" 4" 3.6:" 2.96" 2.78 2.7;2"pu"Gttqt" 32" 35.42" 3.54" "" ""

Mgvgtcpicp",,""?"dgtdgfc"p{cvc"rcfc"vctch"wlk"3'"" """"""""pu""?"vkfcm"dgtdgfc"p{cvc""

"

Fctk"Vcdgn"5"fkmgvcjwk"dcjyc"vcpcocp"

ogtdcw" rcfc" mgfwc" rnqv" mqpugtxcuk"ex situ"

{cpi"ogoknkmk"tcvc/tcvc"vkpiik"vgtgpfcj"fcp"

vgtvkpiik"rcfc"wowt"34"fcp"46"dwncp"dgtcucn"

fctk" rtqxgpcp" {cpi" ucoc0" Rcfc" rnqv"

mqpugtxcuk" ex situ" fk" JR" Uwodgtytkpikp."

Dqpfqyquq." tcvc/tcvc" vkpiik" vgtgpfcj"

dgtcucn" fctk" rtqxgpcp" Ugtco" fcp" vgtvkpiik"

fctk"rtqxgpcp"Ycukqt0"Rcfc"rnqv"mqpugtxcuk"

ex situ"fk"Rgvcm";5."Iwpwpi"Mkfwn."tcvc/tcvc"

vkpiik"vgtgpfcj"dgtcucn"fctk"rtqxgpcp"Ugtco"

fcp" vgtvkpiik" fctk" rtqxgpcp" Pcdktg0" Fctk"

Vcdgn" 6" fkmgvcjwk" dcjyc" vcpcocp" ogtdcw"

rcfc" mgfwc" rnqv" mqpugtxcuk" ex situ" {cpi"

ogoknkmk" tcvc/tcvc" fkcogvgt" vgtgpfcj" rcfc"

wowt"34"fcp"46"dwncp"dgtcucn"fctk"rtqxgpcp"

{cpi" ucoc" {ckvw" rtqxgpcp" Ugtco0"

Ugfcpimcp" tcvc/tcvc" fkcogvgt" vgtvkpiik"

dgtcucn"rtqxgpcp"{cpi"dgtdgfc0"Fctk"Vcdgn"

7"fcp"8"fkmgvcjwk"dcjyc"vgtfcrcv"rgtdgfccp"

Jurnal Pemuliaan Tanaman Hutan

Xqn"9"Pq0"5."Pqxgodgt"4235."39;"/"3;8""

3::"

{cpi" p{cvc" fkcpvctc" rtqxgpcp" {cpi" fkwlk"

wpvwm"ukhcv"vkpiik""fcp"fkcogvgt"vcpcocp"fk"

rnqv" mqpugtxcuk" ex situ" ogtdcw" fk" JR"

Uwodgtytkpikp." Dqpfqyquq" rcfc" wowt" 3"

fcp"4"vcjwp0"Rgtdgfccp"vgtugdwv"ogpwplwm/

mcp" dcjyc" rtqxgpcp" ogodgtkmcp" rgpic/

twj" vgtjcfcr" mctcmvgtkukvkm" vkpiik" fcp"

fkcogvgt"vcpcocp"ogtdcw"rcfc"wowt"3"fcp"

4"vcjwp0"""

Rtqxgpcp" Ugtco" ugncnw" ogoknkmk" pknck"

tcvc/tcvc"vgtgpfcj"wpvwm"mctcmvgtkuvkm"vkpiik"

fcp"fkcogvgt"rcfc"ugowc"wowt"rgpicocvcp"

fk"mgfwc"rnqv" " mqpugtxcuk"ex situ " vgtugdwv0"

Jcukn" wlk"Fwpecp"wpvwm"mctcmvgtkuvkm" vkpiik"

fcp" fkcogvgt" mggpco" rtqxgpcp" ogtdcw"

*Jcnocjgtc" Vkowt." Yckiq." Qtcpudctk."

Ycukqt." Pcdktg." fcp" Ugtco+" rcfc" wowt"""""""

3" jkpiic" 5"v cjwp" rcfc" rnqv" mqpugtxcuk" ex

situ" fk" JR" Uwodgtytkpikp." Dqpfqyquq"

lwic" ogpwplwmmcp" dcjyc" rtqxgpcp" Ugtco"

ogodgpvwm" mgnqorqm" vgtugpfktk" fcp"

vgtrkucj" fctk" rtqxgpcp" nckp" *[wfqjctvqpq."

4234+0" Jcn" kpk" lwic" fkrgtmwcv" fctk" jcukn"

cpcnkuku" kuq¦ko" vgtjcfcr" mggpco" rtqxgpcp"

vgtugdwv0" [wfqjctvqpq" *422:+" ogp{cvcmcp"

dcjyc" jcukn" cpcnkuku" mncuvgt" WRIOC""

dgtfcuctmcp" " Nei’s standard genetic

distance" *3;94+" ogpwplwmmcp" dcjyc"

mggpco" rqrwncuk" ogtdcw" ogodgpvwm" vkic"

mncuvgt" wvcoc0" Rqrwncuk"Jcnvko" fcp"Yckiq"

ogodgpvwm" mncuvgt" rgtvcoc." rqrwncuk"

Pcdktg." Qtcpudctk" fcp" Ycukqt" ogodgpvwm"

mncuvgt" mgfwc0" Rqrwncuk" Ugtco"ogodgpvwm""

mncuvgt"mgvkic"{cpi"lcwj"vgtrkucj"fctk"mgfwc"

mncuvgt" nckpp{c0" Rqnc" rgpignqorqmcp" kpk"

fkfwic" fkrgpictwjk" qngj" uglctcj"

vgtdgpvwmp{c" rwncw" Ugtco0" Ogpwtwv"

Yjkvoqtg" *3;:9+." rcfc" cycnp{c"

Jcnocjgtc." Ugtco" fcp" Rcrwc" ogtwrcmcp"

ucvw" fctcvcp" fctk" fctcvcp" dguct" Cwuvtcnkc/

Pgy" Iwkpgc" {cpi" ogtwrcmcp" dcikcp" fctk"

Australian Gondwanaland0" Rgtdgfccp"

ycmvw" rgokucjcp" Ugtco" fcp" Jcnocjgtc"

fctk" fctcvcp" dguct" Cwuvtcnkc/Pgy" Iwkpgc"

ogpicmkdcvmcp" rgtdgfccp" uvtwmvwt" igpgvkm"

{cpi"lcwj"fk"cpvctc"mgfwcp{c0"Rwncw"Ugtco"

fkfwic" vgtrkucj"fctk"fctcvcp"Cwuvtcnkc/Pgy"

Iwkpgc" lcwj" ngdkj" fcjwnw" fkdcpfkpimcp"

Jcnocjgtc0"Rwncw"Ugtco"{cpi"vgtngrcu"lcwj"

vgtngdkj" fcjwnw" fcp" fkrkucjmcp" qngj" ncwvcp"

ogpicncok" kuqncuk" tgrtqfwmuk" fcp" cfcrvcuk"

nqmcn" vgtjcfcr" nkpimwpicp" ugvgorcv" {cpi"

ogp{gdcdmcp" ugocmkp" ogpkpimcvp{c"

fkhgtgpukcuk"igpgvkm"cpvctc"rqrwncuk"ogtdcw"

fk"Ugtco"fcp"rqrwncuk/rqrwncuk"ogtdcw"fctk"

Jcnocjgtc" fcp" Rcrwc0" Ogpwtwv" Nqxgnguu"

fcp" Jcotkem" *3;:6+." fkhgtgpukcuk" igpgvkm"

cpvct rqrwncuk" fkrgpictwjk" qngj" cfcp{c"

cnktcp" igp" ogncnwk" rgp{gdctcp" ugtdwm" uctk"

fcp" dklk0" Urgukgu" fgpicp" rqrwncuk"

fkumqpvkp{w" fcp" vgtkuqncuk" ogpwplwmmcp"

mgpckmcp"vkpimcv"fkhgtgpukcuk"igpgvkm"mctgpc"

vwtwpp{c" cnktcp" igp0" Rgtdgfccp" uvtwmvwt"

igpgvkm" vgtugdwv" fcrcv" fkgmurtgukmcp"

ogncnwk" rgtdgfccp" mctcmvgtkuvkm" " vcpcocp0"

Ugtco" ogtwrcmcp" rtqxgpcp" {cpi" wpkm"

mctgpc" rcfc" nqmwu" vgtvgpvw" cfc" cngn" {cpi"

jcp{c" fkoknkmk" rqrwncuk" Ugtco" fcp" vkfcm"

Cfcrvcdknkvcu."Rgtvwodwjcp"fcp"Tgigpgtcuk""Rcfc"Rnqv"Mqpugtxcuk"Gz"Ukvw"Ogtdcw"Vtk"Rcowpimcu"[wfqjctvqpq"fcp"Dwtjcp"Kuockn"

3:;"

fkoknkmk" qngj" mgnkoc" rqrwncuk" {cpi" nckp"

*[wfqjctvqpq." 422:+0" Ogpwtwv" ¥qdgn."

et.al.," *3;98+" mgdcp{cmcp" lgpku" vcpcocp"

jwvcp" ogorwp{ck" ucvw" ucorck" dgdgtcrc"

rtqxgpcp" {cpi" ogoknkmk" ukhcv" igpgvkm"

rgpvkpi" {cpi" dgtukhcv" wpkm" ucvw" ucoc" nckp"

fcp" ugdcnkmp{c0"Qngj" mctgpc" kvw." rgmgtlccp"

dguct" rgtvcoc" fcnco" mqpugtxcuk" mqorngmu"

igp" dcik" rctc" rgownkc" vcpcocp" cfcncj"

ogp{gncocvmcp" ukhcv" {cpi" wpkm" fctk" uwcvw"

rtqxgpcp0" Fgpicp" cfcp{c" xctkcuk" igpgvkm"

{cpi" vkpiik" cpvct" rtqxgpcp" ogtdcw" ocmc"

ugocmkp" dguct" lwic" rqvgpuk" fctk" uwodgt"

fc{c" igpgvkm" ogtdcw" {cpi" dkuc"

fkugncocvmcp" fcp" fkocphccvmcp" wpvwm"

ogpfwmwpi" rtqitco" rgownkccp" lgpku"

ogtdcw0""

C. Regenerasi

Tgigpgtcuk" uwodgtfc{c" igpgvkm"

ogtwrcmcp" dcikcp" {cpi" rgpvkpi" fctk"

uvtcvgik"mqpugtxcuk"mctgpc"dgtjcuknp{c"uwcvw"

rtqitco" mqpugtxcuk" jcp{c" fcrcv" fkpknck"

ugvgncj" uwodgtfc{c" vgtugdwv" dgtgigpgtcuk"

*Hkpmgnfc{." 4227+0" Tgigpgtcuk" fkcycnk"

fgpicp" mgocorwcp" uwcvw" vcpcocp" wpvwm"

ogncnwk"ugowc"vcjcrcp"fcnco"rgodwpiccp0"

Rgodwpiccp" rcfc" vcpcocp" jwvcp" {cpi"

dgtwowt" rcplcpi" *rgtgpkcn+" ogtwrcmcp"

rtqugu" {cpi" ucpicv" mqorngmu0" Jcn" kpk"

fkugdcdmcp" cfcp{c" kpvgtcmuk" vcpcocp"

fgpicp" mqpfkuk" nkpimwpicp" ugrcplcpi"

ycmvw" ugvkcr" vcjwp0" Rtqugu" rgodwpiccp"

rcfc" fcuctp{c" ogtwrcmcp" kpvgtcmuk" hcmvqt"

gmuvgtpcn1nkpimwpicp" *ecjc{c." uwjw."

mgngodcdcp+" fcp" hcmvqt" kpvgtpcn" urgtvk"

igpgvkm"*Vjqocu."3;;5+0"""

Vcpcocp" ogtdcw" ogoknkmk" dkugmuwcn"

{ckvw" ogorwp{ck" dwpic" lcpvcp" fcp" dgvkpc"

fcnco" ucvw" dwpic0" Dwpic" ogtdcw"

ogorwp{ck" gorcv" mgnqrcm" dwpic" {cpi"

dgtyctpc" mgjklcwcp." ucvw" ocjmqvc" dwpic"

fcp" vkic" dgpcpi" uctk0" Ocjmqvc" dwpic"

dgtyctpc" rwvkj" *fgpicp" dcikcp" vgpicj"

dgtyctpc" ogtcj+" cvcw" ogtcj" owfc"

ogpfgmcvk"wpiw"cvcw"ogtcj0"Dwpic"vgtngvcm"

fkdcikcp" cmukngt0" Rqnkpcvqt" wvcoc" dwpic"

ogtdcw" cfcncj" ugtcpiic" *ngdcj+." dwtwpi"

fcp" cpikp" *Vjcocp." et al0." 4228+0" Jcukn"

rgpicocvcp" rcfc" rnqv" mqpugtxcuk" ex situ"

ogtdcw" "fk"Iwpwpi"Mkfwn"fcp"Dqpfqyquq"

ogpwplwmmcp" dcjyc" dwpic" ogtdcw"

vgtocuwm" dwpic" jgvgtquvknk" {ckvw" dwpic"

{cpi" ogoknkmk" vcpimck" rwvkm" *uv{nwu+" fcp"

vcpimck" uctk" *hkncogp+" {cpi" vkfcm" ucoc"

rcplcpi0" Rcfc" dwpic"ogtdcw" vcpimck" rwvkm"

ngdkj"rcplcpi"fctk"vcpimck"uctk"*Icodct"3+0" "

Vkrg" dwpic" jgvgtquvknk" ogowpimkpmcp"

vgtlcfkp{c""rgp{gtdwmcp"vgtdwmc"cvcw"mcykp"

ukncpi0"Pknck" tgtcvc"rgtmcykpcp" ukncpi"rcfc"

ownvk"nqmwu"*vo+"wpvwm"rqrwncuk"ogtdcw"fctk"

Ocpqmyctk" fcp"Mgtqo" ucoc" {ckvw" ugdguct"

3.2220"Pknck""tgtcvc"rgtmcykpcp"ukncpi"rcfc"

uwcvw" nqmwu" *vu+" wpvwm"rqrwncuk"Ocpqmyctk"

fcp" Mgtqo" ocukpi/ocukpi" " 2.;:3" fcp""

2.;940" "P knck" " vo" " fcp" " vu" " {cpi" " dguct""

ogpwplwmmcp" dcjyc" mgowpimkpcp"

vgtlcfkp{c" mcykp" ukncpi" nwct" rcfc" mgfwc"

Jurnal Pemuliaan Tanaman Hutan

Xqn"9"Pq0"5."Pqxgodgt"4235."39;"/"3;8""

3;2"

rqrwncuk" vgtugdwv" ucpicv" dguct0" "P knck" vo/vu"

*biparental inbreeding+" wpvwm" rqrwncuk"

Ocpqmyctk" fcp" Mgtqo" ocukpi/ocukpi"

ugdguct" 2.23;"fc p" 2.24:0"P knck" kpk"

ogpikpfkmcukmcp" dcjyc" mgowpimkpcp"

vgtlcfkp{c" rgtmcykpcp" mgtcdcv" cfcncj" mgekn"

{ckvw"4.:"'"wpvwm"rqrwncuk"Mgtqo"fcp"3.;"

'" wpvwm" rqrwncuk" Ocpqmyctk0" Pknck" tr"

*correlation of paternity+"{cpi"tgpfcj"{ckvw"

2.48;" *Mgtqo+" fcp" 2.369" *Ocpqmyctk+"

ogpikpfkmcukmcp" tgpfcjp{c" rtqdcdknkvcu"

full sib" rcfc" mgvwtwpcp" {cpi" dgtcucn" fctk"

ucvw" rqjqp"* Pkpiukj." 422:+0" Vcpcocp"

ogtdcw" rcfc" mgfwc" rnqv" mqpugtxcuk"

dgtdwpic"rgtvcoc"mcnk"rcfc"wowt"6"vcjwp0""

"""" "Icodct"30"Dwpic"ogtdcw"*vcorcm"ucorkpi+"""""""""""""""""Icodct"40"Dwpic"ogtdcw"*vcorcm"cvcu+"

"

Rgodwpiccp"ogtdcw"fk"rnqv"mqpugtxcuk"

ex situ" " fk" Iwpwpi" Mkfwn" fcp" Dqpfqyquq"

ngdkj" egrcv" fkdcpfkpimcp" Ectkvc0" " Ogtdcw"

{cpi" fkvcpco" fk" mgdwp" rgteqdccp" nkvdcpi"

Ectkvc" dgtdwpic" rcfc" wowt" 32"v cjwp0"

Dwpic" ogtdcw" dgtwrc" dwpic" oclgowm"

fcnco" dgpvwm" ocnck." vcpimck" wvcoc""""""""""""

7/3:" eo" fcp" rcplcpi" vclwm" dwpic" 3.7/4.7"

eo0"Dwcj"ogtdcw"dgtdgpvwm"rqnqpi."dwncv."

cvcw"dgtdgpvwm"cicm"rcplcpi0"Dgpkj"ogtdcw"

dgtdgpvwm" dwncv" rkrkj" fcp" dgtyctpc" eqmncv"

vwc" mgogtcj/ogtcjcp0" Dwpic" ogmct" rcfc"

dwncp" Pqxgodgt" ucorck" Lcpwctk" fcp" dwcj"

vwc" rcfc" dwncp"Ogk" ucorck" Ciwuvwu" Dwcj"

ukcr" fkrcpgp" ugvgncj" ocucm" hkukqnqiku"

fgpicp" ektk/ektk" yctpc" dwcj" eqmncv" vwc"

ucorck" mgjkvco/jkvcocp." mwnkv" dwcj" mgtcu"

fcp" dgpkj" dgtyctpc" eqmncv" vwc" mgogtcjcp"

*[wpkctvk."3;;6"fcp"4222+0""

Rtqugu" rgodwpiccp" ogtdcw" rcfc" rnqv"

mqpugtxcuk" ex situ" fk" Iwpwpi" Mkfwn" fcp"

Dqpfqyquq" fkownck" rcfc" owuko"

rgpijwlcp0" Rcfc" rnqv" mqpugtxcuk" ex situ" fk"

Iwpwpi" Mkfwn." mwpewr" dwpic" owpewn" fcp"

ogmct" ugncoc" owuko" rgpijwlcp"

*Pqxgodgt"ucorck"rgtvgpicjcp"Crtkn+"cmcp"

ogodwuwm." tqpvqm" fcp" vkfcm" fcrcv"

dgtmgodcpi" ogplcfk" rqnqpi0" Dwpic" {cpi"

ogmct" rcfc" uccv" fkownckp{c" owuko"

mgoctcw" *cmjkt"Crtkn"cvcw"Ogk+"cmcp"fcrcv"

dgtmgodcpi" ogplcfk" rqnqpi" dwcj0" Rqnqpi"

dwcj" ogtdcw" cmcp" ocucm" cvcw" vwc" rcfc"

stylus ""filamen

mahkota

kelopak

Cfcrvcdknkvcu."Rgtvwodwjcp"fcp"Tgigpgtcuk""Rcfc"Rnqv"Mqpugtxcuk"Gz"Ukvw"Ogtdcw"Vtk"Rcowpimcu"[wfqjctvqpq"fcp"Dwtjcp"Kuockn"

3;3"

dwncp" Ciwuvwu" fcp" rgecj" *over ripe+" rcfc"

rgtvgpicjcp" dwncp" Ugrvgodgt0" Rcfc" rnqv"

mqpugtxcuk"ex situ" fk"Iwpwpi"Mkfwn" rtqugu"

rgodwpiccp" dgtncpiuwpi" ugorwtpc" fcp"

ocorw" ogpijcuknmcp" dklk0" Rqnqpi" dwcj"

ogtdcw" {cpi" uwfcj" ocucm" cvcw" vwc"

dgtyctpc"eqmncv" vwc"jkpiic"jkvco"*Icodct"

5+0" Dklk" ogtdcw" {cpi" uwfcj" vwc" dgtyctpc"

jkvco" fgpicp" dgpvwm" dwncv" rkrkj" cvcw"

rgtugik" rcplcpi" *Icodct" 8+0" Rqnqpi" dwcj"

{cpi" ngycv" ocucm" *over ripe+" ugncplwvp{c"

cmcp"rgecj"*Icodct"6+"fcp""dklk"cmcp"lcvwj0"

Rgp{gdctcp" dklk" ogtdcw" vkfcm" lcwj" fctk"

rqjqp" kpfwmp{c" mctgpc" dklk" ogtdcw" ewmwr"

dgtcv0"Dcjmcp"dklk"ogtdcw"{cpi"lcvwj"uwfcj"

ocorw" dgtmgodcpi" ogplcfk" cpcmcp" cnco"

*Icodct"7+0"

" """""""""""" ""Icodct"50"Dwpic"fcp"rqnqpi"dwcj"ocucm"mqpugtxcuk"""""""""""""""""""""ex situ"ogtdcw"fk"Iwpwpi"Mkfwn""

Icodct"60"Rqnqpi"dwcj"ogtdcw"over ripe "Rcfc"rnqv" rcfc" mqpugtxcuk" ex situ" fk"""Iwpwpi"Mkfwn"

"""""""""""""""""""""""""""""

" "Icodct"70"Dklk"fcp"cpcmcp"ogtdcw"rcfc"rnqv"""""""""""""""""""""Icodct"80"Dgpvwm."yctpc"fcp"wmwtcp"dklk"ogtdcw"""""""""""""""""""mqpugtxcuk"ex situ"fk"Iwpwpi"Mkfwn"""""""""""""""""""""""""""""""""""""""

Rcfc" rnqv" mqpugtxcuk" ex situ" fk"

Dqpfqyquq." mwpewr" dwpic" {cpi" owpewn"

fcp" ogmct" cmcp" ogodwuwm." tqpvqm" fcp"

vkfcm" fcrcv" dgtmgodcpi" ogplcfk" rqnqpi"

dwcj" dckm" ugncoc" owuko" rgpijwlcp"

ocwrwp"mgoctcw"*Icodct"9"fcp":+0""

Jurnal Pemuliaan Tanaman Hutan

Xqn"9"Pq0"5."Pqxgodgt"4235."39;"/"3;8""

3;4"

"""""""""""""""" """""""""""""""""""""""""""""Icodct" 90" Cpvjguku" ogtdcw" ogodwuwm" fcp"

tqpvqm"rcfc"rnqv"mqpugt/xcuk"ex situ"fk"Dqpfqyquq"""""""""""""""""""""""""""""""""""""

"

Icodct" :0"Mwpewr" dwpic"ogtdcw"ogmct"ogodwuwm"fcp" tqpvqm" rcfc" rnqv" mqpugtxcuk" ex situ" fk"Dqpfqyquq"

Rtqugu" rgodwpiccp" ogtdcw" dgnwo"

dgtncpiuwpi" ugorwtpc" fcp" dgnwo" ocorw"

ogpijcuknmcp" dklk" rcfc" rnqv" mqpugtxcuk" ex

situ" fk" Dqpfqyquq0" Vcjcrcp" rgodwpiccp"

{cpi" uwfcj" fkncnwk" vcpcocp" ogtdcw" rcfc" "

rnqv" mqpugtxcuk"ex situ" fk"Dqpfqyquq"dctw"

ucorck" rcfc" vcjcr" cpvjguku0" Mqpfkuk" kpk"

fkfwic" fkrgpictwjk" qngj" hcmvqt" kmnko" fcp"

mqpfkuk"vcpcj"rcfc"mgfwc"rnqv"mqpugtxcuk"ex

situ0" Ewtcj" jwlcp" ogtwrcmcp" ucncj" ucvw"

hcmvqt" kmnko" {cpi" ogorgpictwjk""

rgodwpiccp0" Ewtcj" jwlcp" rcfc" rnqv"

mqpugtxcuk ex situ" fk" Dqpfqyquq" ngdkj"

vkpiik"fkdcpfkpimcp"rnqv"mqpugtxcuk""ex situ"

fk"Iwpwpi"Mkfwn0"Rnqv"mqpugtxcuk"ex situ"fk"

Dqpfqyquq" ogoknkmk" vkrg" kmnko" D" fgpicp"

ewtcj" jwlcp" 4622"o o1vj" fcp" " rnqv"

mqpugtxcuk" ex situ" fk" Iwpwpi" Mkfwn"

ogoknkmk" " vkrg" kmnko"E"fgpicp"ewtcj"jwlcp"

3:22" oo1vj0" Fkucorkpi" kvw." ngdcj" {cpi"

ogtwrcmcp" cigp" rgp{gtdwm" lwic" vkfcm"

dcp{cm"fklworck"rcfc""mqpugtxcuk ex situ"fk"

Dqpfqyquq"{cpi"ogorwp{ck"mqpfkuk"ewcec"

{cpi"ucpicv"dcucj.""

Ugnckp"hcmvqt"kmnko"fcnco"jcn"kpk"ewtcj"

jwlcp." mqpfkuk" vcpcj" rcfc" mgfwc" rnqv"

mqpugtxcuk" lwic" fkfwic" ogorgpictwjk"

mgdgtjcukncp" rtqugu" rgodwpiccp" ogtdcw0"

Ugdcikcp" dguct" vcpcj" rcfc" rnqv" mqpugtxcuk"

ex situ" fk" Iwpwpi" Mkfwn" cfcncj" dgtdcvw"

fgpicp" uqnwo" vcpcj" {cpi" fcpimcn" *mwtcpi"

fctk" 52" eo+0" Mgvgtugfkccp" ckt" fcnco" vcpcj"

ogplcfk" hcmvqt" rgodcvcu" rgtvwodwjcp"

vcpcocp" ogtdcw" fk" Iwpwpi" Mkfwn0"

Ugfcpimcp" rcfc" rnqv" mqpugtxcuk" ex situ" fk"

Dqpfqyquq" uqnwo" vcpcjp{c" ewmwr" fcnco"

*ngdkj" fctk" 72" eo+" fcp" vgtugfkc" ewmwr"

dcp{cm" ckt0" Mgvgtdcvcucp" ckt" fk" Iwpwpi"

Mkfwn"ogodwcv" vcpcocp"ugtkpi"ogpicncok"

egmcocp"*uvtguu+"ckt0"Ogumkrwp"egmcocp"ckt"

Cfcrvcdknkvcu."Rgtvwodwjcp"fcp"Tgigpgtcuk""Rcfc"Rnqv"Mqpugtxcuk"Gz"Ukvw"Ogtdcw"Vtk"Rcowpimcu"[wfqjctvqpq"fcp"Dwtjcp"Kuockn"

3;5"

fcrcv" ogpijcodcv" rgtvwodwjcp" vcpcocp"

ogtdcw" vgvcrk" fk" ukuk" nckp" egmcocp" ckt" kpk"

dkuc" dgtfcorcm" rqukvkh" wpvwm" ogocew"

kpkukcuk" dwpic" vcpcocp" ogtdcw0"

Rgodwpiccp" fk" fcgtcj" vtqrku" ogtwrcmcp"

tgurqp" vgtjcfcr" vwtwpp{c" mgvgtugfkccp" ckt"

fcnco" vcpcj0" Ogpwtwv" Owipkulcj" fcp"

Ugvkcycp" *3;;7+." egmcocp" ckt" {cpi" fkkmwvk"

qngj"jwlcp"ugtkpi"ogtcpiucpi"rgodwpiccp"

vcpcocp"vcjwpcp"vtqrkmc0""""""""""

D. Adaptabilitas

Hkpmgnfg{" fcp" Jcpu" *4229+."

ogp{cvcmcp" dcjyc" wowt" fcp" koqdknkvcu"

vwodwjcp"ogpikornkmcukmcp"dcjyc"ogtgmc"

rgtnw"dgtvcjcp"vgtjcfcr"mqpfkuk"nkpimwpicp"

{cpi" dgtdgfc" ugncoc" jkfwrp{c0" Cfcrvcuk"

vwodwjcp" vgtjcfcr" nkpimwpicpp{c"

ogtwrcmcp" rgtu{ctcvcp1mgdwvwjcp" wpvwm"

fcrcv" dgtvcjcp" jkfwr0" " " Mgdgtcfcrvcukcp"

ogpegtokpmcp" vkpiikp{c" mgocorwcp"

dgtvcjcp" jkfwr" fcp" tgrtqfwmuk" rcfc" uwcvw"

nkpimwpicp"vgtvgpvw0"Dgtfcuctmcp"rctcogvgt"

{cpi" fkcocvk" *rgtugp" jkfwr." rgtvwodwjcp"

vcpcocp" fcp" mgocorwcp" dgtdwpic1"

tgigpgtcuk+" rcfc" mgfwc" rnqv" mqpugtxcuk" gz"

situ." ogtdcw" fcrcv" dgtcfcrvcuk" fgpicp"

ewmwr" dckm" rcfc" mgfwc" rnqv" mqpugtxcuk"

vgtugdwv0" Ugectc" wowo." dgtfcuctmcp" fctk"

dguctcp" pknck" rctcogvgt" vgtugdwv" ocmc"

mgocorwcp" dgtcfcrvcuk" vcpcocp" ogtdcw"

rcfc" rnqv" mqpugtxcuk" gz" situ" fk" Iwpwpi"

Mkfwn"ngdkj"dckm"fkdcpfkpimcp"Dqpfqyquq0""

"

IV. "KESIMPULAN

"30 Vcpcocp" ogtdcw" fctk" vwlwj" rtqxgpcp"

{cpi" fkwlk" ogorwp{ck" mgocorwcp"

dgtcfcrvcuk" fgpicp" ewmwr" dckm" rcfc"

rnqv" mqpugtxcuk" ex situ" " fk" Rgvcm" ;5."

Iwpwpi"Mkfwn"fcp""JR"Uwodgtytkpikp."

Dqpfqyquq0" Mguguwckcp" rgtu{ctcvcp"

vwodwj" cpvctc" jcdkvcv" cnco" ogtdcw"

fgpicp" mqpfkuk" nkpimwpicp" rcfc" mgfwc"

rnqv"mqpugtxcuk ex situ""ogtdcw"vgtugdwv"

fcp" vkpiikp{c" mgtcicocp" igpgvkm"

ogtdcw"vgncj"ogodgtkmcp"rgtcpcp"{cpi"

dguct" vgtjcfcr" mgocorwcp" dgtcfcrvcuk"

vgtugdwv0"

40 Rgtdgfccp" rtqxgpcp" fcp" mctcmvgtkuvkm"

vgorcv" vwodwj" rcfc" mgfwc" rnqv"

mqpugtxcuk ex situ" " ogtdcw" lwic"

ogodgtkmcp"rgpictwj"vgtjcfcr"rgtvwo/

dwjcp" fcp" tgigpgtcuk" ogtdcw0" Cfcp{c"

xctkcuk"rgtvwodwjcp"vkpiik"fcp"fkcogvgt"

fkcpvctc" rtqxgpcp" {cpi" fkwlk" rtqxgpcp"

ogpikpfkmcukmcp" dcjyc" rtqxgpcp"

ogodgtkmcp" rgpictwj" vgtjcfcr" mctcm/

vgtkukvkm" vkpiik" fcp" fkcogvgt" vcpcocp"

ogtdcw0" Rgtvwodwjcp" vkpiik" fcp"

fkcogvgt" vcpcocp" ogtdcw" rcfc" rnqv"

mqpugtxcuk" ex situ" Rgvcm" ;5."I wpwpi"

Mkfwn" ngdkj" dckm" fkdcpfkpimcp" JR"

Uwodgtytkpikp." Dqpfqyquq" rcfc"

ugowc" wowt" rgpicocvcp0" Tgigpgtcuk"

ogtdcw"uwfcj"dgtncpiuwpi"fgpicp"dckm"

fcp"dgtjcukn"rcfc"rnqv"mqpugtxcuk"ex situ"

Rgvcm" ;5."I wpwpi" Mkfwn0" Ugfcpimcp"

Jurnal Pemuliaan Tanaman Hutan

Xqn"9"Pq0"5."Pqxgodgt"4235."39;"/"3;8""

3;6"

rcfc" rnqv" mqpugtxcuk" ex situ" JR"

Uwodgtytkpikp."Dqpfqyquq."tgigpgtcuk"

ogtdcw" dgnwo" dgtjcukn" cvcw" vkfcm"

dgtncpiuwpi"fgpicp"dckm""

V. UCAPAN TERIMA KASIH

Rgpwnku" ogpiwecrmcp" vgtkoc" mcukj" mgrcfc"

ugowc" rkjcm" {cpi" ogodcpvw" mgncpectcp"

rgpgnkvkcp" kpk." mjwuwup{c" mgrcfc" Dcrcm"

Fktq"Gmq"Rtcoqpq."U0Jwv0V"fcp"{cpi"vgncj"

ogodcpvw" fcnco"mgikcvcp" rgpiwmwtcp" fcp"

gpvt{"fcvc"0"

"

DAFTAR PUSTAKA

Hkpmgnfg{." T" cpf" Jcvvgogt." J0J0" 42290"Vtqrkecn"Hqtguv"Igpgvkeu0"Urtkpigt/Xgtnci"Dgtnkp"Jgkfgndgti0"

Hkpmgnfg{."T0"42270"Cp"kpvtqfwevkqp"vq"vtqrkecn"hqtguv" igpgvkeu0" Kpuvkvwvg" qh" Hqtguv"Igpgvkeu" cpf" Hqtguv" Vtgg" Dtggfkpi."Igqti/Cwiwuv/Wpkxgtukv{" I…vvkpigp."D¯uigpygi" 4."F /59299" I…vvkpigp."Igtocp{0"

Ikpycn."J0U"cpf"C0M0"Ocpfcn0"42260"Xctkcvkqp"kp" Itqyvj" Rgthqtocpeg" qh" Acacia nelotica" "Yknnf0" Gz" Fgn0" Rtqxgpcpegu" qh"Yknf" Igqitcrjkecn" Qtkikp"

Cfcrvcdknkvcu."Rgtvwodwjcp"fcp"Tgigpgtcuk""Rcfc"Rnqv"Mqpugtxcuk"Gz"Ukvw"Ogtdcw"Vtk"Rcowpimcu"[wfqjctvqpq"fcp"Dwtjcp"Kuockn"

3;7"

Yjkvoqtg." V0E" *gf+0" 3;:90" Biogeographical evolution of the Malay Archipelago0"Qzhqtf" Hqtguvt{" Kpuvkvwvg." Wpkxgtukv{" qh"Qzhqtf0" ENCTGPFQP" RTGUU/QZHQTF""

[wfqjctvqpq."V0R0""422:0""Uvwfk"Xctkcuk"Igpgvkm""Dgdgtcrc"Rqrwncuk""Ogtdcw"*Intsia"Bijuga"Q0Mv¦g+" " Ogpiiwpcmcp" Rgpcpfc"Kuqgp¦ko" fcp" Rgocphccvcpp{c" fcnco"Rtqitco" Mqpugtxcuk" Igpgvkm0" Vguku"Rtqitco" Uvwfk" Know" Mgjwvcpcp" Lwtwucp."Hcmwnvcu" Mgjwvcpcp." Wpkxgtukvcu" Icflcj"Ocfc."[qi{cmctvc0"

[wfqjctvqpq." V0R0" fcp" Dwtjcp." K0" 42340"""Xctkcuk" Igpgvkm" Wlk" Rtqxgpcp" Ogtdcw"Ucorck"Wowt"Vkic"Vcjwp"fk"Dqpfqyquq."Lcyc" Vkowt0" " Lwtpcn" Rgpgnkvkcp" Jwvcp"Vcpcocp"Xqn08" Pq03." Lwnk" 42340" "" Dcnck"Dguct" Rgpgnkvkcp" Dkqvgmpqnqik" fcp"Rgownkccp" Vcpcocp" Jwvcp0" Dcfcp"Rgpgnkvkcp"fcp"Rgpigodcpicp"Mgjwvcpcp."Fgrctvgogp"Mgjwvcpcp0"

[wpkctvk." P0" 42220" Ogtdcw" *Intsia" urr+0" Cvncu"dgpkj"vcpcocp"jwvcp"Kpfqpgukc0"Rwdnkmcuk"Mjwuwu"Xqn0" 4"""P q0" 50" Dcnck" Vgmpqnqik"Rgtdgpkjcp0" Dcfcp" Nkvdcpi" Mgjwvcpcp"fcp"Rgtmgdwpcp0"Dqiqt0"

[wpkctvk." P0" 3;;60" Vgmpkm" Dwfkfc{c" Ogtdcw"*Intsia" urr+" wpvwm" Rgodcpiwpcp" Jwvcp"Vcpcocp0" Mnqp0" Kphqtocuk" fcp"Mqowpkmcuk" Dgpkj" Xqn" 4"P q0230" Dcnck"Rgpgnkvkcp" fcp" Rgpigodcpicp" Vgmpqnqik"Rgtdgpkjcp0" Dcfcp" Nkvdcpi" Mgjwvcpcp"fcp"Rgtmgdwpcp0"Dqiqt0"

¥qdgn." D0." Oe0Mgg." C0." Jqgm." I0O0." cpf"F{tpguu." E0" "3;980"T gncvkqpujkr" qh"gpxktqpogpv" vq" eqorqukvkqp." uvtwevwtg"cpf"fkxgtukv{"qh"hqtguv"eqoowpkvkgu"qh"vjg"egpvtcn" yguvgtp" ectecfgu" qh"Q tgiqp0"Ecol.Mono0"68