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References Allen, D & C Mellon. 2004. Factors relating to the wintering population of diving duck on the Lough Neagh System. Research report by Allen and Mellon Environmental Ltd, to Environment and Heritage Service, Belfast, 34 pp. Allen, D, C Mellon, I Enlander & G Watson. 2004. Lough Neagh diving ducks: recent changes in wintering populations. Irish Birds 7: 327-336. Anthes, N. 2004. Long-distance migration timing of Tringa sandpipers adjusted to recent climate change. Bird Study 51: 203-211. Atkinson, PW, GE Austin, MM Rehfisch, H Baker, P Cranswick, M Kershaw, J Robinson, RHW Langston, DA Stroud & C van Turnhout. in prep. Identifying declines in waterbirds: the effects of missing data, population variability and count period on the interpretation of long-term survey data. Austin, GE, I Peachel & MM Rehfisch. 2000. Regional trends in coastal wintering waders in Britain. Bird Study 47: 352-371. Austin, GE, SSF Jackson & HJ Mellan. 2004. WeBS Alerts 2000/2001: Changes in numbers of wintering waterbirds in the United Kingdom, its Constituent Countries, Special Protection Areas (SPAs) and Sites of Special Scientific Interest (SSSIs). BTO Report to the WeBS Partners, BTO Research Report 349. Banks, A, D Bolt, I Bullock, B Haycock, A Musgrove, S Newson, N Fairney, W Sanderson, R Schofield, L Smith, R Taylor & S Whitehead. 2004. Ground and aerial monitoring protocols for in shore Special Protection Areas. CCW Marine Monitoring Report No 11, 89 pp. Baillie, SR, JH Marchant, HQP Crick, DG Noble, DE Balmer, LP Beaven, RH Coombes, IS Downie, SN Freeman, AC Joys, DI Leech, MJ Raven, RA Robinson & RM Thewlis. 2005. Breeding Birds in the Wider Countryside: their conservation status 2004. BTO Research Report No. 385. BTO, Thetford. (http://www.bto.org/birdtrends) Bainbridge, IP & CDT Minton. 1978. The migration and mortality of the Curlew in Britain and Ireland. Bird Study 25: 39-50. Banks, A, D Bolt, I Bullock, B Haycock, A Musgrove, S Newson, N Fairney, W Sanderson, R Schofield, L Smith, R Taylor & S Whitehead 2004. Ground and aerial monitoring protocols for in shore Special Protection Areas. CCW Marine Monitoring Report No: 11, 89 pp. Beecroft, R. 1990. Trimley Marshes Nature Reserve. In: Piotrowski, SH. (ed.) Suffolk Birds 1990. The Suffolk Naturalists’ Society, Ipswich. Beekman, JH. 1997. Censuses of NW European Bewick’s Swan population January 1990 and 1995. Swan Specialist Group Newsletter 6: 7-9. Bibby, CJ, ND Burgess, DA Hill & S Mustoe. 2000. Bird Census Techniques. Second Edition. Academic Press, London. Birdlife International. 2004. Birds in Europe. Population Estimates, Trends and Conservation Status. Birdlife Conservation Series 12. Birdlife International. BOURC. 1999. British Ornithologists’ Union Records Committee: 25th Report October 1998. Ibis 141: 175-180. Boyd, H & T Piersma. 2001. Changing balance between survival and recruitment explains population trends in Red Knots Calidris canutus islandica wintering in Britain, 1969-1995. Ardea 89: 301-317. Brown, L & M Smart. 2002. Pintail in the Severn Vale. Worcestershire Record New Series, Number 12 (Newsletter of the Worcestershire Biological Records Centre), April 2002, ISSN 1475-9616. Buck, AL. 1993. An Inventory of UK Estuaries Volume 3, North-west Britain. JNCC, Peterborough. Buck, AL. 1997. An Inventory of UK Estuaries. 7 volumes. JNCC, Peterborough. Buckland, ST, D Anderson, K Burnham, J Laake, D Borchers & L Thomas. 2001. Introduction to distance sampling: estimating abundance of biological populations. Oxford University Press, Oxford. Burton, NHK, AJ Musgrove, MM Rehfisch, A Sutcliffe, & RJ Waters. 2003. Numbers of wintering gulls in the United Kingdom, Channel Islands and Isle of Man: a review of the 1993 and previous Winter Gull Roost Surveys. British Birds 96: 376-401. Camphuysen, CJ, CM Berrevoets, HJWM Cremers, A Dekinger, R Dekker, BJ Ens, TM Have, RKH van der Kats, T Kuiken, MF Leopold, J van der Meer & T Piersma. 2002. Mass mortality of common eiders (Somateria mollissima) in the Dutch Wadden Sea, winter 1999/2000: starvation in a commercially exploited wetland of international importance. Biological Conservation 106: 303-317. Carp, E. 1972. Proceedings of the international conference on the conservation of wetlands and waterfowl. Ramsar, Iran, 30 January - 3 February 1971. IWRB, Slimbridge. Cayford, JT & RJ Waters. 1996. Population estimates for waders Charadrii wintering in Great Britain, 1987/88-1991/92. Biological Conservation 77: 7-17. Chandler, RJ. 1981. Influxes into Britain and Ireland of Red-necked Grebes and other waterbirds during winter 1978/78. British Birds 74: 55-81. References 288

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Page 1: WWC 2000-01: References · PDF fileReferences Allen, D & C Mellon. 2004. ... Neagh System. Research report by Allen and Mellon Environmental Ltd, ... DA Stroud & C van Turnhout. in

ReferencesAllen, D & C Mellon. 2004. Factors relating to the

wintering population of diving duck on the Lough

Neagh System. Research report by Allen and

Mellon Environmental Ltd, to Environment and

Heritage Service, Belfast, 34 pp.

Allen, D, C Mellon, I Enlander & G Watson. 2004.

Lough Neagh diving ducks: recent changes in

wintering populations. Irish Birds 7: 327-336.

Anthes, N. 2004. Long-distance migration timing of

Tringa sandpipers adjusted to recent climate

change. Bird Study 51: 203-211.

Atkinson, PW, GE Austin, MM Rehfisch, H Baker, P

Cranswick, M Kershaw, J Robinson, RHW

Langston, DA Stroud & C van Turnhout. in prep.

Identifying declines in waterbirds: the effects of

missing data, population variability and count

period on the interpretation of long-term survey

data.

Austin, GE, I Peachel & MM Rehfisch. 2000. Regional

trends in coastal wintering waders in Britain. Bird

Study 47: 352-371.

Austin, GE, SSF Jackson & HJ Mellan. 2004. WeBS

Alerts 2000/2001: Changes in numbers of wintering

waterbirds in the United Kingdom, its Constituent

Countries, Special Protection Areas (SPAs) and

Sites of Special Scientific Interest (SSSIs). BTO

Report to the WeBS Partners, BTO Research

Report 349.

Banks, A, D Bolt, I Bullock, B Haycock, A Musgrove,

S Newson, N Fairney, W Sanderson, R Schofield, L

Smith, R Taylor & S Whitehead. 2004. Ground and

aerial monitoring protocols for in shore Special

Protection Areas. CCW Marine Monitoring Report

No 11, 89 pp.

Baillie, SR, JH Marchant, HQP Crick, DG Noble, DE

Balmer, LP Beaven, RH Coombes, IS Downie, SN

Freeman, AC Joys, DI Leech, MJ Raven, RA

Robinson & RM Thewlis. 2005. Breeding Birds in

the Wider Countryside: their conservation status

2004. BTO Research Report No. 385. BTO,

Thetford. (http://www.bto.org/birdtrends)

Bainbridge, IP & CDT Minton. 1978. The migration

and mortality of the Curlew in Britain and Ireland.

Bird Study 25: 39-50.

Banks, A, D Bolt, I Bullock, B Haycock, A Musgrove,

S Newson, N Fairney, W Sanderson, R Schofield, L

Smith, R Taylor & S Whitehead 2004. Ground and

aerial monitoring protocols for in shore Special

Protection Areas. CCW Marine Monitoring Report

No: 11, 89 pp.

Beecroft, R. 1990. Trimley Marshes Nature Reserve.

In: Piotrowski, SH. (ed.) Suffolk Birds 1990. The

Suffolk Naturalists’ Society, Ipswich.

Beekman, JH. 1997. Censuses of NW European

Bewick’s Swan population January 1990 and 1995.

Swan Specialist Group Newsletter 6: 7-9.

Bibby, CJ, ND Burgess, DA Hill & S Mustoe. 2000.

Bird Census Techniques. Second Edition.

Academic Press, London.

Birdlife International. 2004. Birds in Europe.

Population Estimates, Trends and Conservation

Status. Birdlife Conservation Series 12. Birdlife

International.

BOURC. 1999. British Ornithologists’ Union Records

Committee: 25th Report October 1998. Ibis 141:

175-180.

Boyd, H & T Piersma. 2001. Changing balance

between survival and recruitment explains

population trends in Red Knots Calidris canutus

islandica wintering in Britain, 1969-1995. Ardea 89:

301-317.

Brown, L & M Smart. 2002. Pintail in the Severn Vale.

Worcestershire Record New Series, Number 12

(Newsletter of the Worcestershire Biological

Records Centre), April 2002, ISSN 1475-9616.

Buck, AL. 1993. An Inventory of UK Estuaries Volume

3, North-west Britain. JNCC, Peterborough.

Buck, AL. 1997. An Inventory of UK Estuaries. 7

volumes. JNCC, Peterborough.

Buckland, ST, D Anderson, K Burnham, J Laake, D

Borchers & L Thomas. 2001. Introduction to

distance sampling: estimating abundance of

biological populations. Oxford University Press,

Oxford.

Burton, NHK, AJ Musgrove, MM Rehfisch, A Sutcliffe,

& RJ Waters. 2003. Numbers of wintering gulls in

the United Kingdom, Channel Islands and Isle of

Man: a review of the 1993 and previous Winter Gull

Roost Surveys. British Birds 96: 376-401.

Camphuysen, CJ, CM Berrevoets, HJWM Cremers, A

Dekinger, R Dekker, BJ Ens, TM Have, RKH van der

Kats, T Kuiken, MF Leopold, J van der Meer & T

Piersma. 2002. Mass mortality of common eiders

(Somateria mollissima) in the Dutch Wadden Sea,

winter 1999/2000: starvation in a commercially

exploited wetland of international importance.

Biological Conservation 106: 303-317.

Carp, E. 1972. Proceedings of the international

conference on the conservation of wetlands and

waterfowl. Ramsar, Iran, 30 January - 3 February

1971. IWRB, Slimbridge.

Cayford, JT & RJ Waters. 1996. Population estimates

for waders Charadrii wintering in Great Britain,

1987/88-1991/92. Biological Conservation 77: 7-17.

Chandler, RJ. 1981. Influxes into Britain and Ireland

of Red-necked Grebes and other waterbirds during

winter 1978/78. British Birds 74: 55-81.

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GlossaryThe terms listed below are generally restricted to those that have been adopted specifically for use within WeBS

or more widely for monitoring.

Autumn For waders, autumn comprises July to

October inclusive. Due to differences in seasonality

between species (see Monthly fluctuations), a strict

definition of autumn is not used for wildfowl.

Complex site A WeBS site that consists of two or

more sectors.

Core Counts The basic WeBS counts that monitor all

wetlands throughout the UK once per month on

priority dates. Used to determine population

estimates and trends and identify important sites.

Distance survey/analysis A method of survey where

observers travel along a set path (transect) and not

only count but record the distance to any birds seen.

Birds further from the transect will tend to be missed

by the observers, but recording the distance to each

bird enables the number missed to be calculated.

This technique is now used regularly for aerial survey

of marine waters.

Local Organiser Person responsible for co-

ordinating counters and counts at a local level,

normally a county or large estuary, and the usual

point of contact with WeBS partners.

Incomplete counts When presenting counts of an

individual species, a large proportion of the number

of birds was suspected to have been missed, eg due

to part coverage of the site or poor counting

conditions, or when presenting the total number of

birds of all species on the site, a significant

proportion of the total number was missed.

I-WeBS An independent but complementary scheme

operating in the Republic of Ireland to monitor non-

breeding waterbirds, organised by the IWC

BirdWatch Ireland, the National Parks and Wildlife

Service (Ireland) and The Wildfowl & Wetlands

Trust.

Low Tide Counts (LTC) WeBS counts made at low

tide to assess the relative importance of different

parts of individual estuaries as feeding areas for

intertidal waterbirds.

National threshold Used as a generic term to imply

the 1% British threshold for sites in Great Britain, and

the 1% all-Ireland threshold for sites in Northern

Ireland. Similarly, national importance is used as a

generic term to refer to sites in Great Britain and in

Northern Ireland that meet the respective thresholds.

Short-stopping The phenomenon of migratory birds

travelling only part of the way to their former final

wintering destination. For example, birds that

previously wintered in Britain and Ireland will remain

on the Continent (usually because of warmer

weather).

Spring For waders, spring comprises April to June

inclusive. Due to differences in seasonality between

species (see Monthly fluctuations), a strict definition

of spring is not used for wildfowl.

Waterbirds WeBS follows the definition adopted by

Wetlands International. This includes a large number

of families, those occurring regularly in the UK being

divers, grebes, cormorants, herons, storks, ibises and

spoonbills, wildfowl, cranes, rails, waders, gulls and

terns.

WeBS count sector The unit of division of large sites

into areas which can be counted by one person in a

reasonable time period. They are often demarcated

by geographic features to facilitate recognition of the

boundary by counters. The finest level at which data

are recorded.

WeBS count site A biologically meaningful area that

represents a discrete area used by waterbirds such

that birds regularly move within but only occasionally

between sites. The highest level at which count data

are stored.

WeBS count sub-site A grouping of sectors within a

site to facilitate co-ordination. In most cases, sub-

sites also relate to biologically meaningful units for

describing waterbird distribution.

WeBS count unit The area/boundary within which a

count is made. The generic term for sites, sub-sites

and sectors.

Winter For waders, winter comprises November to

March inclusive. Due to differences in seasonality

between species (see Monthly fluctuations), a strict

definition of winter is not used for wildfowl.

1% criterion The Ramsar Convention has

established site selection criteria. Criterion 6 states

that ‘a wetland should be considered internationally

important if it regularly supports 1% of the individuals

in a population of one species or subspecies of

waterbird’. See also national importance.

1% threshold This logically derives from the 1%

criterion and relates to the number of birds that are

used as the nominal 1% of the population for the

purposes of site selection. Thus, an international

population of 300,000 Shelduck has a derived 1%

threshold of 3,000.

Glossary 295

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Appendices

APPENDIX 1. INTERNATIONAL SITE DESIGNATIONS

The Ramsar Convention on Wetlands of International

Importance especially as Waterfowl Habitat requires

each Contracting Party to designate suitable

wetlands, selected on account of their international

significance in terms of ecology, botany, zoology,

limnology or hydrology, for inclusion in a List of

Wetlands of International Importance (known as

Ramsar sites) (Carp 1972, Ramsar Convention

Secretariat 2004). The EU Directive on the

Conservation of Wild Birds (EC/79/409) lays emphasis

on the need to conserve bird habitats as a means of

maintaining populations and that this, in part, should

be achieved by the establishment of a network of

protected areas termed Special Protection Areas

(SPAs) (Stroud et al 1990, 2001). Ramsar sites and

SPAs may be identified using a number of criteria,

including a number of quantitative selection criteria

(see Appendix 2) which draw heavily upon waterbird

counts, especially WeBS and the other data

presented in this report.

The tables below list the number and area of

classified SPAs and Ramsar sites as of 26 May 2005.

More detailed information about protected sites can

be accessed via the JNCC website at

http://www.jncc.gov.uk/page-4. Full listings of sites

currently designated as SPAs and Ramsar sites can

be found at the following links:

http://www.jncc.gov.uk/page-1400http://www.jncc.gov.uk/page-1389

Table A1. Number and area of classified SPAs and designated Ramsar sites in the UK as of 26/5/2005.

Classified SPAs Designated Ramsar sitesCountry No. of sites Site Area (ha) No. of sites Area (ha) England 77 609,249 66 294,427England/Scotland 1 43,637 1 43,637England/Wales 2 37,748 3 39,336Northern Ireland 10 70,660 17 86,831Scotland 139 597,883 50 283,569Wales 17 123,007 7 11,366Total 246 1,482,187 144 758,684

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APPENDIX 2. INTERNATIONAL AND NATIONAL IMPORTANCE

Any site recognised as being of international

ornithological importance is considered for

classification as a Special Protection Area (SPA)

under the EC Directive on the Conservation of Wild

Birds (EC/79/409), whilst a site recognised as an

internationally important wetland qualifies for

designation as a Ramsar site under the Convention

on Wetlands of International Importance especially

as Waterfowl Habitat. Criteria for assessing the

international importance of wetlands have been

agreed by the Contracting Parties to the Ramsar

Convention on Wetlands of International Importance

(Ramsar Convention Bureau 2004). Under Criterion 6,

a wetland is considered internationally important if it

regularly holds at least 1% of the individuals in a

population of one species or subspecies of waterbird,

while Criterion 5 states that any site regularly

supporting 20,000 or more waterbirds also qualifies.

Britain and Ireland’s wildfowl belong, in most cases,

to the northwest European population (Scott & Rose

1996), and the waders to the east Atlantic flyway

population (Stroud et al 2004).

A wetland in Britain is considered nationally

important if it regularly holds 1% or more of the

estimated British population of one species or

subspecies of waterbird, and in Northern Ireland is

important in an all-Ireland context if it holds 1% or

more of the estimated all-Ireland population.

The 1% thresholds for British, all-Ireland and

international waterbird populations, where known,

are listed in Table A2. Thus, any site regularly

supporting at least this number of birds potentially

qualifies for designation under national legislation, or

the EC Bird’s Directive or Ramsar Convention. The

international population for each species and sub-

species is also specified in the table. However, it

should be noted that, where 1% of the national

population is less than 50 birds, 50 is normally used

as a minimum qualifying threshold for the

designation of sites of national or international

importance (Stroud et al 2001).

1% thresholds have not been derived for

introduced species since it would not be appropriate

to establish protected sites (eg SSSIs) for these

species.

Sources of qualifying levels represent the most

up-to-date figures following recent reviews: for

wildfowl in Britain see Kershaw & Cranswick (2003);

for waders in Britain see Rehfisch et al (2003a); for

gulls in Britain see Burton et al (2003); for all-Ireland

importance for divers see Danielsen et al (1993) and

for other waterbirds see Whilde (in prep) cited in

Way et al (1993). International criteria follow

Wetlands International (2002).

It was agreed at the 1996 meeting of the Ramsar

Convention in Brisbane that population estimates will

be reviewed by Wetlands International every three

years and 1% thresholds revised every nine years

(Rose & Stroud 1994; Ramsar Resolution VI.4).

Note the estimates for Britain and internationally

have been revised since the last Wildfowl & Wader

Counts, and some population names have been

changed also (following Wetlands International

2002). It should be noted that for some populations,

where the British total is the international total, the

precise figure given for the estimates may differ

because of different rounding conventions applied in

the relevant publications.

Table A2. 1% thresholds for national and international importance.

Species/sub-species Population Great Britain All-Ireland InternationalMute Swan Britain 375 - 380

Ireland - 100 100Bewick's Swan bewickii, NW Europe (non-br) 81 *25 290Whooper Swan Iceland (br) 57 100 210Bean Goose fabalis (Taiga) *4 + 1,000

rossicus (Tundra) 1 6,000Pink-footed Goose Greenland, Iceland (br) 2,400 + 2,400European White-fronted Goose albifrons, Baltic-North Sea 58 + 10,000Greenland White-fronted Goose flavirostris 209 140 300Greylag Goose anser, Iceland (br) 819 *40 1,000

anser, NW Scotland 96 - 90Barnacle Goose E Greenland (br) 450 75 540

Svalbard (br) 220 + 230Dark-bellied Brent Goose bernicla, W Siberia (br) 981 + 2,200Light-bellied Brent Goose hrota, Ireland (non-br) + 200 200

hrota, Svalbard, N Greenland (br) *30 + 50Shelduck NW Europe (br) 782 70 3,000Wigeon NW Europe (non-br) 4,060 1,250 15,000Gadwall strepera, NW Europe (br) 171 + 600Teal NW Europe (non-br) 1,920 650 4,000Mallard platyrhynchos, NW Europe (non-br) 3,520 500 **20,000Pintail NW Europe (non-br) 279 60 600Garganey W Africa (non-br) + + **20,000

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Species/sub-species Population Great Britain All-Ireland InternationalShoveler NW & C Europe (non-br) 148 65 400Red-crested Pochard C Europe & W Mediterranean + + 500Pochard NE & NW Europe (non-br) 595 400 3,500Tufted Duck NW Europe (non-br) 901 400 12,000Scaup marila, W Europe (non-br) 76 *30 3,100Eider 1 mollissima, Britain, Ireland 730 *20 750Eider 1 mollissima, Baltic, Wadden Sea - - 10,300Long-tailed Duck W Siberia, N Europe (br) 160 + **20,000Common Scoter nigra 500 *40 16,000Velvet Scoter fusca, Baltic, W Europe (non-br) *30 + 10,000Goldeneye clangula, NW, Central Europe (non-br) 249 110 4,000Smew NW & C Europe (non-br) *4 + 400Red-breasted Merganser NW & C Europe (non-br) 98 *20 1,700Goosander 2 merganser, Scotland, N England, Wales 161 + 160Goosander 2 merganser, NW & C Europe (non-br) - - 2,500Red-throated Diver NW Europe (non-br) 49 *10 10,000Black-throated Diver arctica *7 *1 10,000Great Northern Diver NW Europe (non-br) *30 ? 50Little Grebe ruficollis 78 ? 3,400Great Crested Grebe cristatus 159 *30 4,800Red-necked Grebe grisegena, NW Europe (non-br) *2 ? 1,000Slavonian Grebe auritus, NW Europe (large billed) *7 ? 35Black-necked Grebe nigricollis, Europe, N Africa *1 ? 2,800Cormorant carbo, NW Europe 230 ? 1,200Shag aristotelis ? ? 2,400Little Egret garzetta, W Europe, NW Africa ? ? 1,300Grey Heron cinerea, W Europe, NW Africa (br) ? ? 2,700Moorhen chloropus, Europe, N Africa (br) 7500 ? **20,000Coot atra, NW Europe (non-br) 1,730 250 17,500Oystercatcher ostralegus, Europe, NW Africa 3,200 500 10,200Avocet W Europe (br) *35 + 730Little Ringed Plover curonicus, Europe, W Africa ? ? 2,400Ringed Plover hiaticula, Europe & N Africa (non-br) 330 125 730Golden Plover apricaria - - 650

altifrons, Iceland & Faeroes, E Atlantic 2,500 2,000 9,300Grey Plover E Atlantic (non-br) 530 *40 2,500Lapwing Europe (br) **20,000 2,500 **20,000Knot islandica 2,800 375 4,500

canutus 3,400Sanderling E Atlantic, W & S Africa (non-br) 210 *35 1,200 passage 300Little Stint Europe & West Africa (non-br) ? ? 2,000Curlew Sandpiper W Africa (non-br) ? ? 7,400Purple Sandpiper maritima, E Atlantic 180 *10 750Dunlin arctica 330

schinzii, Iceland (br) 9,500schinzii, Baltic (br) 40schinzii, Britain & Ireland (br) 250alpina, W Europe (non-br) 5,600 1,250 13,300

passage 2,000Ruff W Africa (non-br) *7 + ?Jack Snipe NE Europe (br) ? 250 ?Snipe gallinago, Europe (br) ? ? **20,000Woodcock Europe (br) ? ? **20,000Black-tailed Godwit islandica 150 90 350Bar-tailed Godwit lapponica 620 175 1,200Whimbrel islandicus + + 6,100Curlew arquata 1,500 875 4,200Spotted Redshank Europe (br) 1 + 1,000Redshank 1,200 245

totanus, E Atlantic (br) 2,500robusta 650

Redshank brittanica 1,300

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Species/sub-species Population Great Britain All-Ireland International passage 1,200Greenshank Europe (br) *6 *9 3,100Green Sandpiper Europe (br) ? ? 14,500Common Sandpiper NW & C Europe (br) ? ? 17,000Turnstone interpres, NE Canada, Greenland (br) 500 225 1,000Mediterranean Gull Black Sea, Med’nean, Europe, Africa ? ? 8,400Little Gull N, C & E Europe (br) ? ? 840Black-headed Gull N & C Europe (br) 19,000 ? **20,000Common Gull canus 9,000 ? 17,000Lesser Black-backed Gull graellsii 500 ? 5,300Herring Gull argentatus 4,500 ? 13,000

argenteus 11,000Yellow-legged Gull cachinnans 10,000

michahellis 7,000Great Black-backed Gull NE Atlantic 400 ? 4,700Kittiwake tridactyla, E Atlantic (br) ? ? **20,000Little Tern albifrons, W Europe (br) ? ? 340Black Tern niger ? ? 4,000Sandwich Tern sandvicensis, W Europe (br) ? ? 1,700Common Tern hirundo, N, E Europe (br) ? ? 6,400

? Population size not accurately known+ Population too small for meaningful figure to be obtained * Where 1% of the British or all-Ireland wintering population is less than 50 birds, 50 is normally used as a minimum qualifying level for national

or all-Ireland importance** A site regularly holding more than 20,000 waterbirds qualifies as internationally important by virtue of absolute numbers 1 The UK’s SPA and Ramsar Scientific Working Group recommends that a figure of 15,500 is used as the international 1% threshold for site-

selection (see species account for Eider for further detail) 2 The UK’s SPA and Ramsar Scientific Working Group recommends that a figure of 2,700 is used as the international 1% threshold for site-

selection (see species account for Goosander for further detail)

Appendices 299

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APPENDIX 3. ANALYSES

This appendix provides additional detail about the

analyses used in this report to that presented in

Analyses and lists the index values used to produce

the graphs of annual and monthly indices in the

species accounts.

Data availability

The count scheme first begun in 1947 has developed

considerably over time (see Cranswick et al 1997). In

particular, coverage of species and area has

expanded during this time. The first year for which

data for certain species or areas are available for use

in analyses are given below:

Table A3. First year of availability of WeBS Core Count data for different species and areas.

Wildfowl in GB 1960 on computer (collected since 1947)Waders in UK 1969/70Great Crested Grebe 1982/83Coot 1982/83Little Grebe 1985/86Cormorant 1986/87Wildfowl in Northern Ireland 1986/87All other species (rare grebes, divers, rarities, gulls, terns) 1993/94

National totals for goose populations

Figures presented in Tables A4-A9 for total counts of

the various goose populations are derived initially

from WeBS Core Counts, but are replaced by results

of dedicated censuses (see Survey Methods, Analyses

and Coverage for appropriate references, methods

and dates) where these provide better counts.

Several goose populations are identified according to

location (and totals derived by summing counts from

particular WeBS regions) where they cannot be

separated in the field by appearance.

Table A4. Data used for national totals for Goose populations.

Bean Goose WeBS Core Counts in all months.

Pink-footed Goose October and November counts replaced by summed counts from the co-ordinatednational censuses.

European White-fronted Goose WeBS Core Counts in all months.

Greenland White-fronted Goose November and March counts replaced by summed counts from the co-ordinatedlate autumn and late spring international censuses, respectively.

Greylag Goose: Iceland

WeBS Core Counts from all WeBS regions in Scotland except those on the westcoast (see NW Scotland population) plus Northumberland and North Cumbria.October and November counts replaced by summed counts from the co-ordinatednational censuses.

NW Scotland WeBS Core Counts from WeBS regions Islay/Jura/Colonsay,Mull/Lismore/Coll/Tiree, Skye, Highland Southwest and North and South OuterHebrides. August and February counts replaced by summed counts from co-ordinated censuses of Outer Hebrides. August 1997 count replaced by total fromfull national survey.

naturalised WeBS Core Counts for all sites in Wales and England, except for Northumberlandand North Cumbria.

Note that Icelandic and NW Scotland populations overlap in WeBS regions Orkney and North Highland. NW Scotland birds counted byWeBS in these regions will be included Icelandic population totals. Note also that up to 2,340 naturalised birds occur in Scotland(Delany 1993) and others in Northumberland and North Cumbria

Canada Goose WeBS Core Counts in all months

Barnacle Goose: Greenland

WeBS Core Counts from all WeBS regions on Scottish west coast, plus Shetlandand Orkney. November and March counts replaced by summed counts from theco-ordinated late autumn and late spring censuses in Argyll plus the monthlymaximum count from Hoy, Orkney.

Appendices300

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Barnacle Goose: Svalbard

WeBS Core Counts from WeBS regions Dumfries & Galloway, North Cumbria,Northumberland, Borders, Lothians, Central, Fife, Perth & Kinross, Angus,Grampian, Moray and SE Highland. Dumfries & Galloway and North CumbriaWeBS Core Counts replaced by Solway-wide counts and censuses betweenOctober and March.

naturalised WeBS Core Counts for all WeBS regions in Wales and England, except forNorthumberland and North Cumbria.

Dark-bellied Brent Goose WeBS Core Counts, plus additional counts of inland areas in January and February

Light-bellied Brent Goose: Canada

WeBS Core Counts for sites in Northern Ireland, Wales and WeBS regionsShetland, Orkney, Highland North, Western Isles, Skye, Highland Southwest,Islay/Jura/Colonsay, Argyll West Mainland, Mull/Lismore/Coll/Tiree, Dumbarton/SEArgyll, Renfrew, Lanarkshire/Strathkelvin, Ayrshire & Arran, Dumfries & GallowayWest, Cornwall, Devon and the Channel Islands. October count in NorthernIreland replaced by counts in Northern Ireland made during all-Ireland census.

Svalbard WeBS Core Counts from regions not used to compile Canada population totals(see above).

Annual Indices

Annual indices using WeBS data are calculated using

a Generalised Additive Model (GAM) (see Methods

and Atkinson et al in prep). Index values are derived

from sites where at least 50% of the maximum

possible number of counts, bearing in mind that

different months are used for different species, were

complete. Index values provided extend back to

1966/67 for wildfowl and 1971/72 for waders,

representing the first years in which coverage was

deemed sufficient for data to be included in the

calculation of the index. A number of species were

only first included in WeBS in the 1980s, whilst

counts of wildfowl in Northern Ireland only began in

earnest in 1985/86.

Underhill (1989) recommended that, where

possible, the index is based on counts from more

than one month. The months chosen for each

species are given below. The most appropriate

grouping of months on which to base the annual

index for waders is December, January and February,

the period when the wintering population in Britain

and Northern Ireland is most stable (Prys-Jones et al

1994). However, the peak abundance of different

wildfowl occur in different months according to

species, and thus different months and different

numbers of months were selected for each (Kirby et

al 1995).

The selection of months for calculating indices

for wildfowl and their allies was made by first

calculating monthly index values for all months

September to March, and selecting that with the

highest index value and any adjacent months with

overlapping consistency intervals. Data from all years

from 1966/67 onwards were used for calculating the

index for each of these species, as recommended in

Kirby et al (1995), or from the years in which data

were first available for species added to the scheme

subsequently (see above). Caution is urged in

particular regarding the first few years’ index values

for these species only recently included in the

scheme; missing counts may have been incorrectly

recorded as nil counts, giving rise to anomalous

index values.

Due to more stable populations of waders

during the winter, the months December to February

are chosen for calculation of index values for all

waders for which there are suitable data. Due to the

small number of sites in Northern Ireland, data are

combined for analysis at the UK level.

Table A5. Months used in calculating indices for wildfowl species in Great Britain and Northern Ireland (indicated using thefirst letter of the months September to March).

Species GB NIMute Swan SONDJFM SONDJBewick's Swan JF NDJFWhooper Swan ND ONDJFMPink-footed Goose O or N -European White-fronted JF -Greenland White-fronted N or M N or M Greylag Goose: Icelandic O or N -

naturalised S -Canada Goose S -Barnacle Goose: Svalbard any month -Dark-bellied Brent Goose DJF -Light-bellied Brent Goose - SONDJFMShelduck JF DJFMWigeon J SONDJFMGadwall SONDJFM SONDJ

Species GB NITeal D DJMallard D SOPintail ONDJ ONDJFMShoveler SO SONDJFMPochard NDJ NDJFTufted Duck NDJF ONDJFMGoldeneye F DJFMRed-breasted Merganser ONDJFM SONDJFMGoosander DJF -Ruddy Duck SONDJFM -Little Grebe SO SONGreat Crested Grebe SON SONDJFMCormorant SONDJFM SONDCoot SONDJ SONDJ

Appendices 301

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2002/032001/022000/011999/001998/991997/981996/971995/961994/951993/941992/931991/921990/911989/901988/891987/881986/871985/861984/851983/841982/831981/821980/811979/801978/791977/781976/771975/761974/751973/741972/731971/721970/711969/701968/691967/681966/67Y

ear

100 96 92 97 95 89 85 83 74 74 73 72 76 70 62 60 51 50 45 45 50 47 44 46 45 42 43 44 46 48 50 47 48 50 49 51 57M

S

100 22 81 38 46219202264197225247292265225180131332219289166146148159142142134109 78 56 65 43 73104 50 51 64 63B

S

100230 98109 76112127100 89 80101103146125130 88 98 83 51 55 80 45 72 78 52 57 47 63 74 52 67 68 57 79 46 48 71W

S

100126115101109112111 95123106 94110 91 87 84 82 64 61 41 48 42 43 45 38 37 33 34 35 42 39 32 31 34 35 31 32 36P

G

100126 97170164245230272201278 88321233290309361311286331224267297264193371211290237117313124184428494538388135

EW

100 95108106113105107110 98 87 78 83 78 74 65 61 58 57 48 42 37

NW

100142130123135128128133139160158142185134175169164172103131130154145130123108 90101111123117103104100 98 85 97 JI

100106104104 89 82 77 73 61 52 54 48 45 47 48 39 38 38 36 27 23 18 14 11 8 10 7 4 5 3 6 3 2 1 1 1 4 JE

100 93101 86 89 76 93 72 57 59 69 77 70 70 79 74 63 63 57 44 47 41 38 34 32 28 25 18 19 20 24 14 16 16 12 12 8C

G

100 83 84 91 92 84 81 72 63 50 46 47 43 41 43 40 37 37 37 30 30 29 32 27 31 24 25 21 18 18 15 13 11 14 15 13 13Y

S

100106124128104131133121134164139184147116145128128126107108113 88 85 93 81 64 59 40 44 56 42 42 43 33 32 39 32D

B

100110105100109120153151118138241275255199186197161150114119119161159130153 92105 90 89 95 94 84 71121100 95 88S

U

100 94100 90103 89122107 98 89 87 93 72 73 75 75 92 85 83 43 44 57 57 55 69 47 62 45 48 59 55 49 60 69 63 69 59

WN

Table

A6.G

reatBritainannualindex

valuesfor

wildfow

l (seeT

ableA

7for

speciescodes).

100100 89100 83 87 77 81 72 59 52 51 56 53 54 50 45 39 35 36 34 31 25 23 20 20 12 12 14 10 12 12 11 5 6 4 4G

A

100151104137123134122151107125101116118147108100 82 89 85106123135101 88 88 78 70 65 57 67 52 50 46 53 44 26 36 T

.

100108101111117103111130116114131133138149149166149168138160154139146146148117143118139131145151175124155142173M

A

100 85 79 70100 87 96108 94 83106130105132125140 92129163131145121142117 93170 98 99115135 74 56 59 40 48 32 28P

T

100113105118102122125125100 92 94122120106 92 91101100101 93 93 85 83 93 86 87 79 64 79105 89 83 53 60 75 52 53S

V

100 86106118120109107130117107105108111117119104115101116122116 94111111106130112147154136137121111106119135131P

O

100102108102100 98103 93 89 89 88 89 92 86105 86 92 91 87 84 88 86 99 96 86 88 80105 97102105 97 84 71 73 69 63T

U

100111145 89 91173240249146130109104250 57150117164 43116123195380449422401482300

1089153125271579218013021442178817621650

SP

100105107127130127133127115113123114127103 98103102 78 87 74 76 94 76 84 72 59 61 68 74 80 72 71 72 49 47 61 75G

N

100123105132120131113130119 94102 99 95 87 94 85 70 70 63 56 58 62 56 60 52 54 60 46 47 45 39 40 36 38 27 30 24R

M

100126 89140146153253196100123120135140116111106112 92 85 97 69 79 82 67 98 64 62 72 46 83 51 75 68 89 64 61 41G

D

100101 92107 88 94 95 80 68 54 62 69 68 66 63 57 52 50 49 45 33 30 34 19 13 11 11 6 3 3 2 1 1 1 1 0 0R

Y

100104101102 96 84 95 97 88 86 80 77 77 90 90 78 81 81 85

CO

100101 95101 82 79 80 97 72 64 55 58 59 58 51 20

LG

100108111100100 97 94 99 92 88 95 88 89 90 76 70 68 68 81

GG

100109100105 97 91100100 90 92 85 83 92 77 75

CA

Appendices302

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CA 67 86 63 54 81 74 73 78 81 111

129

127 94 104

100

GG 58 76 74 76 68 80 97 62 139

125

103

146 95 113

116

138

100

LG 20 112

114 87 96 109

114

137

147

107 98 94 107

114

113

100

CO

114

147

139

187

172

172

187 99 195

224

166

163

162

208

188

134

100

RY 92 90 94 46 58 108 91 77 104

147

200 84 139

149

216

100

RM 35 107 91 116 99 96 121

102

116

149

102

132

112 97 89 95 100

GN

285

335

271

259

331

357

315

217

233

237

187

159

226

194

249

189

100

SP 87 82 92 42 70 121

179

117

171

160

141

183

197

207

171

192

100

TU 99 192

202

213

203

227

211

217

216

257

211

185

170

198

263

162

100

PO

234

250

357

324

330

353

256

224

215

304

240

215

213

272

248

188

100

SV 63 103

114

100 84 110 80 129

106

108 98 76 89 108 90 107

100

Red

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PT 6 50 75 43 67 79 62 66 42 45 65 65 87 80 55 56 100

RM

GD

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CA

MA

101 89 108

127

120

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101 98 123

151

107

104

121

101

108

119

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T.

52 126

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132

153

115 80 86 87 91 102

110

121

110

108

115

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GA 38 101

105

152

119

137

161

139

171

128

134 89 119 98 126

130

100

WN

118

153

193

151

174

160

123

116

123

101

125

133

109

160

114

118

100

SU 25 54 89 68 83 65 60 68 71 85 82 92 90 91 84 97 100

Can

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Ligh

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QN 7 86 82 84 102

104 75 71 86 83 87 100

101 96 112 92 100

QN

WN

GA

T.

MA

PT SV PO TU

SP GN

CG 0 58 83 88 67 53 67 31 80 39 84 39 46 57 64 63 100

WS 56 77 86 59 64 60 65 34 65 63 73 55 55 56 40 61 100

BS 28

1842

3240

9573

0580

1439

7319

2794

2710

6818

9529

77 877

568

159

177

232

100

MS 87 106

115

127

123

127

134

116

143

151

134

151

166

158

148

145

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Mut

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90/9

119

91/9

219

92/9

319

93/9

419

94/9

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95/9

619

96/9

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B

Appendices 303

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Table A8. UK annual index values for waders.

Year OC AV RP GV KN SS DN BW BA CU RK TT1974/75 79 1 154 40 109 98 152 34 75 95 109 1081975/76 102 1 169 36 104 92 156 29 82 98 116 1271976/77 98 0 123 41 94 66 147 28 98 84 105 1331977/78 86 0 129 34 65 64 116 32 73 74 89 1321978/79 82 1 126 45 79 92 114 27 87 73 91 1481979/80 91 2 134 52 103 96 125 30 106 80 101 1221980/81 102 1 122 71 108 69 118 29 84 83 90 1171981/82 99 2 108 53 84 53 102 27 111 79 79 1201982/83 91 3 113 58 108 66 98 33 95 78 77 1141983/84 100 3 128 62 96 72 99 34 86 81 88 1331984/85 98 5 123 71 107 57 92 36 108 80 89 1331985/86 116 4 138 78 135 79 100 45 107 87 97 1661986/87 110 4 126 82 107 69 82 30 105 86 95 1491987/88 121 5 136 104 120 78 90 38 92 99 121 1781988/89 118 9 178 114 118 81 111 54 87 94 116 1551989/90 116 13 137 97 111 64 110 54 81 99 117 1581990/91 128 12 139 115 112 70 141 47 106 93 102 1441991/92 115 21 133 119 109 87 133 58 84 115 116 1551992/93 112 34 139 114 129 68 107 58 85 120 109 1361993/94 94 39 128 120 98 54 108 58 78 103 102 1351994/95 94 58 129 145 102 73 122 61 76 122 116 1371995/96 92 52 144 116 107 60 116 63 92 91 98 1271996/97 116 47 124 153 113 106 149 81 142 95 104 1191997/98 102 66 113 131 106 73 122 96 86 126 114 1141998/99 97 65 114 117 102 96 112 91 89 110 111 1191999/00 100 90 114 115 87 94 99 100 84 125 111 1132000/01 105 87 114 131 128 88 104 85 72 125 115 1202001/02 95 117 90 102 113 85 88 98 97 98 109 862002/03 100 100 100 100 100 100 100 100 100 100 100 100

Wader species codes OC OystercatcherAV Avocet RP Ringed PloverGV Grey PloverKN KnotSS Sanderling

DN Dunlin BW Black-tailed GodwitBA Bar-tailed GodwitCU Curlew RK RedshankTT Turnstone

Appendices304

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Table A9. Great Britain monthly index values for wildfowl 2002/03.

Sep Oct Nov Dec Jan Feb MarBS 1 0 7 51 61 100 2WS 1 6 96 84 35 100 43EW 0 1 13 49 100 21 0DB 0 26 80 79 97 100 28SU 74 100 66 60 68 58 48WN 12 69 100 89 95 84 36GA 79 100 100 99 100 98 55T. 36 71 80 77 100 66 35MA 89 98 97 97 100 72 39PT 5 34 100 75 89 71 9SV 60 92 93 94 93 85 100PO 27 40 90 94 100 81 17TU 87 89 100 98 91 92 73SP 5 10 13 37 100 51 26GN 2 8 66 80 100 100 87GD 40 43 80 97 100 83 43LG 93 100 82 73 70 63 52GG 100 95 81 66 69 65 73CA 90 100 83 80 78 72 63CO 88 100 95 94 81 61 38

Table A10. Northern Ireland monthly index values for wildfowl in 2002/03.

Sep Oct Nov Dec Jan Feb MarBS 0 0 0 100 0 83 0WS 0 0 97 0 100 0 6PB 45 100 88 17 16 12 12SU 4 21 47 62 100 59 39WN 17 82 100 71 71 40 59GA 92 100 71 45 46 90 86T. 69 75 58 45 50 100 57MA 100 84 46 39 55 34 21PT 23 44 100 92 40 69 11SV 34 42 85 82 100 29 15PO 5 22 64 63 100 45 7TU 23 44 72 82 100 79 60SP 0 1 3 18 70 100 16GN 2 3 91 50 81 56 100LG 54 100 64 52 73 29 26GG 88 100 43 33 38 28 64CA 85 100 30 37 42 42 40CO 89 100 53 38 45 30 23

Appendices 305

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APPENDIX 4. WEBS NATIONAL ALERTS

Table A11. Percentage change and Alerts over short- (5 years), medium- (10 years) and long-term (25 years or maximum available if less than 25 years) of counts in Great Britain (wildfowl and waders) and in the United Kingdom (waders). Symbols:

medium alert, high alert, medium increase, high increase, O no substantial change (from Austin et al 2004).

Alert Percentage changeGreat Britain

Firstwinter

Lastwinter 5-

year10-

year25-

yearmax-year

5-year

10-year

25-year

max-year

Mute Swan 74/75 00/01 O O 14 31 98Bewick's Swan 74/75 00/01 O -15 -37 99Whooper Swan 74/75 00/01 O 57 27 132European White-fronted Goose 74/75 00/01 -49 -58 -50Dark-bellied Brent Goose 74/75 00/01 O O -13 -24 90Shelduck 74/75 00/01 O O O -20 -22 7Wigeon 74/75 00/01 O O -3 27 73Gadwall 74/75 00/01 O 24 77 699Teal 74/75 00/01 O O 6 11 94Mallard 74/75 00/01 O -12 -28 -27Pintail 74/75 00/01 O O -18 -30 -24Shoveler 74/75 00/01 O O 3 3 60Pochard 74/75 00/01 O O -12 -8 -28Tufted Duck 74/75 00/01 O O O 8 13 8Goldeneye 74/75 00/01 O O O -14 -6 5Red-breasted Merganser 74/75 00/01 O O -18 7 80Goosander 74/75 00/01 O -25 -6 56Little Grebe 85/86 00/01 O 15 69 544Great Crested Grebe 82/83 00/01 O O 11 22 67Cormorant 86/87 00/01 O O -1 13 163Coot 82/83 00/01 O O O 11 24 31Oystercatcher 74/75 00/01 O O O 0 -12 8Avocet 74/75 00/01 73 223 >2000Ringed Plover 74/75 00/01 O -16 -25 -28Grey Plover 74/75 00/01 O O -17 2 196Knot 74/75 00/01 O O O 6 -5 15Sanderling 74/75 00/01 O O O 18 18 -6Dunlin 74/75 00/01 O -24 -26 -39Black-tailed Godwit 74/75 00/01 O 17 65 188Bar-tailed Godwit 74/75 00/01 O O O -24 -22 -13Curlew 74/75 00/01 O O O 14 17 31Redshank 74/75 00/01 O O O 6 1 -1Turnstone 74/75 00/01 O O -7 -25 -10

United Kingdom

Oystercatcher 74/75 00/01 O O O 2 -10 11Avocet 74/75 00/01 73 223 >2000Ringed Plover 74/75 00/01 O O -15 -24 -27Grey Plover 74/75 00/01 O O -17 2 196Knot 74/75 00/01 O O O 5 -4 10Sanderling 74/75 00/01 O O O 19 20 -4Dunlin 74/75 00/01 O -24 -25 -39Black-tailed Godwit 74/75 00/01 O 17 65 187Bar-tailed Godwit 74/75 00/01 O O -26 -23 -17Curlew 74/75 00/01 O O O 15 17 29Redshank 74/75 00/01 O O O 8 3 3Turnstone 74/75 00/01 O O -7 -25 -10

Appendices306

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APPENDIX 5. ERRATA TO WILDFOWL & WADER COUNTS 2000/01

The following errors in the 2000/01 report have come to light:

In accounts for most divers, grebes, Cormorant, herons, wildfowl, gulls and Kingfisher, it was

stated that annual site maxima were derived from the 12-month period data July to June; the

period used, however, was April to March. For the current report (2001-03) July to June has been

used, altering some peak counts (those that had occurred May or June) previously presented for

some species eg gulls.

Headings for site importance in tables for terns, summer visitors and passage species should

have read 1996 to 2000, rather than 96/97 to 2000/01, to indicate the 12-month period (January to

December) from which annual maxima were derived.

Page 9 Site importance: annual maxima for terns were derived from the 12-month period January to

December, not July to June as incorrectly stated.

Page 48 Christchurch Harbour was erroneously listed in the table of Mute Swan sites under ‘Important

sites not counted’.

Page 53 Figure 15 legend should have read ‘Annual indices for Pink-footed Goose in GB’.

Page 135 The records of Bonaparte’s Gulls were errors; none was recorded by WeBS in 2000/01.

Appendices 307

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APPENDIX 6. LOCATIONS OF WeBS COUNT SITES MENTIONED IN THIS REPORT

The location of all counts sites or areas mentioned in

this report are given here. Sites are listed

alphabetically, with the 1 km square OS grid

reference for the centre of the site, the habitat (H)

and the county or district. Note that this is not an

exhaustive list of WeBS sites counted in 2001–03,

simply those mentioned by name in this report.

Figure A1 shows the location of many of the more

important sites for waterbirds.

Key to habitat codes (the predominant habitat type is given for complex sites containing many different habitats)L LakeR ReservoirP Gravel or sand pitV RiverC Canal

M MarshS Sewage treatment worksE EstuaryO Open coastN Non-wetland

Site Site1-km sq 1-km sqH HWeBS Region WeBS RegionAbberton Reservoir TL9818 R EssexAberdeen Beach (Dee

Mouth to Don Mouth)NJ9507 O Grampian

Aberlady Bay NT4581 E LothianAdur Estuary TQ2006 E West Sussex Afan Estuary & Port Talbot Harbour

SS7488 E WestGlamorgan

Aignish Bay NB4932 O Western Isles Alde Complex TM4257 E SuffolkAlexandra Park TQ8010 R East Sussex Allington Gravel Pits SU4717 P HampshireAlnmouth NU2510 E N’th’mberlandAlt Estuary SD2903 E MerseysideAlton Water TM1436 R SuffolkAmpton Water TL877 L SuffolkAngler's Country Park Lake SE3716 P W YorkshireAnstruther Bay NO5603 O FifeAppin/Erriska/B’loch/Lismore NM9043 O StrathclydeApplecross Bay - Sand NG688 O HighlandAqualate Mere SJ7720 L StaffordshireArdleigh Reservoir TM0328 R EssexArdivachar Point NF7345 O Western Isles Ardrossan to Farland Head NS2045 O StrathclydeArlington Reservoir TQ5307 R SussexArran NR9535 O StrathclydeArun Valley TQ0314 V West Sussex Ashford Common Waterworks

TQ0869 S Surrey

Ashleworth Ham SO8326 M GlosAttenborough Gravel Pits SK5234 P NottsAuchenharvie Golf course NS2641 L Ayshire/AranAudenshaw Reservoirs SJ9196 R Gr. M’chesterAxe Estuary (Devon) SY250 E DevonAyr to North Troon NS3232 O StrathclydeBainton Pits TF1107 P CambsBalgray Reservoir NS5157 R RenfrewBallyroney Lake J229382 L DownBallysaggart Lough H7961 L TyroneBalranald RSPB Reserve NF7169 L Western Isles Bann Estuary C7935 E LondonderryBarcombe Mills Reservoir TQ434 R East Sussex Bardolf Water Meadows SY7796 M DorsetBardney Pits TF1168 P LincolnshireBarmston Ponds NZ3356 L Durham

Baron’s Haugh NS7555 L StrathclydeBarons Folly NT6426 L BordersBarrow Gurney Reservoir ST5367 R AvonBarton Broad TG3621 L NorfolkBarton Pits SK2017 P StaffordshireBaston Langtoft Gravel Pits TF1212 P LincolnshireBay of Sandoyne to Holme Sound

HY4800 O Orkney

Beadnell to Seahouses NU2231 O N’th’mberlandBeaulieu Estuary SZ4298 E HampshireBeauly Firth NH5848 E HighlandBeddington Sewage Farm TQ2966 S Gtr LondonBedfont & Ashford GPs TQ0872 P Gtr LondonBelfast Lough J4083 E DownBelvide Reservoir SJ8610 R StaffordshireBenacre Broad TM5383 L SuffolkBenbecula NF8150 N Western IslesBesthorpe & Girton GPs SK8165 P NottsBewl Water TQ6733 R SussexBirgham Haugh NT7938 N BordersBittell Reservoirs SP0275 R Hereford &

WorcsBlack Cart Water NS4767 M StrathclydeBlackmoorfoot Reservoir SE0912 R W YorkshireBlackwater Estuary TL9307 E EssexBlagdon Lake ST5150 R AvonBlatherwyke Lake SP9796 L NorthantsBlenheim Park Lake SP4316 L OxfordshireBlithfield Reservoir SK0524 R StaffordshireBlucks Pool to Freshwater West

SR8898 O Dyfed

Blunham Gravel Pits TL1551 P BedfordshireBlyth Estuary(Northumberland)

NZ3082 E N’th’mberland

Blyth Estuary (Suffolk) TM4675 E SuffolkBoghill Fields (Coleraine) C8734 N LondonderryBolton-on-Swale Gravel Pits SE2498 P N YorkshireBosherston Lakes SR9794 L DyfedBothal Pond NZ2487 L N’th’mberlandBough Beech Reservoir TQ4947 R KentBoultham Mere SK9571 P LincolnshireBrandesburton Ponds West TA0948 P HumbersideBrading Harbour SZ6388 E Isle of Wight Bradley Pools SK225 L Derbyshire

Appendices308

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Site 1-km sq H WeBS RegionBraewick Loch HU2478 L ShetlandBramshill Park Lake SK7560 L HampshireBranahuie Saltings NB4631 O Western Isles Brent Reservoir TQ2287 R Gtr LondonBressay Sound HU4842 O ShetlandBreydon Water & Berney Marshes

TG4907 E Norfolk

Broadford Bay NG6523 O SkyeBroad Water Canal J1462 C AntrimBrogborough Pit SP9739 P BedfordshireBuckden/Stirtloe Gravel Pits TL2066 P CambsBurghfield Gravel Pits SU6870 P BerkshireBurry Inlet SS5096 E W Glamorgan,

DyfedBusbridge Lakes SU9742 L SurreyBush River: Deepstown C9434 V AntrimBute NS0761 L StrathclydeCaerlaverock WWT NY0565 E D & GallowayCaistron Quarry NU0001 P N’th’mberlandCaithness Lochs ND1859 L HighlandCamas Dubh-aird NG783 O HighlandCambois to Newbiggin NZ3084 O N’th’mberlandCamel Estuary SW9474 E CornwallCameron Reservoir NO4711 R FifeCanary Road H8755 M ArmaghCannop Ponds SO6010 L GlosCardigan Bay SH5020 O Gwynedd,

DyfedCarlingford Lough J2013 E DownCarlingford to Newcastle J3718 O DownCarmarthen Bay SN2501 E DyfedCarse of Stirling NS7196 N CentralCarsebreck & Rhynd Lochs NN8609 L TaysideCarsington Water SK2451 R DerbyshireCastle Howard Lake SE7170 L N YorkshireCastle Loch (Lochmaben) NY0881 L Dumfries &

GallowayCastle Marshes Reserve TM4791 M SuffolkCemlyn Bay & Lagoon SH3393 O GwyneddChasewater SK0307 R W MidlandsChat Moss SJ7096 M Gtr M’chesterCheddar Reservoir ST4454 R SomersetChelmarsh Reservoir SO7387 R ShropshireCherwell Valley SP4929 R OxfordshireChetwynd Pool SJ7420 L ShropshireChew Valley Lake ST5659 R AvonChichester Gravel Pits SU8703 P West Sussex Chichester Harbour SU7700 E West Sussex Chillington Hall Pool SJ8550 L StaffordshireChristchurch Harbour SZ1792 E DorsetChurch Wilne Reservoir SK4632 R DerbyshireClachan/Whitehouse NR7959 N StrathclydeClatto Reservoir NO3607 R FifeClea Lakes J506557 L DownCleddau Estuary SN0005 E DyfedCley Marsh TG0644 M NorfolkClifford Hill Gravel Pits SP8061 P NorthantsClumber Park Lake SK6374 L NottsClwyd Estuary SJ0079 E Clwyd

Site 1-km sq H WeBS Region

J704337L

N’th’mberland

Cromarty Firth TG3719 L

REssex

Cuckmere EstuarySU9145

PDevon

Deben EstuarySJ2675

ECheshire

Derwent ReservoirTL7796

Doon EstuaryNH7384 E

M

Clyde Estuary NS3576 E StrathclydeCochrage Loch NO1350 L Perth & K’rossColl NM2055 N StrathclydeColliford Reservoir SX1871 R CornwallColne Estuary TM0614 E EssexColne Valley Gravel Pits TQ0489 P Gtr LondonColney Gravel Pits TG1708 P NorfolkColonsay/Oronsay NR3896 N StrathclydeColt Crag Reservoir NY948 R N’th’mberlandColwick Country Park SK6039 L NottsColwyn Bay SH9079 O ClwydConnaught Water TQ4095 L EssexCoombe Country Park SP3974 L WarwickshireCoombe Hill Canal SO8626 C GlosCors Caron (CorsTregaron)

SN6863 M Dyfed

Cors Crugyll SH3376 M AngleseyCotswold Water Park (East) SU1999 P Glos, Oxon Cotswold Water Park (West)

SU0595 P Glos, Wilts

Craigalea to Newcastle O DownCresswell Pond NZ2894 N’th’mberlandCresswell to Chevington Burn

NZ2895 O

Crichel Lake ST9907 L DorsetCriddling Stubbs Quarry Pool

SE5120 P W Yorkshire

NH7771 E HighlandCrome's Broad NorfolkCropston Reservoir SK5410 LeicestershireCrouch-Roach Estuary TQ8496 ECrowdy Reservoir SX1483 R CornwallCroxall Pits SK1814 P Staffordshire

TV517 E East Sussex Cuttmil Ponds L SurreyDagenham Chase GP TQ5186 Gtr LondonDart Estuary SX846 EDarwell Reservoir TQ7121 R SussexDean Heritage Museum Lake

SO6610 L Glos

TM2942 E SuffolkDee Estuary (England/Wales)

E Merseyside,Cheshire,Clwyd

Dee Estuary (Scotland) NJ9505 GrampianDee Flood Meadows SJ4159 VDeeping St James GPs TF1808 P LincolnshireDengie Flats TM0300 E Essex

NZ0251 R DurhamDidlington Lakes P NorfolkDinnet Lochs (Lochs Davan & Kinord)

NJ4800 L Grampian

Dinton Pastures SU7872 M BerkshireDitchford Gravel Pits SP9468 P NorthantsDon Mouth to Ythan Mouth NJ9815 O Grampian

NS3219 O StrathclydeDornoch Firth HighlandDoxey Marshes SSSI SJ9024 StaffordshireDrakelow Gravel Pit SK2320 P DerbyshireDraycote Water SP4469 R Warwickshire

Appendices 309

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Site 1-km sq H WeBS Region

Drummond PondNZ2796

EDown

Durham Coast SN6394

ENorthants

East Park HY7241

ONorfolk

Eccup ReservoirNO4719

LDevon

Entrance to Deer/Shapinsay Sounds

P

Drumgay Lough H2448 L FermanaghNN8518 L Tayside

Druridge Pool L N’th’mberlandDuddon Estuary SD2081 CumbriaDundrum Bay J4235 EDungeness Gravel Pits TR0619 P KentDupplin Loch NO0320 L Tayside

NZ4349 O DurhamDyfi Estuary E DyfedDysynni Estuary SH5702 GwyneddEarls Barton Gravel Pits SP8966 PEarlsferry Links to Anstruther

NO5302 O Fife

East Chevington Pools NZ2799 L N’th’mberlandTA1213 L Humberside

East Sanday Coast O OrkneyEast Unst HU6508 ShetlandEast Wretham Meres TL9088 LEasterloch/Uyeasound HP5901 O ShetlandEasting/Sand Wick HP6201 O Shetland

SE2941 R W Yorkshire Eden Estuary E FifeEgilsay HY4730 OrkneyElfordleigh Golf & Country Club

SX5458 L

Ellesmere Lakes SJ4035 L ShropshireEmberton Gravel Pits SP8850 P Bucks

HY5513 O Orkney

Erme Estuary SX6249 E DevonEversley Cross & Yateley GP SU8601 HampshireExe Estuary SX9883 E DevonEyebrook Reservoir SP8595 R LeicestershireFairburn Ings SE4627 P N YorkshireFairfield SSSI TQ9626 M KentFal Complex SW8541 E CornwallFala Flow NT4258 L LothianFarmoor Reservoirs SP4406 R OxfordshireFarmwood Pool SJ8173 L CheshireFarne Islands NU2136 O N’th’mberlandFen Drayton Gravel Pits TL3470 P CambsFernworthy Reservoir SX6684 R DevonFiddler’s Ferry Power Station Lagoons

SJ5585 P Cheshire

Fincastle Loch NN8762 L TaysideFindhorn Bay NJ0462 E GrampianFleet Pond SU__55 L HampshireFleet/Wey SY6976 E DorsetFonthill Lake ST9331 L WiltshireForeland SZ6584 O Isle of Wight Forest of Dean Ponds SO6010 L GwentFort Georgo to Nairn NH7958 E HighlandFort Henry Ponds & Exton Park Lake

SK9311 L Leicestershire

Forth Estuary NT2080 E Lothian,Central, Fife

Forth & Teith Valley NS7595 N CentralForyd Bay SH4559 E GwyneddFowey Estuary SX1254 E Cornwall

Site 1-km sq H WeBS Region

Strathclyde

Frampton Pools SO757 P GlosGaddon Loch NO2812 L FifeGadloch NS6471 L StrathclydeGare Loch. NS2585 L StrathclydeGatton Park TQ273 L SurreyGerrans Bay SW8937 O CornwallGigha, Kintrye NR6450 N ArgyllGirvan to Turnberry NS2002 O StrathclydeGladhouse Reservoir NT2953 R LothianGlynde Levels TQ4709 M East Sussex Godmanchester Gravel Pit TL2672 P CambsGoring TQ1001 O West SussexGourock to Largs NS1867 O Argyll & Bute Grafham Water TL1568 R CambsGrand Western Canal:

Greenway Bridge to Ayshford Bridge

ST0213 C Devon

Great Pool Westwood Park SO8763 L Hereford & Worcs

Grimsthorpe Lake TF0222 L LincolnshireGrindon Lough NY8067 L N’th’mberlandGroby Pool SK5208 L LeicestershireGrouville Marsh WV6949 M Channel Isles Gruinard Bay NG9393 O HighlandGuernsey Shore WV27 O Channel Isles Gunton Park Lake TG2234 L NorfolkHacosay, Bluemull & Colgrave Sounds

NU5697 O Shetland

Haddo House Lakes NJ8734 L GrampianHagnaby Lock Fen TF3459 L LincolnshireHallington Reservoir NY976 R N’th’mberlandHamford Water TM2225 E EssexHamilton Low Parks & Strathclyde Park

NS7257 L Strathclyde

Hampton & Kempton Reservoirs

TQ1269 R Gtr London

Hanningfield Reservoir TQ7398 R EssexHardley Flood TM3899 M NorfolkHarewood Lake SE3144 L W YorkshireHarrold-Odell Country Park SP9557 P BedfordshireHarrow Lodge Park TQ5286 L Gtr LondonHoughton Regis Chalk Pit TL0123 P BedfordshireHaweswater Reservoir NY4814 R CumbriaHay-a-Park Gravel Pits SE3658 P N YorkshireHayle Estuary SW5537 E CornwallHeadley Mill Pond SU8138 L HampshireHeaton Park Reservoir SD8205 R Gtr M'chester Heigham Holmes TG4420 M NorfolkHelford Estuary SW7526 E CornwallHen Reedbeds TM4777 V SuffolkHerne Bay TR1768 O KentHickling Broad TG4121 L NorfolkHilfield Park Reservoir TQ1596 R HertfordshireHillsborough Forest Lake J2458 L DownHillside Marsh SU7550 M HampshireHirsel Lake NT8240 L BerwickshireHogganfield Loch NS6467 LHolbeach St Matthew TF4134 O LincolnshireHolburn Moss NU0536 L N’th’mberland

Appendices310

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Site 1-km sq H WeBS Region

TG3116

O

Site 1-km sq H WeBS Region

Langtoft West End GPs

Lancashire

NS4388

Holkham & Burnham Overy Fresh Marshes

TF8644 M Norfolk

Holkham Bay TF8845 E NorfolkHolkham Lake TF8843 L NorfolkHolland Marshes TM2117 M EssexHollowell Reservoir SP6872 R NorthantsHolme TF7044 O NorfolkHolme Pierrepoint GPs SK6239 P NottsHoly Loch to Toward Point NS1674 O ArgyllHope Carr Reserve SJ6698 N Gtr M’chesterHornsea Mere TA1947 L HumbersideHorsey Mere TG4422 L NorfolkHoughton Green Pool SJ6292 L CheshireHoveringham Gravel Pits SK707 P NottsHoveton Great Broad L NorfolkHowick to Beadnell NU2327 O N’th’mberlandHule Moss NT7149 L BordersHumber Estuary TA2020 E Humberside,

LincolnshireHunterston Sands NS1848 O Ayshire &

ArranHuntingdon Racecourse GPs TL1972 P CambsHurleston Reservoir SJ6255 R CheshireHurworth Burn Reservoir NZ4033 R DurhamIngrebourne Valley TQ5485 L Gtr M’chesterInland Sea SH2779 E GwyneddInner Moray Firth NH6752 E HighlandIsland of Papa Westray HY4952 N OrkneyIslands of South Barra NL6189 N Western Isles Islay NR3560 N StrathclydeIsle of Cumbrae NS1656 O StrathclydeIslesteps NX9772 V D & Galloway Jersey Shore WV6249 Channel Isles Jura NR5672 N StrathclydeKedleston Park Lake SK3141 L DerbyshireKeills Peninsular & Isle of Danna

NR7385 N Argyll

Kenfig Pools SS7981 L Mid & S Glamorgan

Kennington Park TQ5681 L EssexKessingland Levels TM5185 L SuffolkKilconquhar Loch NO4801 L FifeKillimster Loch ND3056 L HighlandKillough Harbour J5437 O DownKilkeel to Lee Stone Point J3214 O DownKirkabister/Dury Voe HU4550 O ShetlandKing George V Reservoir TQ3796 R Gtr LondonKing George VI Reservoir TQ0473 R SurreyKing’s Dyke Pits TL2397 P CambsKings Bromley Gravel Pits SK1116 P StaffordshireKings Mill Reservoir SK5159 R NottsKingsbridge Estuary SX7411 E DevonKinnordy Loch NO3654 L AngusKirkby-on-Bain Gravel Pits TF2360 P LincolnshireKnight & Bessborough Rsvrs TQ1268 R SurreyKyle of Durness NC375 E HighlandKyle of Tongue & Tongue Bay

NC5859 O Highland

Lackford Gravel Pits TL7971 P Suffolk

Lade Sands TR0921 O KentLake of Menteith NN5700 L CentralLakenheath Fen TL7186 M SuffolkLancaster Canal SD4766 C LancashireLangford Lowfields GPs SK8259 P NottsLangstone Harbour SU6902 E Hampshire

TF1111 P LincolnshireLarne Lough D4200 E AntrimLee Valley Gravel Pits TL3702 P Hertfordshire,

EssexLeighton Moss SD4874 LLeighton & Roundhill Rsvrs SE1678 R N YorkshireLentran NH5747 E HighlandLeybourne/New Hythe GPs TQ6959 P KentLinacre Reservoirs SK3372 R DerbyshireLindisfarne NU1041 E N’th’mberlandLittle Loch Broom NH0491 O HighlandLittle Paxton Gravel Pits TL1963 P CambsLivermere TL8771 L SuffolkLlangorse Lake SO1326 L PowysLlanrhystud - Morfa Mawr SH4571 M AngleseyLlyn Alaw SH3986 R AngleseyLlyn Coron SH3770 L AngleseyLlyn Padrig SH3672 L AngleseyLlyn Traffwll SH3276 L GwyneddLlynnau Y Fali SH3177 L AngleseyLlys-y-fran Reservoir SN0324 R DyfedLoch a’Phuill NL9541 L StrathclydeLoch Ashie NH624 R HighlandLoch Bee (South Uist) NF7744 L Western Isles Loch Bhasapoll NL9746 L TireeLoch Broom NH098 O HighlandLoch Coalisport NR7677 L StrathclydeLoch Ellrig NS8874 L CentralLoch Eriboll NC4359 O HighlandLoch Ewe NG8788 L HighlandLoch Eye NH8379 L HighlandLoch Fleet Complex NH7896 E HighlandLoch Fyne NR8881 L ArgyllLoch Gairloch NG7975 O HighlandLoch Garten NH9718 L HighlandLoch Gelly NT2092 L FifeLoch Grogary, Loch nam

Feithean & Hougharry Lochan

NF7171 L Western Isles

Loch Gruinart NR2971 E StrathclydeLoch Indaal NR3261 E StrathclydeLoch Insh & Spey Marshes NH8304 L HighlandLoch Ken NX7168 R D & Galloway Loch Kishorn NG830 O HighlandLoch Leven NO1401 L TaysideLoch Linnhe: CamasShallachain

NM9862 L Highland

Loch Linnhe: Corran Ferry - Onich

NN011 O Highland

Loch Lomond L StrathclydeLoch Long NO2938 L AngusLoch Long/Loch Goil NS2192 L StrathclydeLoch Mahaick NN7006 L Central

Appendices 311

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Site 1-km sq H WeBS Region

L

Loch of Tankerness

Loch Sheil: West NM7169

Loch Mullion NN9833 L TaysideLoch of Beith HU3689 L ShetlandLoch of Boardhouse HY2725 L OrkneyLoch of Brow HU3815 L ShetlandLoch of Clumlie HU4017 ShetlandLoch of Harray HY2915 L OrkneyLoch of Lintrathen NO2754 L TaysideLoch of Sabiston HY2922 L OrkneyLoch of Skene NJ7807 L GrampianLoch of Spiggie HU3716 L ShetlandLoch of Stenness NY2812 L OrkneyLoch of Strathbeg NK0758 L GrampianLoch of Swannay HY3127 L Orkney

HY5109 L OrkneyLoch of the Clans NH9353 L HighlandLoch of Wester ND3259 L HighlandLoch nan Gabhar NM9663 L HighlandLoch Ordais & Port Mhor Bragar

NB2849 O Western Isles

Loch Ore NT1695 L FifeLoch Paible (N. Uist) NF7268 L Western Isles Loch Ryan NX0565 E D & Galloway

L HighlandLoch Spynie NJ2366 L GrampianLoch Tullybelton NO0034 L TaysideLoch Urrahag NB3247 L Western IslesLoch Watten ND2256 L HighlandLochs Beg & Scridain NM5027 L StrathclydeLochs Heilen & Mey ND2568 L HighlandLoe Pool SW6424 L CornwallLondon Wetland Centre TQ2277 R Gtr LondonLong Loch NS4752 L StrathclydeLongnewton Reservoir NZ3616 R ClevelandLongside Lake TQ0168 P SurreyLongueville Marsh WV6748 M Channel Isles Lonsdale Road Reservoir TQ2177 R Gtr LondonLooe Estuary SX2553 E CornwallLossie Estuary NJ2470 E GrampianLost & Golding Hill &

Baldwins Hill PondsTQ4297 L Essex

Lough Aghery J2853 L DownLough Foyle C6025 E LondonderryLough Money J5345 L DownLoughs Neagh & Beg J0575 L Down, Antrim,

Londonderry,Tyrone,Armagh

Lower Derwent Valley SE6938 M HumbersideLower Lough Erne H1060 L FermanaghLower Teviot Valley NT6928 V BordersLower Windrush Valley GPs SP4004 P OxfordshireLunning/Lunna Holm HU5375 O ShetlandLynford Gravel Pit TL8194 P NorfolkMachrihanish NR6522 N StrathclydeMaer Lake SK2070 M CornwallMalltraeth Marsh RSPB SH4571 M AngleseyMarizion Marsh SW5131 L CornwallMarsh Lane Gravel Pits TL3069 P CambsMarston Sewage Treatment

Works

SK9042 S Lincolnshire

Site

Orkney

1-km sq

HY4010

H

N

WeBS Region

Orkney

Martin Mere SD4105 L LancashireMasham Gravel Pits SE2182 P N YorkshireMeadow Lane Gravel Pits TL3270 P CambsMedway Estuary TQ8471 E KentMelbost Sands & Tong Saltings

NB4534 E Western Isles

Mere Sands Wood NR SD4415 L LancashireMersehead RSPB Reserve NX905 E D & Galloway Mersey Estuary SJ4578 E CheshireMiddle Tame Valley GPs SP2096

SM8421

E

Nosterfield Gravel Pits

P Staffordshire,Warwickshire

Middle Yare Marshes TG3504 M NorfolkMillbrook Clay Pit TL0041 P BedfordshireMilldam & Balfour Mains Pools

HY4817 L Orkney

Minsmere TM4666 L SuffolkMoine Mhor NR8293 N ArgyllMonach Isles NF6262 O Western Isles Monikie Country Park NO5038 R TaysideMontrose Basin NO6958 E TaysideMoray Coast NJ3067 O GrampianMoray Firth NH8060 E HighlandMorecambe Bay SD4070 E Lancashire,

CumbriaMunlochy Bay NH6752 E HighlandNE Glamorgan Moorland Pools

SO0808 L E Glamorgan

Nene Washes TF3300 M CambsNew Road Pits Tl1549 P BedfordshireNewgale Beach O DyfedNewport Pagnell Gravel Pits SP885 P BucksNewtown Estuary SZ4291 E Isle of Wight Nocton Fen TF1065 M LincolnshireNorbury Pond SJ9185 L Gtr M'chester North Bay (South Uist) NF7546 O Western Isles North Cave Gravel Pits SE8833 P HumbersideNorth Fetlar HU6295 O ShetlandNorth Norfolk Coast TF8546 NorfolkNorth Ronaldsay HY7655 N OrkneyNorth Ronaldsay Lochs HY7655 L OrkneyNorth Sutherland NC5060 N HighlandNorth Uist NF8370 N Western Isles North Uist (West Coast) NF7070 O Western Isles North Warren & Thorpeness Mere

TM4658 L Suffolk

North West Solent SZ3395 E HampshireNorton Marsh TF8244 M Norfolk

SE2880 P N YorkshireNunnery Lakes TL8781 L NorfolkOld Moor Wetlands (Wath Main Ings)

SE4302 L S Yorkshire

Orwell Estuary TM2238 E SuffolkOsterley Park Lakes TL1478 L Gtr LondonOtmoor SP564 M OxfordshireOuse Washes TL5394 M CambsOuse & Lairo Water HY5019 L OrkneyOuter Ards J6663 O Down

Appendices312

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Site Site1-km sq 1-km sqH HWeBS Region WeBS RegionPagham Harbour SZ8796 E West Sussex Panshanger Flash TL2812 L HertfordshirePar Sands Pools SX0853 L CornwallPassfield Lake SU8234 L HampshirePeatmoor Lagoon SU1186 L WiltshirePegwell Bay TR3563 E KentPitsford Reservoir SP7669 R NorthantsPool of Virkie HU3911 O ShetlandPoole Harbour SY9988 E DorsetPort Meadow SP4908 M OxfordshirePortsmouth Harbour SU6204 E HampshirePortworthy Mica Dam SX5660 P DevonPriory Country Park TL0749 P BedfordshirePriory Water SK7118 L LeicestershirePugneys Country Park Lakes SE3317 P W YorkshirePulfin Bog TA0544 P HumbersidePurfleet Chalk Pit TQ5778 P EssexQueen Elizabeth II Reservoir TQ1167 R SurreyQueen Mary Reservoir TQ0769 R SurreyQueen Mother Reservoir TQ0076 R BerkshireQueens Valley Reservoir WV5652 R Channel Isles Quendale Bay HU3712 O ShetlandRadwell Gravel Pits TL017 P BedfordshireRanworth & Cockshoot Broads

TG3515 L Norfolk

Ravensthorpe Reservoir SP6770 R NorthantsRavenstruther NS9343 L LanarkshireReddish Hall SJ6989 L Gtr M’chesterRedmires Reservoirs SK2685 R S Yorkshire Red Point to Port Henderson

NG7470 O Highland

Reedham Water TG3618 L NorfolkRescobie Loch NO5151 L AngusRevesby Reservoir TF3067 R LincolnshireRhunahaorine NR7049 N ArgyllRibble Estuary SD3825 E LancashireR. Avon: Fordingbridge to Ringwood

SU1510 M Hampshire

R. Avon: Ringwood to Christchurch

SZ1499 M Hampshire

R. Avon: Salisbury to Fordingbridge

SU1721 V Wiltshire

R. Cam: Kingfishers Bridge TL5473 V CambsR Cam: Upware-Dimmocks Cote

TL5371 V Cambs

R. Clyde: Carstairs Junction NS9744 V StrathclydeR. Clyde: Carstairs to Thankerton

NS9842 V Strathclyde

R. Devon: Kersiepow Ponds NS8996 V CentralR. Devon: Tullibody Bridge NS8596 V CentralR. Eamont: Watersmeet - Pooley Bridge

NY5329 V Cumbria

R. Earn: Lawhill Oxbows NN9517 V TaysideR. Earn: Millands Marsh & Floods

NN9917 V Perth& K’ross

R. Eden: Warcop to Little Salkeld

NY6424 V Cumbria

R. Foyle: Grange C3606 V TyroneR. Idle: Bawtry to Miserton SK7195 V NottsR. Irwell SD7903 V Gtr M'chester

R. Lagan: Flatfield J1961 V DownR. Nith: Keltonbank toNunholm and Dalscone

NX9774 V D & Galloway

R. Severn/Vyrnwy Confluence

SJ3215 V Shropshire

R. Spey: Boat of Balliefirth NH9922 V HighlandR. Tay: Dunkeld NO0042 V TaysideR. Teith: Daldorn to Deanston

NN6902 V Central

R. Test: Broadlands Estate SU350 V HampshireR. Test: Fullerton to Stockbridge

SU3535 V Hampshire

R. Teviot: Kalemouth to Roxborough

NT7030 V Borders

R. Tweed: Kelso to Coldstream

NT7737 V Borders

R. Tweed: Magdalenehall NT6331 V BordersR. Wye: Bakewell to Haddon

SK2366 V Derbyshire

R. Wye: Putson SO5138 V Hereford & Worcs

Rivers Eamont & Eden:Honeypot to Edenhall

NY5631 V Cumbria

Rodbourne Sewage Works SU1385 S WiltshireRomney Marsh TR050 M KentRossie Bog NO2711 M FifeRostherne Mere SJ7484 L CheshireRough Firth NX8453 E D & Galloway Rova Head to Kirkabister HU4842 O ShetlandRoxton Gravel Pits TL1653 P BedfordshireRufford Lake SK6465 L NottsRutland Water SK9207 R LeicestershireRyde Pier to Puckpool Point SZ6092 O Isle of Wight Rye Harbour & Pett Level TQ9418 E East Sussex Sandbach Flashes SJ7259 L CheshireScaling Dam Reservoir NZ7412 R ClevelandScapa Flow HY4000 O OrkneyScolt Head TF8046 E NorfolkSeahouses to Budle Point NU2231 O N’th’mberlandSeaton Gravel Pits TR2258 P KentSennowe Park Lakes TF9825 L NorfolkSevern Estuary ST5058 E Glos, Avon,

Somerset,Gwent, E Glam

Shetland Isles HU4060 N ShetlandShipley Country Park SK4343 R DerbyshireShipton-on-Cherwell Quarry SP4717 P OxfordshireShrigley Lake J518544 L DownShustoke Reservoir SP2391 R WarwickshireSiblyback Reservoir SX2370 R CornwallSkinflats NS9284 E CentralSlains Lochs NK0230 L GrampianSlamannan Plateau NS8474 N CentralSnetterton Gravel Pits TL9991 P NorfolkSnettisham TF6535 E NorfolkSolway Estuary NY1060 E Cumbria, D &

GallowaySolway Firth NY0048 E Cumbria, D &

Galloway

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Site Site1-km sq 1-km sqH HWeBS Region WeBS RegionSomerset Levels ST4040 M SomersetSomersham Gravel Pits TL3779 P CambsSonning Gravel Pit SU7475 P OxfordshireSoudley Ponds SO6611 L GlosSound of Barra NF7509 O Western Isles Sound of Gigha NR6950 O ArgyllSound of Harris NF9788 O Western Isles Sound of Jura NR6575 O StrathclydeSound of Tarransay NG0498 O Western Isles South Fetlar HU6287 O ShetlandSouth Ford NF7747 O Western Isles Southill Lake TL1443 L BedfordshireSouth Iver Gravel Pits TQ0377 P BucksSouth Lancashire Mosses SD4015 M LancashireSouth Mainland HU4234 N ShetlandSouth Stoke TQ0210 V West Sussex South Uist NF8032 N Western Isles South Uist (West Coast) NF7332 O Western Isles South Walls ND3089 N OrkneySouth West Lancashire SD4015 N LancashireSouth Yell Sound HU4081 O ShetlandSouthampton Water SU4507 E HampshireSpade Oak Gravel Pit SU8887 P BucksSpey Mouth NJ3465 E GrampianSpittal to Cocklawburn NU0250 O N’th’mberlandSprotbrough Flash SE5300 L S YorkshireSt Andrews Bay NO5121 O FifeSt Benet’s Levels TG3815 M NorfolkSt Johns Loch ND2272 L HighlandSt Mary’s Island NZ3475 O N’th’mberlandStaines Moor Gravel Pits TQ0373 P SurreyStaines Reservoirs TQ0575 R SurreyStanford Reservoir SP6080 R LeicestershireStanford Training Area TL8695 L NorfolkStanwick Gravel Pits SP9773 P NorthantsStevenston Point (Irvine to Saltcoats)

NS2839 E Strathclyde

Stewartby Lake TL0042 P BedfordshireStiffkey Fen TF9843 L NorfolkStockgrove Country Park SP9229 L BedfordshireStodmarsh NNR & Collards Lagoon

TR2061 L Kent

Stoke Newington Reservoirs TQ3287 R Gtr LondonStour Estuary TM1732 E Essex, SuffolkStrangford Lough J5560 E DownStranraer Lochs NX1161 L D & GallowayStratfield Saye SU7061 L HampshireStrathearn NN8819 N TaysideStrawberry Hill Ponds TQ4196 L EssexStudland Bay SZ0383 O DorsetSullom Voe HU3773 O ShetlandSutton & Lound Gravel Pits SK6985 P NottsSwale Estuary TQ9765 E KentSwanbourne Lake TQ0108 L West Sussex Swanholme Lake SK9468

OW Yorkshire

Tamar Complex

NH9487 OP

Devon

Tay/Isla Valley NZ5528

ENorfolk

Thanet Coast HU5565

ENorfolk

Thrapston Gravel Pit NX7362

Tiree NL9741 TF7544

R

Traeth Lafan (Lavan Sands) NG0798

NHertfordshire

Tweed Estuary SU7875

E

Lothian

Upper Loch TorridonH3231

RDown

Uyea Sound HU2548

OGlos

L LincolnshireSwansea Bay SS6391 W Glamorgan Swillington Ings SE3828 PSwithland Reservoir SK5513 R LeicestershireTalkin Tarn NY5458 L Cumbria

SX4363 E Devon,Cornwall

Tarbat Ness to Rockfield HighlandTattershall Pits TF2057 LinconshireTaw-Torridge Estuary SS4733 ETay Estuary NO3225 E Fife, Tayside Tayinloan NS6945 N Ayshire &

ArranNO1438 L Tayside

Tees Estuary E ClevelandTeign Estuary SX8772 DevonTerrington East TF5526 OTerringtonWest TF5026 O NorfolkThames Estuary TQ7880 E Kent, Essex,

Gtr LondonTR2669 O Kent

The Houb (Whalsay) O ShetlandThe Strand (Colonsay) NR360 ArgyllThompson Water TL9194 LThoresby Lake SK6370 L NottsThorpe Water Park TQ0268 P Surrey

SP9979 P NorthantsThreave Estate V D & GallowayThreipmuir & Harlaw Rsvrs NT1764 R LothianThursley Lake SU9239 L SurreyThurso Bay ND1167 O HighlandThwaite Flat & Roanhead Ponds

SD214 L Cumbria

N StrathclydeTitchwell O NorfolkToft Newton Reservoir TF0387 LincolnshireTophill Low Reservoirs TA0748 R HumbersideTottenhill Gravel Pits TF6311 P NorfolkTraeth Coch (Red Wharf Bay)

SH5480 E Anglesey

SH6474 E GwyneddTraigh Luskentyre E Western Isles Traighear NF8276 Western IslesTring Reservoirs SP9113 RTrinity Broads TG4614 L NorfolkTundry Pond SU7752 L Hampshire

NT9853 E N’th’mberlandTwyford Gravel Pits P BerkshireTyne Estuary NZ2464 N’th’mberland,

DurhamTyninghame Estuary NT6379 ETyrella Shore J4735 O DownUpper Cowgill Reservoir NT0029 R Strathclyde

NG8955 L HighlandUpper Lough Erne L FermanaghUpper Tamar Reservoir SS2812 DevonUpper Quoile River J4745 VUpper Tay NN9557 N TaysideUpton Warren LNR SO9367 L Hereford &

WorcsHP5902 O Shetland

Vaila Sound/Gruting Voe O ShetlandVirkie/Quendale HU3911 ShetlandWalmore Common SO7425 M

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Site 1-km sq H WeBS RegionWalthamstow Reservoirs TQ3589 R Gtr LondonWalton Lock SJ6086 C CheshireWarkworth Lane Ponds

E

NZ2793 L N’th’mberlandWarren and Lion GorgesWash TF5540 Lincolnshire,

NorfolkWash Strategic Area Watermead Gravel Pits SK6008 P LeicestershireWater Sound ND4394 O OrkneyWatermill Broad TL7796 P NorfolkWaulkmill Glen and Littleton Reservoirs

NS5257 R Strathclyde

Weirwood Reservoir TQ3934 R East Sussex Welbeck Estate SK5773 L NottsWellington Country Park SU7362 L HampshireWellington Gravel Pits SO5047 P Hereford &

WorcsWells-next-the-Sea TF9145 O NorfolkWemyss Bay to Fairlie NS2057 O Ayshire &

ArranWest Voe of Sumburgh HU3909 O ShetlandWest Water Reservoir NT1252 R BordersWest Whalsay and Sounds HU5565 O ShetlandWestfield Marshes ND0664 M HighlandWeybread Pits TM2481 P Suffolk

Site 1-km sq H WeBS RegionWhiteness to Scarvister HU4335 O ShetlandWhiteness to Skelda Ness HU3543 O ShetlandWhitlingham Country Park TL2508 L NorfolkWhitrig Standing Water NT6234 M BordersWicken Fen TL5570 L CambsWidewall Bay ND4292 O OrkneyWigan Flashes SD5803 L Gtr M'chester Wigtown Bay NX4456 E D & Galloway Willen Lake SP8741 R BucksWilliam Girling Reservoir TQ3694 R Gtr LondonWillington TL1050 R BedfordshireWindermere SD3995 L CumbriaWintersett & Cold Hiendley Reservoirs

SE374 R W Yorkshire

Woburn Park Lakes SP9632 L BedfordshireWoolston Eyes SJ6588 P CheshireWraysbury Gravel Pits TQ0073 P BerkshireWraysbury Pond TQ0073 P BerkshireWraysbury Reservoir TQ0274 R SurreyWynyard Lake NZ4224 L ClevelandYar Estuary SZ3588 E Isle of Wight Yealm Estuary SX5449 E DevonYell Sound HU4285 O ShetlandYthan Estuary NK0026 E Grampian

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Key to Figure A1

Abberton Reservoir 169Adur Estuary 243Alaw Reservoir 125Alde Complex 163Alnmouth 73Alt Estuary 115Artro Estuary 131Attenborough GPs 140Auchencairn Bay 81Avon Estuary 219Axe Estuary 225Ballo Reservoir 33Bann Estuary 89Baston/Langtoft GPs 145Beaulieu Estuary 231Belfast Lough 98Belvide Reservoir 136Berney Marshes 159Black & White Lochs (Loch Inch) 87Blackwater Estuary 170Blagdon Lake 195Blithfield Reservoir 137Blyth (Northumberland) Estuary 75Blyth (Suffolk) Estuary 161Brading Harbour 237Braint Estuary 128Breydon Water 160Burry Inlet 200Camel Estuary 206Cameron Reservoir 32Carlingford Lough 93Carmarthen Bay 201Carron Valley Reservoir 41Carsebreck & Rhynd Lochs 40Castle Lo. (Lochmaben) 79Castle Semple & Barr Lochs 47Cefni Estuary 127Cheshunt Gravel Pits 181Chew Valley Lake 194Chichester Harbour 240Christchurch Harbour 229Clandeboye Lake 97Cleddau Estuary 204Clwyd Estuary 119Coll 57 Colne Estuary 168Colwyn Bay 120Conwy Estuary 121Coquet Estuary 74Corby Loch 20Cotswold Water Park East 190Cotswold Water Park West 191Cowgill Reservoirs 60Cromarty Firth 9Crombie Reservoir 27Crouch/Roach Estuary 177Cuckmere Estuary 245Dart Estuary 221

Deben Estuary 164Dee (Eng/Wal) Estuary 118Dee (Scotland) Estuary 22Deeping St James GPs 146Dengie Flats 171Deveron Estuary 14Dinnet Lochs 24Don Estuary 21Dorchester Gravel Pits 188Dornoch Firth 7Draycote Water 139Drummond Pond 39Duddon Estuary 101Dulas Bay 124Dundrum Bay 94Dungeness Gravel Pits 248Dupplin Lochs 35Durham Coast 76Dyfi Estuary 134Dysynni Estuary 133Eden Estuary 31Ellesmere Group 135Erme Estuary 218Exe Estuary 223Eyebrook Reservoir 143Fairburn Ings 109Fal Complex 212Fala Flow 64Fedderate Reservoir 15Fen Drayton Gravel Pit 149Fiddlers Ferry Lagoons 113Fleet Bay 84Fleet/Wey 226Forth Estuary 62Foryd Bay 129Fowey Estuary 213Gadloch 42 Gannel Estuary 207Gladhouse Reservoir 63Grafham Water 147Guernsey Shore 209Gunton Park 155Haddo House Lakes 17Hamford Water 167Hanningfield Reservoir 178Hay-a-Park Gravel Pits 104Hayle Estuary 208Helford Estuary 211Hickling Broad 156Hirsel Lake 69Holborn Moss 72Hornsea Mere 107Hoselaw Loch 68Hule Moss 66Humber Estuary 108Hunterston Estuary 49Inland Sea 126Inner Clyde Estuary 46Inner Moray Firth 10Irt/Mite/Esk Estuary 100Irvine/Garnock Estuary 48Jersey Shore 210King George V Res 180Kingsbridge Estuary 220Kinnordy Loch 28Kirkcudbright Bay 83

Lackford Gravel Pits 152Lake of Menteith 44Langstone Harbour 239Larne Lough 99Lavan Sands 122Lindisfarne 71Little Paxton GPs 148Llandegfedd Reservoir 197Loch Clunie 36Loch Druidibeg 59Loch Eye 8Loch Fleet 6Loch Garten 25Loch Gilp 51Loch Gruinart 56Loch Indaal 55Loch Ken 82Loch Leven 34Loch Lomond: Endrick Mouth 45Loch Mahaick Doune 43Loch na Cille 52Loch of Boardhouse 2Loch of Harray 3Loch of Lintrathen 29Loch of Skene 23Loch of Spiggie 1Loch of Stenness 4Loch of Strathbeg 16Loch of the Lowes 37Loch Quien 50Loch Ryan 88Loch Spynie 11Loch Tullybelton 38Loch Watten 5Looe Estuary 214Lossie Estuary 12Lough Foyle 90Loughs Neagh & Beg 92Lower Derwent Valley 105Lower Windrush Valley GPs 189Luce Bay 86Machrihanish 54Martin Mere 116Mawddach Estuary 132Medina Estuary 235Medway Estuary 175Meikle Loch Slains 19Mersey Estuary 114

Yar Estuary 233

Mid Avon Valley 228Middle Tame Valley GPs 138Middle Yare Marshes 158Minsmere 162Montrose Basin 26Morecambe Bay 103Nene Washes 150Nevern Estuary 203Newhaven Estuary 244Newtown Estuary 234North Norfolk Coast 154North-West Solent 230Ogmore Estuary 198Orwell Estuary 165Otter Estuary 224

Ouse Washes 151Outer Ards Shoreline 96Pagham Harbour 241Pegwell Bay 173Pitsford Reservoir 142Plym Estuary 216Poole Harbour 227Portsmouth Harbour 238Pulborough/Amberley Brooks 242Queen Mary Reservoir 182Red Wharf Bay 123Rhunahaorine 53 Ribble Estuary 117River Tweed: Kelso toColdstream 67 Rostherne Mere 111Rough Firth 80Rutland Water 144Rye Harbour/Pett Levels 246Sevenoaks Wildfowl Reserve 179Severn Estuary 193Solway Estuary 78Somerset Levels 196Southampton Water 232Spey Estuary 13St Benets Levels 157Staines Reservoir 183Stour Estuary 166Strangford Lough 95Stratfield Saye 186Swale Estuary 174Swansea Bay 199Swithland Reservoir 141Tamar Complex 215Taw/Torridge Estuary 205Tay Estuary 30Tees Estuary 77Teifi Estuary 202Teign Estuary 222Thames Estuary 176Thanet Coast 172The Wash 153Theale Gravel Pits 187Thorpe Water Park 184Tiree 58 Tophill Low Reservoirs 106Traeth Bach 130Tweed Estuary 70Tyninghame Estuary 65Upper Lough Erne 91Walland Marsh 247Walmore Common 192Wath & Broomhill Ings 110West Water Reservoir 61Wigtown Bay 85Windermere 102 Woolston Eyes 112Wootton Estuary 236Wraysbury Gravel Pits 185

Yealm Estuary 217Ythan Estuary 18

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Figure A1. Location of important WeBS sites. Circles show the central position of 248 key WeBS sites, in the UK and theChannel Islands. Sites chosen include most internationally important sites, but also sites of regional importance in areas withfew wetlands or few sites counted by WeBS. Thus, inclusion of a site does not imply any measure of relative conservationimportance.

1

234

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Appendices 317