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TheHerpetologyof
ErieCounty,
Pennsylvania:A
BibliographyRevised2ndEdition
Brian S. Gray and Mark Lethaby
Special Publication of the Natural History Museum at the Tom Ridge Environmental
Center, Number 1
2
3
Special Publication of the Natural History Museum at the Tom Ridge
Environmental Center
The Herpetology of Erie County, Pennsylvania: A
Bibliography
Revised 2nd
Edition
Compiled by
Brian S. Gray
and
Mark Lethaby
Natural History Museum at the Tom Ridge Environmental Center, 301 Peninsula
Dr., Suite 3, Erie, PA 16505
Number 1 Erie, Pennsylvania 2017
Cover image: Smooth Greensnake, Opheodrys vernalis from Erie County, Pennsylvania.
4
Introduction
Since the first edition of The herpetology of Erie County, Pennsylvania: a
bibliography (Gray and Lethaby 2012), numerous articles and books have been published
that are pertinent to the literature of the region’s amphibians and reptiles. The purpose of
this revision is to provide a comprehensive and updated list of publications for use by
researchers interested in Erie County’s herpetofauna. We have made every effort to
include all major works on the herpetology of Erie County. Included are the works of
Atkinson (1901) and Surface (1906; 1908; 1913) which are among the earliest to note
amphibians and or reptiles specifically from sites in Erie County, Pennsylvania. The
earliest publication to utilize an Erie County specimen, however, may have been that of
LeSueur (1817) in his description of Graptemys geographica (Lindeman 2009). While
the bibliography is quite extensive, we did not attempt to list everything, such as articles
in local newspapers, and unpublished reports, although some of the more significant of
these are included. The authors welcome any literature citations that we may have
missed, so that we may consider their inclusion in future editions of this bibliography.
Milk Snake, Lampropeltis triangulum from West Springfield, Erie County.
5
Bibliography of Erie County Herpetology
Anderson, E. 1983. Turtle attracts statewide attention. Erie Morning News, Sept. 15.
Anderson, E. 1989a. Nature’s way: Amphibians and turtles. Erie Morning News.
April 13, 1989.
Anderson, E. 1989b. Nature’s way: The lucky frog. Erie Morning News. April 17, 1989.
Anderson, E. 2009. Nature's Way. Turtle focus of research. Erie Times-News Monday,
August 31, 2009. 4B.
Anderson, E. and J. Stull. 1995. Nature Watch: 1985-1995 Erie, Pennsylvania and nearby
areas. RNS Printing. Erie, PA 125 pp.
Aquatic Ecology Associates. 1977a. First interim data report for the period March 3,
1977 to May 20, 1977 for United States Steel Corporation. 600 Grant Street,
Pittsburg, Pennsylvania 15230, May 31, 1977.
Aquatic Ecology Associates. 1977b. Aquatic and terrestrial ecology studies, Lake Front
Plant, Conneaut, Ohio, summary report for the period May 21, 1977 to June 15,
1977. Ecology Associates, Pittsburgh, Pennsylvania.
Atkinson, D. A. 1901. The reptiles of Allegheny County, Pennsylvania. Annals of the
Carnegie Museum 1:145-157.
Bartlett, J. 1999. Reptile study leaves unsettled questions. Erie Times Morning News.
Monday, December 27, 1999, 4B.
Bartlett, J. 2000. Turtles, ducks might carry PCBs, mercury. Erie Times News. April 24,
2000. 1B and 8B.
Bartlett, J. 2001. Park puzzled by turtle deaths. Erie Times News vol.1 (193): 1A and 7A.
April 13, 2001.
Bartlett, J. 2002. Lake Erie mudpuppies dying. Erie Times News vol. 2 (272): 1A and
7A. July 1, 2002.
Bartlett, J. 2003. Park staff a little shell-shocked. Erie Times News 3 (247): 1A and 4A.
June 10, 2003.
Bates, S. P.1884. History of Erie County, Pennsylvania: Containing a history of the
county; its townships, towns, villages, schools, churches, industries, etc.; portraits
of early settlers and prominent men; biographies; history of Pennsylvania,
6
statistical and miscellaneous matter, etc., etc. Warner, Beers and Co., Chicago.
Black, D. 1998. ‘Hunters’ play shell game of turtle round-up. Erie Times News Sunday,
August 16, 1998.
Bogart, J. and M. W. Klemens. 2008. Additional distributional records of Ambystoma
laterale, A. jeffersonianum (Amphibia: Caudata) and their unisexual kleptogens in
Northeastern North America. American Museum Novitates, Number 3627. 58 pp.
Brewer, M.A. and J. L. Schnars. 2007. The response of neonate snapping turtle
(Chelydra serpentina serpentina) hatchlings to a moisture gradient. Regional
Science Consortium at the Tom Ridge Environmental Center at Presque Isle, 3rd
Annual Research Symposium (abstract).
Campbell, J.M., J.K. Bissell, H.N. Cunningham, Jr., E.C. Masteller, and J. Stull.
Biodiversity of Presque Isle. Pp. 405-422. In: Majumdar, S.K., F.J. Brenner, J.E.
Lovich, J.F. Schalles, and E.W. Miller (eds.). Biological Diversity: Problems and
Challenges. The Pennsylvania Academy of Sciences, Easton, Pennsylvania.
Carnegie Museum of Natural History: collection and distribution records. Provided by
Stephen Rogers, collection manager.
Collins, H. H. 1932. Observations on the Life History of the Vermilion-spotted Newt,
Triturus viridescens, in Western Pennsylvania Proc. Pa. Acad. Sci. 6:164-165.
Conant, R. 1975. Peterson field guide to reptiles and amphibians of eastern and central
North America. Houghton Mifflin, Boston.
Ernst C. 1985. Blanding’s Turtle. In: Genoways, H. H., and F. J. Brenner (eds). Species
of Special Concern in Pennsylvania. Special Publication of Carnegie Museum of
Natural History, No. 11. Pittsburgh, PA.
Ernst, C. H. and E. M. Ernst. 1971. The taxonomic status and zoogeography of the
Painted Turtle, Chrysemys picta, in Pennsylvania. Herpetologica 27(4):390-396.
Ernst, C. H. and J. E. Lovich. 2009. Turtles of the United States and Canada. Second
edition. Johns Hopkins University Press, Baltimore. 827 pp.
Fontanella, F. M., C. R. Feldman, M. E. Siddall, and F. T. Burbrink. 2008.
Phylogeography of Diadophis punctatus: extensive lineage diversity and repeated
patterns of historical demography in a transcontinental snake. Molecular
Phylogenetics and Evolution 46(2008):1049-1070.
Gray, B. 1999. Observations of amphibians and reptiles at Asbury Woods and along the
Greenway Trail: With a preliminary list of species. The Gray Press (privately
printed), Erie, PA. 18 pp. +10 plates.
7
Gray, B. 2000. Scaleless snake. Reptiles Magazine Vol.8(12):6
Gray, B. S. 2002a. Observations on the herpetofauna of the Asbury Woods Greenway in
Pennsylvania. Bull. Chicago Herp. Soc. 37(2):21-24.
Gray, B. S. 2002b. Natural History Notes: Thamnophis sirtalis sirtalis (Eastern Garter
Snake) Diet. Herpetol. Rev. 33(2):142-143.
Gray, B. S. 2002c. Natural History Notes: Storeria dekayi dekayi (Northern Brownsnake)
and S. occipitomaculata occipitomaculata (Northern Red-bellied Snake).
Morphology. Herpetol. Rev. 33(3):216.
Gray, B. S. 2002d. An addition to the herpetofauna of the Asbury Woods Greenway,
Erie, Pennsylvania. Bull. Chicago Herp. Soc. 37(11):196-197.
Gray, B. S. 2004. Report on a population of spotted turtles, Clemmys guttata in western
Erie County, Pennsylvania. Bull. Chicago Herp. Soc. 39(2):21-29.
Gray, B. S. 2005a. Checklist of the amphibians & reptiles of the Asbury Woods
Greenway. Pamphlet. Privately printed.
Gray, B. S. 2005b. Note on the distribution of the Short-headed Gartersnake (Thamnophis
brachystoma) in Erie County, Pennsylvania. Bull. Chicago Herp. Soc. 40(6):105-
106.
Spotted Salamander, Ambystoma maculatum from Asbury Woods, Erie County.
Immature Grey Treefrog, Hyla
versicolor from Asbury Woods, Erie
County.
Gray, B. S. 2006. The reptiles and
amphibians of the Asbury Woods
Greenway, Erie County, Pennsylvania.
Bull. Maryland Herp. Soc. 42 (2):115-
126.
Gray, B. S. 2007a. The herpetofauna of a
national superfund site in Erie,
Pennsylvania. Bull. Maryland Herp. Soc.
Vol. 43(3):129-133.
Gray, B. S. 2007b. Inventory of
amphibians along Wolf Run, Erie,
Pennsylvania. Bull. Chicago Herp. Soc.
Vol. 42(12):194.
Gray, B. S. 2008a. A note on site fidelity for ecdysis in the Northern Brown Snake,
Storeria dekayi dekayi. Bull. Chicago Herp. Soc. 43(3):
Gray, B. S. 2008b. Observations on the diet of the Shorthead Garter Snake, Thamnophis
brachystoma. J. Kansas Herpetol. No. 25 (March 2008):24-28.
Gray, B. S. 2008c. A study of carapace and plastron patterns in the spotted turtle,
Clemmys guttata, and their use as a technique for individual recognition. Bull.
Chicago Herp. Soc. 43(7):109-114.
Gray, B. S. 2009a. Arboreal ecdysis in the Eastern Garter Snake (Thamnophis sirtalis
sirtalis). J. Kansas Herpetol. 29:11.
Gray, B. S. 2009b. Recent observations of the herpetofauna of a former National
Superfund Site in Erie County, Pennsylvania. J. Kansas Herpetol. 31:9-11.
Gray, B. S. 2009c. Anomalous scutellation in an Eastern Garter Snake from Erie County,
Pennsylvania. Bull. Chicago Herp. Soc. 44(9):137-138.
Gray, B. S. 2009d. Estimating snout to vent length from data acquired from the shed
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skins of the Northern Brown Snake, Storeria dekayi dekayi. J. Kansas Herpetol.
32:17-19.
Gray, B. S. 2009e. Aberrant pattern in the Allegheny Dusky Salamander
(Desmognathus ochrophaeus) from Erie County, Pennsylvania. Bull. Chicago
Herp. Soc. 44(11):166.
Gray, B. S. 2010. Distribution of native and exotic earthworms in the eastern US:
implications for the ecology of vermivorous snakes. Bull. Chicago Herp. Soc.
45(5):73-86.
Gray, B. S. 2011a. Longevity and growth of a captive Shorthead Garter Snake,
Thamnophis brachystoma. J. Kansas Herpetol. 38:10-11.
Gray, B. S. 2011b. Seasonal activity and natural history observations of five snake
species from the Central Lowland Province of Erie County, Pennsylvania. J.
Kansas Herpetol. 39:14-21.
Gray, B. S. 2011c. Random shots (photograph of a Spring Peeper) in Erie Reader.
September 21 – October 4, 2011. Vol. 1(19):2.
Gray, B. S. 2011d. Random shots (photograph of a Blanding’s Turtle) in Erie Reader.
October 5 – October 18, 2011. Vol. 1(20):2.
Gray, B. S. 2011e. A Guide to the Reptiles of Erie County, Pennsylvania. Privately
Printed. Erie, PA. 88 pp.
Gray, B. S. 2011f. A Guide to the Amphibians of Erie County, Pennsylvania. Privately
Printed. Erie, PA. 76 Pp.
Gray, B. S. 2012. A note on the diet of the Common Garter Snake, Thamnophis sirtalis,
in Pennsylvania. Bull. Chicago Herp. Soc. 47(8):97-98.
Gray, B. S. 2013 a. Aggregations of Brownsnakes, Storeria dekayi (Holbrook, 1836), at a
site in northwestern Pennsylvania. Bull. Chicago Herp. Soc. 48(3):33-38.
Gray, B. S. 2013 b. Natural history notes: Storeria dekayi (Dekay’s Brownsnake) Diet.
Herpetol. Rev. 44(3):527.
Gray, B. S. 2013c. The diet of Dekay’s Brownsnake, Storeria dekayi at a site in Erie
County, Pennsylvania. Regional Science Consortium at the Tom Ridge
Environmental Center at Presque Isle, 9th
annual research symposium proceedings
(abstract of poster presentation).
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Common Five-lined Skink, Plestiodon fasciatus.
Gray, B. S. 2013 d. Observations on litters of Dekay’s Brownsnake, Storeria dekayi from
an urban population in northwest Pennsylvania. Bull. Maryland Herp. Soc.
49(1-4):30-39.
Gray, B. S. 2013 e. The relative importance of slugs and earthworms in the diet of
Dekay’s Brownsnake, Storeria dekayi, from northwestern Pennsylvania. Bull.
Chicago Herp. Soc. 48(12):157-165.
Gray, B. S. 2014 a. Natural history of Dekay’s Brownsnake, Storeria dekayi (Holbrook
1836) at a site in northwestern Pennsylvania. J. N. Amer. Herpetol. 2014(1):28-
39.
Gray, B. S. 2014 b. Natural history notes: Storeria dekayi (Dekay’s Brownsnake)
Cold-tolerance. Herpetol. Rev. 45(1):148.
Gray, B. S. 2014 c. Seasonal activity and temperature relationships of the eastern
gartersnake, Thamnophis sirtalis sirtalis (Linnaeus, 1758), from an urban
population in Erie County, Pennsylvania. Bull. Maryland Herp. Soc. 50(1-
2):22-33.
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Gray, B. S. 2014 d. Kyphoscoliosis in a Dekay’s Brownsnake, Storeria dekayi, from Erie
County, Pennsylvania, USA. Bull. Chicago Herp. Soc. 49(9):127-128.
Gray, B. S. 2014 e. An introduction to identifying prey remains in fecal samples of
snakes of the genus Storeria. Bull. Chicago Herp. Soc. 49(10):142-146.
Gray, B. S. 2014 f. The defensive behaviors of Dekay’s Brownsnake, Storeria dekayi:
An illustrated review of a diverse repertoire. Collinsorum 3(2-4):20-27.
Gray, B. S. 2015 a. Natural history notes: Storeria dekayi (Dekay’s Brownsnake) Diet.
Herpetol. Rev. 46(1):108.
Gray, B. S. 2015 b. A Study of the Defensive Behaviors of Free-ranging Dekay’s
Brownsnakes, Storeria dekayi (Holbrook, 1836). J. N. American Herpetol.
2015(1):43-52.
Gray, B. S. 2015 c. Natural History Observations of the Common Gartersnake,
Thamnophis sirtalis from Northwest Pennsylvania. Bull. Chicago Herp. Soc.
50(5):61-72.
Gray, B.S. 2015 d. The scavenging of snake carcasses by the Spotted Snake Millipede,
Blaniulus guttulatus. Collinsorum 4(2):3.
Gray, B. S. 2015 e. Natural history notes: Storeria dekayi (Dekay’s Brownsnake)
Population size, density and biomass. Herpetol. Rev. 46(4):650-651.
Gray, B. S. 2016. Associations between Dekay’s Brownsnake, Storeria dekayi, and
invertebrates. Bull. Chicago Herp. Soc. 51(2):25-28.
Gray, B. S. and P. Curtis. 2003. A note on sexual dichromatism in Clemmys guttata from
Erie County, Pennsylvania. Bull. Chicago Herp. Soc. 38(6):120-121.
Gray, B. S. and M. Lethaby. 2004. A den site utilized by the Northern Red-bellied Snake,
Storeria occipitomaculata occipitomaculata, in Pennsylvania. Bull. Maryland
Herp. Soc. 40(3):94-96.
Gray, B. S. and M. Lethaby. 2008. The amphibians and reptiles of Erie County,
Pennsylvania. Bull. Maryland Herp. Soc. 44(2):49-69.
Gray, B. S. and M. Lethaby. 2010. Observations of limb abnormalities in
amphibians from Erie County, Pennsylvania. J. Kansas Herpetol. 35:14-16.
Gray, B. S. and M. Lethaby. 2012. An addendum to The Amphibians and Reptiles of
Erie County, Pennsylvania. Bull. Maryland Herp. Soc. 48(1-4):11-19.
Gray, B. S., M. Lethaby, N. McAllister, and S. Bloomstine. 2013. Natural history notes.
12
Storeria occipitomaculata (Red-bellied Snake) Diet. Herpetol. Rev. 44(2):335.
Gray, B., H. M. Smith, and D. Chiszar. 2003. Further anomalies in the litters of a garter
snake from a hazardous waste site. Bull. Chicago Herp. Soc. 38(1):4-6.
Gray, B., H. M. Smith, J. Woodling, and D. Chiszar. 2001. Some bizarre effects on
snakes, supposedly from pollution, at a site in Pennsylvania. Bull. Chicago Herp.
Soc. 36(7):144-148.
Green, D. M. and C. Parent. 2003. Variable and asymmetric introgression in a hybrid
zone in the toads, Bufo americanus and Bufo fowleri. Copeia 2003(1):34-43.
Hahn, T. 2011. A jaw-dropper. Erie Times-News. Saturday, July 2, 2011.
Hudson, H.B. 1930. The distribution and habitat preference of the urodele amphibian
Triturus viridescens, at Presque Isle, Erie, Pennsylvania. Proc. Pa.
Acad. Sci. 4:55-58.
Hughes, J.R. and J.L. Schnars. 2007. Geographic Movements of the Common Snapping
Turtle (Chelydra serpentina serpentina) on Presque Isle State Park, Erie
Pennsylvania. Regional Science Consortium at the Tom Ridge Environmental
Center at Presque Isle, 3rd Annual Research Symposium (abstract).
Hughes, J.R. and J.L. Schnars. 2007. A Herpetological Inventory of Presque Isle State
Park, Erie, Pennsylvania Regional Science Consortium at the Tom Ridge
Environmental Center at Presque Isle, 3rd Annual Research Symposium
(abstract).
Hughes, J. R. and J. L. Schnars. 2008. Geographic movements of the common snapping
turtle (Chelydra s. serpentina) on Presque Isle State Park, Erie, Pennsylvania.
Proceedings of the Regional Science Consortium at the Tom Ridge
Environmental Center at Presque Isle, 4th
Annual Research Symposium
(Abstract).
Hulse, A. C. and K. L. Hulse. 1992. Geographic distribution. New county records for
amphibians and reptiles from Pennsylvania. Herpetol. Rev. 23(2):62-64.
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Pennsylvania and the Northeast. Ithaca and London. Cornell University Press.
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Regional Science Consortium at the Tom Ridge Environmental Center at Presque
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Lake Erie. Herpetologica 44:451-458.
13
Snapping Turtle, Chelydra serpentina from Millcreek Township, Erie County.
LeSueur, C. A. 1817. An account of an American species of tortoise, not noticed in the
systems. Journal of the Academy of Natural Sciences of Philadelphia 1:86-87.
Lethaby, M. 1990. Geographic distribution. Ambystoma jeffersonianum. Herpetol. Rev.
21(4):94.
Lethaby, M. 2001. Amphibians and reptiles on Presque Isle State Park: Summary of
current knowledge.Unpublished report to Presque Isle State Park.
Lethaby, M. 2004. Natural History Notes: Thamnophis brachystoma. Maximum size.
Herpetol. Rev. 35(1):73.
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42(1):115.
Lethaby, M. 2011b. Geographic distribution: Pantherophis spiloides. Herpetol. Rev.
42(1):115.
14
Lethaby, M. 2011c. Natural History Notes: Thamnophis brachystoma. Reproduction:
maximum litter size. Herpetol. Rev. 42(4):620.
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15
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Schnars, J. L. 2006. Using non-invasive techniques to determine mercury levels in
17
Snapping Turtle (Chelydra serpentina serpentina) tissues. Regional Science
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Swanson, P. L. 1948. Notes on the amphibians of Venango County, Pennsylvania.
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19
Northern Ribbonsnake, Thamnophis sauritus septentrionalis from Millcreek
Township, Erie County.
20
