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Sinan Canan, PhD [email protected] The Sense of Hearing

Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

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Page 1: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Sinan Canan, PhD [email protected]

The Sense of Hearing

Page 2: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Sound Waves

  Pressure waves   Compressions/expensions   Propagation in 3 dimensions

Page 3: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Sound Waves

  Pressure waves   Compressions/expensions   Propagation in 3 dimensions

Page 4: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Sound waves

  Frequency – “tone” [Hz]   Amplitude   Intensity   Timbre

Page 5: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Sound waves

  Sound intensity and “decibel” (dB)   Hearing treshold = 0 dB = 1.10-12 W/m2

  Bel = log [Pi/Pref]   dBel = 10 x log [Pi/Pref]

  10 dB = 1.10-11 W/m2 (treshold x 10)   20 dB = 1.10-10 W/m2 (treshold x 100)   30 dB = 1.10-9 W/m2 (treshold x 1,000)   40 dB = 1.10-8 W/m2 (treshold x 10,000)   50 dB = 1.10-7 W/m2 (treshold x 100,000)…

Page 6: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Sound waves

  Sound intensity and “decibel”

Source Intensity dB X Treshold Treshold 1.10-12 W/m2 0 100 Rustle of a leaf 1.10-11 W/m2 10 101 Whisper 1.10-10 W/m2 20 102 Regular talk 1.10-6 W/m2 60 106 Intense traffic 1.10-5 W/m2 70 107 Vacuum cleaner 1.10-4 W/m2 80 108 Large orchestra 6,3.10-3 W/m2 98 109.8 Max volume in an MP3 Player 1.10-2 W/m2 100 1010 Front row of a rock concert 1.10-1 W/m2 110 1011 Takeoff of a military jet 1.102 W/m2 140 1014 Eardrum damage 1.104 W/m2 160 1016

Page 7: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Sound waves   Hearing loss due to age

Page 8: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Ear

Outer ear Middle ear

Inner ear

Page 9: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Outer ear

  Congregating sound   Source localization   “Transfer function”

Page 10: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Outer Ear– “Transfer Function”

Page 11: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Eardrum

Page 12: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Eardrum

Normal Otitis media

Page 13: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Middle Ear

1.  Malleus (hammer) 2.  Incus (anvil) 3.  Stapes (stirrup) 4.  Eardrum 5.  Round window 6.  Eustachian tube

Page 14: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Middle Ear

1.  Malleus (hammer) 2.  Malleus ligament 3.  Incus (anvil) 4.  Incus ligament 5.  M. stapedius 6.  Stapes’ base 7.  Eardrum 8.  Eustachian tube 9.  M. Tensor tympani 10. Chorda tympani

Page 15: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Ossicular reflexes 1.  Malleus (hammer) 2.  Malleus ligament 3.  Incus (anvil) 4.  Incus ligament 5.  M. stapedius 6.  Stapes’ basis 7.  Eardrum 8.  Eustachian tube 9.  M. Tensor tympani 10. Chorda tympani

V

VII

Page 16: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Inner ear

Page 17: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Cochlea

Page 18: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Cochlea

Page 19: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Koklea

Scala vestibuli

Scala tympani

Scala media

Stria vascularis

Organ of Corti

Basilar membrane

Page 20: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Cochlea

Stria vascularis

Page 21: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Cochlea and the Organ of Corti

Page 22: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Organ of Corti

Page 23: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Basilar Membrane

Page 24: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Frequency analysis of sound Frequency selectivity of the basilar membrane

Page 25: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Frequency analysis of sound Frequency selectivity of the basilar membrane

Page 26: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Frequency analysis of sound Frequency selectivity of the basilar membrane

Page 27: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Frequency analysis of sound

Page 28: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Hair cells

  Inner hair cells   Outer hair cells

Page 29: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Organ of Corti

Page 30: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Hair cells

Tepe bağlantısı miyozin

Kanal

Page 31: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Hair cells

Destek hücreleri Deiter hücreleri

Dış tüy hücreleri İç tüy hücreleri

Destek hücreleri

Page 32: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Hair cells

Page 33: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Hair cells Outer hair cells

Inner hair cell

Page 34: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Hair cells

Outer hair cells: “Cochlear amplificator”

Page 35: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Hair cells

Transmitters of inner hair cells

Page 36: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Hair cells

Transmitters of the outer hair cells

Page 37: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Auditory pathways

Superior olivary

nucleus

Page 38: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Auditory pathways

  Superior olivary complex: First comparison of the sounds coming from each ear – source localization   Lateral superior olivary nucleus: Amplitude variations in

high pitches   Medial superior olivary nucleus: Time delays in lower

pitches

Page 39: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Auditory pathways

Electrocochleogram (ECoG)

Brainstem and cortical auditory

evoked potentials

Page 40: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Auditory pathways Electrocochleogram

Page 41: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Auditory cortex

Page 42: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

Auditory cortex

Page 43: Sense of Hearing - Ankara Yıldırım Beyazıt University Sense of Hearing . Sound Waves

The End...

Music is a higher revelation than all wisdom and philosophy… Ludwig van Beethoven