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Regional Summaries of Walleye Life History Characteristics Based on Ontario’s Fall Walleye Index Netting (FWIN) Program
1993 to 2001
Diagnostics and Sampling Standards Working Group Percid Community Synthesis
December 2002
G.E. Morgan and M.D. Malette Cooperative Freshwater Ecology Unit
Department of Biology, Laurentian University Sudbury, Ontario P3E 2C6
R.S. Kushneriuk Centre for Northern Forest Ecosystem Research
Ontario Ministry of Natural Resources 955 Oliver Road
Thunder Bay, Ontario P7B 5E1
S.E. Mann Northwest Region Science and Information
Regional Ageing Laboratory Ontario Ministry of Natural Resources 479 Government Road (Highway 17)
Dryden, Ontario P8N 2Z4
MNR 51729 (200 k P.R. 03 01 25) ISBN 0-7794-4081-1
TABLE OF CONTENTS
1.0 Introduction……………………………………………………………………. 1
2.0 The FWIN Database………………………………………………………….. 4
3.0 Parameter Estimation and Table Format…………………………………... 4 3.1 Basic Waterbody Descriptors……………………………………….. 4 3.2 Relative Abundance………………………………………………….. 5 3.3 Mortality………………………………………………………………... 5 3.4 Weight-Length Relationship and Condition………………………... 5 3.5 Length-at-Age…………………………………………………………. 6 3.6 Weight-at-Age…………………………………………………………. 6 3.7 Growth…………………………………………………………………. 6 3.8 Maturity………………………………………………….……………... 7 3.9 Fecundity………………………………………………………………. 8 3.10 Reproductive Characteristics………………………………………... 8 4.0 Analysis and Discussion……………………………………………………... 9 4.1 Sexual Dimorphism…………………………………………………… 9 4.2 Regional Differences…………………………………………………. 11 5.0 Summary………………………………………………………………………. 12 6.0 Acknowledgements…………………………………………………………… 12 7.0 References…………………………………………………………………….. 13
Appendix: Regional Benchmark Tables
Northwest Region (NW) Figure 2 – Map of Northwestern Ontario Table NW-1. Waterbody descriptors Table NW-2. Walleye relative abundance Table NW-3a. Female mortality Table NW-3b. Male mortality Table NW-4a. Female weight-length relationship Table NW-4b. Male weight-length relationship Table NW-5a. Female length-at-age Table NW-5b. Male length-at-age Table NW-6a. Female weight-at-age Table NW-6b. Male weight-at-age Table NW-7a. Female growth Table NW-7b. Male growth Table NW-8a. Female maturity Table NW-8b. Male maturity Table NW-9. Relative fecundity
Table NW-10. Fecundity-length relationship Table NW-11a. Female reproductive characteristics Table NW-11b. Male reproductive characteristics Table NW-12. Provincial benchmarks
Northeast Region (NE) Figure 3 – Map of Northeastern Ontario
Table NE-1. Waterbody descriptors Table NE-2. Walleye relative abundance Table NE-3a. Female mortality Table NE-3b. Male mortality Table NE-4a. Female weight-length relationship Table NE-4b. Male weight-length relationship Table NE-5a. Female length-at-age Table NE-5b. Male length-at-age Table NE-6a. Female weight-at-age Table NE-6b. Male weight-at-age Table NE-7a. Female growth Table NE-7b. Male growth Table NE-8a. Female maturity Table NE-8b. Male maturity Table NE-9. Relative fecundity Table NE-10. Fecundity-length relationship Table NE-11a. Female reproductive characteristics Table NE-11b. Male reproductive characteristics Table NE-12. Provincial benchmarks
South-central Region (SC) Figure 4 – Map of Southcentral Ontario
Table SC-1. Waterbody descriptors Table SC-2. Walleye relative abundance Table SC-3a. Female mortality Table SC-3b. Male mortality Table SC-4a. Female weight-length relationship Table SC-4b. Male weight-length relationship Table SC-5a. Female length-at-age Table SC-5b. Male length-at-age Table SC-6a. Female weight-at-age Table SC-6b. Male weight-at-age Table SC-7a. Female growth Table SC-7b. Male growth Table SC-8a. Female maturity Table SC-8b. Male maturity Table SC-9. Relative fecundity Table SC-10. Fecundity-length relationship Table SC-11a. Female reproductive characteristics Table SC-11b. Male reproductive characteristics Table SC-12. Provincial benchmarks
1
1.0 Introduction
This report presents regional summaries of life history characteristics of walleye (Stizostedion vitreum vitreum) populations from 390 Ontario lakes and large rivers. These waterbodies were sampled between 1993 and 2001 using the fall walleye index netting (FWIN) standard (Morgan 2002). The results for each region are presented as tables which describe the waterbody characteristics and the following walleye biological attributes: - relative abundance - annual adult mortality by sex - weight-length relationship and condition by sex - length-at-age by sex - weight-at-age by sex - growth by sex - sexual maturity by sex - relative fecundity - fecundity-length relationship - reproductive condition by sex
All data used in this report were derived from FWIN projects submitted to the Cooperative Freshwater Ecology Unit, Laurentian University and archived by the Ontario Ministry of Natural Resources (OMNR). These data were entered, archived, and managed using the OMNR FISHNET software package (Lester et al. 1989). Only data that were collected using the FWIN standard and managed by FISHNET are included in the regional summaries.
FWIN was first operationally tested in 1993 with pilot studies at Lac des Mille Lacs (OMNR northwest region) and the French River (OMNR south-central region). Since that time, the number of projects increased yearly (Table 1).
Table 1. FWIN projects completed in Ontario from 1993 to 2001.
Year Northwest Region
NortheastRegion1.
South-centralRegion1.
Ontario
1993 1 1 2
1994 3 2 1 6
1995 6 1 2 9
1996 9 6 6 21
1997 23 11 7 41
1998 34 26 10 70
1999 19 40 7 66
2000 64 37 11 112
2001 23 24 68 115
1. All northeast region waterbodies in Fishing Division 15 are included in the south-central region data summaries.
2
The large number of FWIN projects in the northwest region in 2000, the northeast region in 1999 and 2000, and the south-central region in 2001 were related to pilot “status of resource” studies and regional fisheries management initiatives. The provincial FWIN database contains 442 completed and archived FISHNET projects from 390 different water bodies. Almost three-quarters of the FWIN projects (and waterbodies) were in the northwest and northeast region (Table 2).
Table 2. FWIN projects and waterbodies sampled in Ontario from 1993 to 2001.
Region Number of FWIN Projects Number of Water bodies
Northwest 182 160
Northeast 147 131
South-central 113 99
Total 442 390
The life history characteristics and benchmarks summarized in this report are intended to be a source document for area biologists responsible for managing their walleye resource. These provide a means of comparing the results from individual populations of interest to other populations within the region or in other regions across the province. For any waterbody, where a standard FWIN survey has been conducted, a rapid, point-in-time assessment of the walleye population status is possible by comparing measured variables with known reference values.
In this report, we derive reference values (i.e., regional benchmarks) to encourage this type of assessment. Results of future sampling efforts can be compared against these walleye benchmarks for key life history parameters. We believe that these benchmarks would be useful to many walleye biologists because it permits them to look at their particular populations in the context of Ontario walleye populations generally. These benchmarks represent the average values along with some measures of variability from a large set of Ontario waterbodies, widely distributed across the province (Figure 1). The regional benchmarks are organized in a series of 11 tables. The provincial benchmarks are summarized in one table (Table 12 which is included at the end of each regions benchmark tables). Data from a project was excluded from a table if any of the required data fields were missing or estimators could not be calculated. (i.e., the number of fish sampled was too small). At the end of each table are summary statistics for comparative purposes (i.e., the regional benchmarks).
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2.0 The FWIN Database
The master FWIN database was created through a unique combination of FISHNET files and compendium output (Kushneriuk 1996). Data from individual FISHNET files were stored in ACCESS® while the compendium output files were compiled by project and stored in ORACLE®. Automated data transformations were performed in ORACLE® to create the final working tables (e.g., the re-ordering of male, female, and combined sex data). All ACCESS® and ORACLE®
tables were combined into a new master FWIN database where querying of the data occurred to create the regional benchmark tables.
Each life history character was estimated using a consistent set of defendable methods. Many of these life history parameters are related to fish age so every effort has been made to ensure consistent tissue collection and age interpretation. Calcified structures collected from the FWIN projects over the years varied across the province. Whenever possible the sagitta or otolith was the first choice for walleye age interpretation. The method used to prepare the otoliths was the crack and burn technique described by Christensen (1964). Annuli were very distinct for age interpretation, especially for very old fish compared to the dorsal spine or scale methods. If no otoliths were collected, the next preferred structure for age interpretation was the dorsal spine. The spine was coated with a two-part epoxy and then sectioned along the transverse plane at the base of the structure. Age interpretation was difficult for very old walleye due to piling of annuli along the edge of the structure. Scales were only used for very young fish (during their rapid growth period and before sexual maturation). Scales were impressed into acetate slides and interpreted in a microfiche reader. The Northwest Science and Information Ageing Lab in Dryden, Ontario prepared and aged over 95% of the total projects in the FWIN database.
3.0 Parameter Estimation and Table Format
3.1 Basic Waterbody Descriptors (Table 1)
Various attributes of the waterbodies were listed alphabetically. Latitude, longitude, surface area, mean depth, maximum depth, and Secchi depth were derived from the Aquatic Habitat Inventory Database (contained in FISHNET) or updated from information provided by OMNR area biologists. Climate was characterized by growing-degree-days above 5oC which was estimated for each individual waterbody from the Ontario climate model (Mackey et al. 1996). The regional benchmarks did not include projects where only a portion of a waterbody was sampled. A map of the waterbodies sampled in each region from 1993 to 2001 is provided (Figures 2 to 4).
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3.2 Relative Abundance (Table 2)
The average catch or relative abundance of walleye caught, standard error, sample size, number of nets that caught walleye, and the average size of the walleye caught (expressed as total length in mm and wet weight in grams) from each project were recorded in this table and arranged by waterbody name in alphabetical order. The relative abundance reported was the geometric mean (i.e., the average of the logarithmically transformed (base 10) individual observations increased by one). For any waterbody that had been sampled in more than one year, data from the most recent year was used as input for deriving regional benchmarks.
3.3 Mortality (Table 3)
Observed female (Table 3a) and male (Table 3b) annual adult mortality was described by the Robson and Chapman estimator for fish age 5 and older (Robson and Chapman 1961, Ricker 1975 p. 31):
1-+=
TN
TS
where S is annual adult survival, T = N1 + 2N2 + 3N3…, N = N0 + N1 + N2 +…, and N is the number of fish whose age is greater than or equal to age 5.
Annual adult mortality (A), the proportion of fish that dies in one year, was then calculated as A=1-S and expressed as a percent. A regression analysis indicated that the combined sex estimate was a good predictor of female and male mortality and we used these results to generate additional female and male estimates. The tables were arranged from low to high values using the estimated total annual mortality. The regional benchmarks were derived using all the data (including multiple years from an individual waterbody).
3.4 Weight-Length Relationship and Condition (Table 4)
These tables give estimates of weight-length parameters for females (Table 4a) and males (Table 4b) based on a geometric mean predictive regression model fitted to logarithmically transformed (base 10) individual observations of wet weight (grams) versus total length (mm).
LW10
log+10
log=10
log
where W is wet weight in grams, is the intercept, is the slope, and L is total length in mm. The equation presented in the table is W=10 L .
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Only projects with 30 or more observations, a correlation coefficient greater than or equal to 0.90 (i.e., R2 0.90), and having their derived regression models with slopes and intercepts significantly different from 0 (p<0.05) were included in these tables. Values were sorted from low to high using the slope parameter ( ). The regional benchmarks were derived using all the data (including multiple years from an individual waterbody).
Condition was calculated as the arithmetic average of the individual ratios of observed wet weight-to-predicted wet weight. Observed wet weight was the actual weight recorded in the FISHNET file. Predicted wet weight was derived from each regions median regression model (i.e., the regional weight-length benchmark relationship) by estimating the wet weight of a walleye based on the observed total length in the FISHNET file.
3.5 Length-at-Age (Table 5)
Total length-at-age data are presented for females (Table 5a) and males (Table 5b). Only arithmetic mean values with a minimum sample size of 4 individual observations or more were included in these tables. Values were sorted from low to high using total length-at-age 1, 2 and 3. The regional benchmarks were derived using all the data (including multiple years from an individual waterbody).
3.6 Weight-at-Age (Table 6)
Wet weight-at-age data are presented for females (Table 6a) and males (Table 6b). Only arithmetic mean values with a minimum sample size of 4 individual observations or more were included in these tables. Values were sorted from low to high using total weight-at-age 1, 2 and 3. The regional benchmarks were derived using all the data (including multiple years from an individual waterbody).
3.7 Growth (Table 7)
The growth parameter tables (Table 7a for females and Table 7b for males) reports results from a parameter constrained growth model based on an empirical method of fitting the Von Bertalanffy growth equation:
)1(=)( ottK
t eLL
where Lt is total length in mm at age t, L is asymptotic total length in mm, K is the Brody growth coefficient (i.e., relative growth rate), and to adjusts the time
7
scale to incorporate a non-zero length at age=0. The value of to was constrained as -1.
Kushneriuk et al. (1996) expanded on the ideas of Pauly (1984) and Payne et al.
(1990) to estimate asymptotic length using the geometric mean length (_
L ) of the largest p% of the fish in a sample, where p is a variable that depends on sample size. Asymptotic length was estimated using:
)0006.0-0223.00732.1( 2_
ppLL
where p is the sampling fraction expressed as a percent. A 1% sample was used wherever the total number of fish aged by sex was greater than 500. For smaller sample sizes, the largest 5 fish and the above formula was used to estimate L .
Omega ( ) is early growth and corresponds to the growth rate near to (and is calculated as K•L ) (Gallucci and Quinn 1979). Pre-maturation growth (h) was defined as the growth rate to 350mm total length for combined sexes (Lester et al. 2000).
1+
350=
350T
mmh
where T350 was interpolated from mean total length in mm of the two consecutive age groups whose lengths bounded 350mm.
Values were sorted from low to high using L . The regional benchmarks were derived using all the data (including multiple years from an individual waterbody).
3.8 Maturity (Table 8)
Maturation was visually assessed by internal examination of the gonads from individually sampled fish (Duffy et al. 2000). Logarithmically transformed (base 10) age and size-at-maturity schedules (i.e., the percent mature at age or length) were fitted to a logistic regression function for each population. Age and size at 10, 50, and 90% maturity were then derived from the equation. The data was arranged from youngest to oldest age-at-50% maturity. Results are presented for females in Table 8a and males in Table 8b. The regional benchmarks were derived using all the data (including multiple years from an individual waterbody).
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3.9 Fecundity (Table 9 and 10)
Mature ovaries were collected in the field and preserved for later counting. Fecundities were estimated gravimetrically. Ovary wet weight (in grams) was determined for the preserved samples, then eggs were counted in three sub-samples, each of approximately 0.30 grams, taken from anterior, medial, and posterior sections of the ovary. The arithmetic mean number of eggs per gram thus obtained was multiplied by the preserved ovary wet weight to determine fecundity. Relative fecundity (RF) was calculated as:
W
eggsofNumberRF =
where W is wet weight in grams.
This data was arranged from lowest to highest in Table 9. Only arithmetic mean values with a minimum sample size of 4 individual observations or more were included.
Table 10 gives estimates of fecundity-total length parameters based on a geometric mean predictive regression model fitted to logarithmically transformed (Napierian logarithms) individual observations of number of eggs versus total length (mm).
Lfecundity ln+ln=ln
where fecundity is the number of eggs, is the intercept, is the slope, and L is total length in mm. The equation presented in the tables is Fecundity=e L .
Only projects having their derived regression models with slopes and intercepts significantly different from 0 (p<0.05) were included in this table. Values were sorted from low to high using the slope parameter ( ). The regional benchmarks were derived using all the data (including multiple years from an individual waterbody).
3.10 Reproductive Characteristics (Table 11)
Other measures of reproductive condition are summarized in Table 11a for females and Table 11b for males. Gonads were removed from the walleye and weighed with an electronic balance that was accurate to at least 1 decimal place. The fall gonadosomatic index (GSI) of sexually mature fish was calculated as:
100×=Weight)Gonad-tWeigh(Total
WeightGonadGSI
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where gonad weight and total weight are in grams.
Fat was gently stripped by hand from the membranes surrounding the digestive tract, organs and body cavity. Visceral fat was weighed with an electronic balance that was accurate to 2 decimal places. The visceral fat index (VFI) of sexually mature fish was calculated as:
100×=WeightTotal
WeightFatVisceralVFI
where visceral fat weight and total weight are in grams.
The index of reproductive effort (R) proposed by Lester et al. (2000) was also calculated for females and males:
L
hR
3=
where h is pre-maturation growth (mm•year-1) and L is asymptotic total length in mm.
Only arithmetic mean values with a minimum sample size of 3 individual observations or more were included in these tables. The tables are ordered from lowest GSI to highest GSI. The regional benchmarks were derived using all the data (including multiple years from an individual waterbody).
4.0 Analysis and Discussion
4.1 Sexual Dimorphism
The analysis of the 1993 to 2001 FWIN database is unique because most comparable studies of life history variation have been interspecific, rather than intraspecific comparisons of population. In this study we have tried to improve our understanding of the variation in walleye life history by increasing the number of populations and life history parameters compared to Lester et al. (2000).
Female and male life history parameters were tested for the null hypothesis that these data were from a normally distributed population using the Kolmogorov-Smirnov one-sample test (Daniel 1978). Mean population differences between females and males from normally distributed data were compared using the paired-sample t test for dependent samples (Zar 1999). For data that were not normally distributed, the nonparametric Wilcoxon paired-sample test (Daniel
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1978, Zar 1999) was employed to test for differences between females and males. The mean difference and the 95% confidence interval for statistically significant (p<0.05) comparisons between the sexes were calculated. The linear dependence of variation in a life history parameter between females and males was tested using simple linear correlation (i.e., we considered the linear relationship between females and males, but neither is assumed to be functionally dependent upon the other) (Zar 1999). Statistical comparisons were performed using STATISTICA data analysis software system (StatSoft 2001).
The mean differences between female and male walleye were statistically significant (p<0.01) for all life history parameters except relative condition (p=0.80) (Table 3). Female walleye had lower mortality, higher asymptotic size, faster early growth, older age at sexual maturity, larger size at sexual maturity, higher allocation to sexual by-products (i.e., higher gonadosomatic index), and higher storage of lipid reserves (i.e., higher visceral fat index) compared to males from the same population. Variation between females and males was statistically correlated for all parameters (p<0.01).
Table 3. Comparison of female and male walleye life history parameters.
Life History Parameter (Sample Size)
Females versus Males Are they correlated? (Mean difference, ±95%C.I.1.)
Adult mortality (A%) (n=269)
Females=27.9%*Males=30.1%
Yes (r=0.52, p<0.01), (2.3%, 0.8 to 3.8%)
Relative condition (n=225)
Females=1.03Males=1.03
Yes (r=0.81, p<0.01)
Asymptotic length (L )(n=211)
Females=742mm*Males=615mm
Yes (r=0.57, p<0.01), (127mm, 119 to 137mm)
Brody coefficient (K) (n=211)
Females=0.155yr-1*Males=0.179yr-1
Yes (r=0.55, p<0.01), (0.024yr-1, 0.020 to 0.029yr-1)
Omega ( )(n=211)
Females=113.88mm•yr-1*Males=109.33mm•yr-1
Yes (r=0.77, p<0.01), (4.55mm•yr-1, 2.60 to 6.45mm•yr-1)
Age-at-50% maturity (n=217)
Females=4.6 years*Males=3.0 years
Yes (r=0.79, p<0.01) (1.6 years, 1.5 to 1.7 years)
Size-at-50% maturity (n=217)
Females=442mm*Males=348mm
Yes (r=0.48, p<0.01), (94mm, 88 to 98mm)
Gonadosomatic index (n=147)
Females=4.4%*Males=2.9%
Yes (r=0.23, p<0.01), (1.5%, 1.2 to 1.7%)
Visceral fat index (n=147)
Females=1.8%*Males=1.4%
Yes (r=0.84, p<0.01), (0.5%, 0.4 to 0.6%)
1. 95% Confidence interval of mean difference. * Significant difference (p<0.01) between females and males.
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4.2 Regional Differences
Life history parameters and population characteristics (i.e., geometric mean walleye relative abundance) for female and male walleye from the northwest, northeast and south-central region FWIN projects were compared using a one-way analysis of variance (Zar 1999). If a significant (p<0.05) difference in means was detected a multiple comparison Tukey test was employed to determine where differences between regions existed (Zar 1999). Statistical comparisons were performed using STATISTICA data analysis software system (StatSoft 2001). Results are summarized in Table 4.
Table 4. Comparison of regional walleye life history parameters.
Regional Average1.
Life History Parameter Northwest Northeast
South-central
OntarioAverage
Female adult mortality (A%) 28 (a) 29 (a) 29 (a) 282.
Male adult mortality (A%) 30 (a) 31 (a) 31 (a) 30
Female relative condition 1.04 (a) 1.04 (a) 1.02 (a) 1.03
Male relative condition 1.06 (b) 0.99 (a) 1.03 (b) 1.03
Female asymptotic length (L , mm) 733 (a) 748 (ab) 778 (b) 744
Male asymptotic length (L , mm) 613 (a) 612 (a) 646 (b) 618
Female Brody coefficient (K , yr-1) 0.155 (a) 0.159 (a) 0.172 (a) 0.158
Male Brody coefficient (K, yr-1) 0.176 (a) 0.182 (a) 0.188 (a) 0.180
Female Omega ( , mm•yr-1) 112.14 (a) 116.99 (a) 131.84 (b) 116.40
Male Omega ( , mm•yr-1) 106.82 (a) 110.67 (a) 120.25 (b) 110.57
Pre-maturation growth (h, mm•yr-1) 89.37 (a) 89.78 (a) 109.54 (b) 93.00
Female age-at-50% maturity (years) 4.8 (b) 4.5 (b) 3.6 (a) 4.5
Male age-at-50% maturity (years) 3.5 (c) 2.9 (b) 2.2 (a) 3.03.
Female size-at-50% maturity (mm) 441 (a) 440 (a) 445 (a) 441
Male size-at-50% maturity (mm) 360 (b) 340 (a) 342 (a) 349
Female gonadosomatic index (%) 3.1 (a) 4.1 (a) 4.8 (b) 4.4
Male gonadosomatic index (%) 3.1 (ab) 2.6 (a) 3.2 (b) 2.9
Relative fecundity (eggs•g-1) 41.33 (a) 49.74 (b) 54.36 (b) 49.79
Female visceral fat index (%) 3.1 (b) 2.1 (ab) 1.6 (a) 1.9
Male visceral fat index (%) 2.1 (a) 1.4 (a) 1.3 (a) 1.4
Relative abundance (number•net-1) 10.7 (c) 6.4 (b) 2.8 (a) 7.5
1. Similar letters represent the similarity (p>0.05) of the population means that they estimate. Different letters represent populations that have a mean different (p<0.05) from any other population. Means are ranked from low to high (i.e., from a to c).
2. Bold font style indicates no significant difference (p>0.05) in means between the three regions. 3. Underlined font style indicates significant difference (p<0.05) between each regions mean.
There were no significant differences between the regional means for female and male adult mortality, female relative condition, female and male Brody growth coefficients, female size-at-50% maturity, and male visceral fat index. The regional means for male age-at-50% maturity and relative abundance (geometric
12
mean) were significantly different from each other. For all other life history parameters, except male size-at-50% maturity and male relative condition, there generally appeared to be no difference between northwest and northeast region means. However the south-central region means were significantly different from the other two regions. The mean male size-at-50% maturity for northwest region walleye populations was significantly different from the other two regions while the mean male relative condition for northeast region walleye populations was significantly different from the other two regions.
5.0 Summary
This report demonstrates the utility of gathering sufficient, standard samples to estimate walleye life history parameters. Females differ significantly from males and future assessment should recognize that data should be collected and analyzed separately for the sexes. Walleye populations in the northwest and northeast region displayed similar life history parameter means and information from these northern populations may be pooled for future analysis. South-central population means appeared to be different and should be treated separately. The regions differed uniquely in only three parameter means: relative abundance, male relative condition, and male age-at-50% maturity.
6.0 Acknowledgements
Heartfelt thanks goes to the numerous biologists, field technicians, and volunteers whose contributions have made the FWIN program such a success. Growing-degree-day estimates were kindly provided by Terry Marshall. Constructive comments from Chuck Brady, John Gunn, Bryan Henderson, Peter Hulsman, Steve Kerr, Rob Korver, Nigel Lester, and Darryl McLeod improved the manuscript greatly. We also gratefully acknowledge Tracy Wakefield for formatting the final document.
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7.0 References
Christensen, J.M. 1964. Burning of otoliths, a technique for age determination of Soles and other fish. Journal du Conseil Permanent International pour l’Exploration del la Mer 29(1):73-81.
Daniel, W.W. 1978. Applied nonparametric statistics. Houghton Mifflin Company, Boston, USA. 503 p.
Duffy, M.J., J.L. McNulty, and T.E. Mosindy. 2000. Identification of sex, maturity, and gonad condition of walleye (Stizostedion vitreum vitreum). Northwest Science and Technology Field Guide: FG-05. Ontario Ministry of Natural Resources, Northwest Region Science and Technology Unit, Thunder Bay, Ontario. 33p.
Gallucci, V.F. and T.J. Quinn. 1979. Reparameterizing, fitting, and testing a simple growth model. Transactions of the American Fisheries Society 108:14-25.
Kushneriuk, R.S. 1996. FISHNET LITE (computer analysis software system). Ontario Ministry of Natural Resources, Peterborough, Ontario.
Kushneriuk, R.S., N.P. Lester, and R.M. Korver. 1996. A compendium of life history characteristics of walleye in Ontario waters. Percid Community Synthesis. Ontario Ministry of Natural Resources. 45 p + lake summaries.
Lester, N.P., R.S. Kushneriuk, S. Orsatti, and D.G. Oliver. 1989. FISHNET user manual. Ontario Ministry of Natural Resources. Toronto.
Lester, N.P., B.J. Shuter, R.S. Kushneriuk, and T.R. Marshall. 2000. Life history variation in Ontario walleye populations: implications for safe rates of fishing. Percid Community Synthesis. Population and Yield Characteristics Working Group. Ontario Ministry of Natural Resources. 34 p.
Mackey, B.D., D. McKenney, Y-Q. Yang, J. McMahon, and M. Hutchinson. 1996. Site regions revisited: a climatic analysis of Hills’ site regions for the province of Ontario using a parametric method. Canadian Journal of Forest Research 26:333-354.
Morgan. G.E. 2002. Manual of instructions – fall walleye index netting (FWIN). Percid Community Synthesis, Diagnostics and Sampling Standards Working Group. Ontario Ministry of Natural Resources. 34 p.
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Payne, N.R., R.M. Korver, D.S. MacLennan, S.J. Nepszy, B.J. Shuter, T.J. Stewart, and E.R. Thomas. 1990. The harvest potential and dynamics of lake trout populations in Ontario. Lake Trout Synthesis. Ontario Ministry of Natural Resources. 72 p.
Pauly, D. 1984. Fish population dynamics in tropical waters: a manual for use with programmable calculators. International Center for Living Resources Management. Manila, Philippines. 325 p.
Ricker. W.E. 1975. Computation and interpretation of biological statistics of fish populations. Bulletin 191. Department of the Environment, Fisheries and Marine Service, Ottawa. 382 p.
Robson, D.S. and D.G. Chapman. 1961. Catch curves and mortality rates. Transactions of the American Fisheries Society 90:181-189.
StatSoft, Inc. (2001). STATISTICA (data analysis software system), version 6. www.statsoft.com.
Zar, J.H. 1999. Biostatistical analysis, fourth edition. Prentice Hall, New Jersey, USA. 663 p. + appendices.
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sta
nda
rd 1
99
3 t
o 2
00
1.
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Ab
ram
2
00
1
50
03
26
-91
55
54
2
04
0.5
1
2.1
3
6.6
2
.0
15
25
Ag
ima
k
19
95
4
92
31
3-9
14
04
7
12
22
.1
3.2
1
5.8
2
.7
14
87
Alp
h
19
97
5
01
91
3-8
63
83
9
46
9.9
3
.4
14
.3
3.1
1
18
8
An
de
rs
19
96
4
85
85
4-8
85
44
6
16
4.5
7
.5
66
.0
3.2
1
21
0
Ara
1
99
8
50
32
55
-87
27
24
4
92
6.8
4
.6
18
.0
5.2
1
16
5
Ba
ll 1
99
7 &
19
98
5
01
72
6-9
35
93
1
29
15
.4
9.5
3
0.5
1
.4
15
51
Ba
r 2
00
0
49
03
00
-91
12
00
1
32
.3
7.5
2
2.0
3
.0
13
76
Ba
rr
20
00
4
90
34
1-9
15
74
4
10
1.0
5
.8
15
.9
3.9
1
39
5
Be
ak
20
00
4
91
65
3-9
21
51
7
52
4.3
7
.7
30
.0
3.8
1
43
1
Be
ave
rho
use
1
99
6
48
32
36
-92
06
09
2
00
0.8
2
3.6
6
3.4
3
.4
15
51
Be
nd
ing
1
99
7
49
19
13
-92
08
15
1
13
5.7
1
2.5
4
5.8
4
.6
15
64
Bo
ffin
2
00
0
48
45
03
-93
28
10
5
44
.7
3.4
1
2.8
4
.8
15
00
Bo
tsfo
rd
20
01
5
00
84
2-9
13
83
2
14
47
.2
7.2
2
4.7
2
.6
15
00
Bra
dsh
aw
2
00
0
48
51
00
-91
25
00
1
44
.5
2.5
8
.0
5.4
1
38
3
Bre
nn
an
2
00
0
50
29
27
-89
54
25
2
57
5.3
1
.8
20
.7
4.0
1
25
1
Bre
we
r 1
99
9
47
17
26
-91
57
20
1
02
.0
7.1
1
5.9
1
.7
15
54
Bro
ad
1
99
8
50
26
31
-92
37
47
6
15
.1
2.6
5
.8
1.4
1
50
1
Bu
rditt
19
98
4
85
70
4-9
34
64
2
14
20
.5
9.7
2
3.5
5
.9
16
41
Bu
rro
ws
20
00
4
95
70
8-8
64
41
8
17
61
.3
2.9
1
8.3
2
.3
11
80
Ca
lm
19
98
4
84
62
7-9
20
45
0
24
80
.9
13
.9
65
.3
2.3
1
53
7
Ch
ill
20
00
4
90
80
0-9
10
10
0
16
4.0
6
.1
34
.0
3.0
1
37
5
Ch
urc
hill
1
99
8
50
47
39
-91
10
32
4
17
8.8
5
.2
22
.6
3.4
1
36
7
Clo
ud
1
99
9
48
08
07
-89
32
34
4
20
.9
9.0
1
6.5
3
.7
13
06
Co
mp
an
ion
3.
20
00
4
85
40
0-9
11
40
0
51
5.6
5
.1
27
.5
2.4
1
38
0
Cro
oke
d P
ine
1
99
8
48
47
04
-91
05
36
1
60
4.0
6
.1
17
.4
4.4
1
40
7
Cu
ttle
1
99
8
48
52
49
-93
07
39
5
62
.5
5.1
1
6.8
3
.1
16
39
Da
rby
20
00
4
83
63
5-9
20
84
1
85
.8
4.1
1
1.6
2
.2
15
00
NW
- 2
Ta
ble
NW
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
de
s M
ille
Lacs
19
93
, 1
99
4
& 1
99
9
48
50
57
-90
30
37
2
41
01
.0
6.8
2
4.4
1
.8
13
22
De
sp
air
2
00
0
48
52
25
-93
39
29
9
11
.4
6.1
9
.2
2.8
1
50
0
De
vio
us
19
99
4
90
90
6-8
74
72
8
13
3.5
2
.3
12
.5
2.7
1
09
7
Dim
ple
2
00
0
49
14
05
-91
55
59
3
73
.5
12
.3
25
.0
4.7
1
38
7
Do
g
19
97
4
84
55
0-8
93
23
3
14
80
4.3
2
9.6
1
17
.0
2.5
1
23
2
Do
re
19
98
4
83
60
0-9
12
70
0
No
da
ta
No
da
ta
No
da
ta
No
da
ta
14
46
Do
ve
tail
19
96
4
85
24
8-9
20
22
4
79
4.8
2
.3
27
.0
2.0
1
52
9
Dru
m
20
00
4
91
73
1-9
20
95
9
10
3.1
4
.8
17
.0
3.3
1
42
4
Ea
gle
2
00
0
49
40
53
-93
04
10
2
76
91
.0
6.8
3
3.5
2
.9
15
79
Elb
ow
1
99
5
48
44
43
-91
00
51
5
83
.6
22
.5
58
.6
5.6
1
39
3
Eltru
t 2
00
0
49
00
20
-92
24
48
2
40
1.4
3
.7
12
.2
1.7
1
59
1
Elv
a
19
98
4
95
23
6-9
10
95
3
16
5.1
1
.4
5.2
2
.7
14
69
Exp
an
se
1
99
8
50
20
27
-91
47
00
8
52
.1
3.9
1
3.1
3
.1
14
76
Fa
cto
r 2
00
0
48
42
02
-92
03
53
6
29
.7
11
.9
49
.4
5.9
1
46
3
Fa
vo
ura
ble
1
99
5
52
50
57
-93
54
55
7
11
7.6
4
.0
13
.0
2.1
1
27
0
Fin
layso
n
19
97
4
85
43
3-9
13
52
2
14
55
.3
16
.3
51
.9
2.3
1
46
1
Fo
otp
rin
t 2
00
0
48
54
00
-93
36
30
4
97
.0
7.4
1
7.4
3
.4
15
00
Ga
mm
on
2
00
0
51
00
31
-94
44
03
1
21
8.5
7
.3
21
.0
3.5
1
47
6
Ga
msb
y
20
01
4
93
10
4-8
71
05
4
60
1.8
1
.7
7.6
2
.3
11
16
Ga
rde
n
19
97
4
93
12
5-8
94
93
4
18
03
.8
7.0
2
2.3
1
.7
12
44
Ge
ikie
2
00
0
49
30
00
-89
19
00
N
o d
ata
N
o d
ata
N
o d
ata
N
o d
ata
1
23
8
Ge
org
ia
19
99
4
91
93
4-8
75
31
0
58
0.8
5
.2
13
.7
2.5
1
11
1
Ha
rris
2
00
1
50
16
46
-90
38
38
4
19
.4
6.4
2
8.0
4
.0
13
59
He
nd
ers
on
1
99
7
48
49
00
-90
18
00
1
50
.9
2.5
6
.5
2.5
1
28
4
Hic
ks
19
99
4
84
85
0-8
90
52
4
19
8.3
N
o d
ata
6
.4
0.8
1
18
4
Hu
sb
an
d
20
00
4
85
11
1-9
12
63
5
15
5.9
5
.1
11
.6
2.9
1
38
3
Hu
tch
iso
n
20
00
4
94
70
9-8
65
70
9
21
5.3
6
.1
23
.8
2.6
1
14
9
NW
- 3
Ta
ble
NW
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Ica
rus
20
00
4
81
31
8-9
03
32
9
72
5.2
1
1.6
4
2.7
3
.4
13
37
Ice
1
99
7
48
51
00
-90
15
00
1
12
.0
1.6
3
.0
1.0
1
26
6
Iro
n R
an
ge
1
99
8
48
11
24
-90
19
48
2
33
.1
2.5
6
.1
3.6
1
28
7
Ja
c S
ag
a
20
00
4
91
31
4-9
15
71
5
19
.5
5.0
1
5.0
3
.0
14
03
Ja
ckfish
2
00
0
48
56
26
-93
35
46
1
88
9.9
6
.0
21
.4
3.6
1
50
0
Jo
ne
s
20
00
4
90
60
4-9
22
63
5
38
9.1
5
.9
17
.9
1.5
1
45
6
Ju
tte
n
20
01
5
01
95
9-9
03
24
5
66
8.9
1
7.6
2
5.0
4
.1
13
46
Ka
op
skik
am
ak
20
00
4
90
70
8-9
25
02
9
12
66
.7
7.2
3
9.0
4
.0
15
00
Ka
pkic
hi
19
98
5
12
81
7-9
02
25
7
12
77
.2
3.7
1
3.1
2
.9
12
33
Ka
wa
wia
g
20
00
4
91
25
6-9
25
12
4
46
8.2
8
.4
35
.1
3.4
1
49
9
Ke
cku
sh
2
00
0
48
56
44
-91
14
15
1
36
.8
5.2
1
3.7
2
.5
13
78
Ke
ike
wa
bik
1
99
7
49
49
35
-92
10
05
8
79
.4
3.6
7
.6
1.9
1
55
4
Ke
no
ga
mis
is
19
98
4
94
12
8-8
65
24
5
41
65
.2
1.9
9
.2
2.0
1
16
0
Ke
no
rain
2
00
0
49
01
41
-92
19
35
5
45
.5
6.1
1
3.7
3
.2
14
47
Ke
no
zh
e
20
00
4
91
04
7-9
25
10
7
20
6.4
2
.9
5.2
2
.7
14
99
Kilb
urn
2
00
0
50
41
34
-94
28
50
1
35
9.8
8
.6
61
.9
2.4
1
50
6
Kir
k
20
00
4
83
90
0-9
20
70
0
86
.2
3.3
1
0.0
2
.0
15
00
Klo
tz
20
01
4
94
80
7-8
55
14
6
83
0.1
7
.8
19
.8
3.2
1
27
5
Ku
ku
ku
s
20
00
4
94
61
9-9
14
11
0
41
28
.6
5.0
1
9.5
2
.4
15
05
La
c la
Cro
ix
19
99
4
82
10
0-9
20
90
0
57
71
.0
34
.0
54
.0
3.2
1
58
2
Lig
ht
20
00
4
85
63
8-9
11
83
2
18
7.2
4
.1
13
.7
2.5
1
38
0
Little
Eye
2
00
0
48
50
36
-91
46
51
7
5.4
7
.5
26
.5
2.0
1
38
3
Little
Sa
nd
ford
2
00
0
49
06
00
-91
36
28
2
92
1.7
3
6.8
1
14
.7
11
.7
13
80
Little
Tro
ut
19
98
5
10
31
0-9
31
42
0
36
65
.5
6.7
1
6.5
3
.7
14
02
Little
Tu
rtle
2
00
0
48
47
28
-92
41
57
2
22
5.0
3
.6
9.2
1
.8
16
22
Lo
ng
leg
ge
d
19
97
5
04
70
2-9
41
02
7
69
15
.5
8.9
3
5.4
2
.7
14
81
Lo
un
t 1
99
7 &
19
98
5
01
00
6-9
41
83
0
30
14
.4
7.0
3
2.0
1
.7
15
86
Ma
invill
e
19
98
4
85
20
6-9
31
33
8
86
7.7
3
.9
10
.7
2.3
1
64
2
NW
- 4
Ta
ble
NW
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Ma
ko
kib
ata
n
20
01
5
11
62
9-8
72
05
3
58
19
.1
6.1
1
3.0
2
.3
11
71
Ma
nio
n
20
01
4
85
44
1-9
21
85
6
11
97
.9
5.6
1
2.0
3
.0
15
77
Ma
no
mim
2
00
0
48
52
00
-93
43
00
1
26
.3
3.0
8
.0
2.9
1
50
0
Ma
rmo
in
19
94
4
85
23
0-9
13
25
0
10
51
5.7
6
.8
30
.5
2.4
1
46
3
McA
ree
1
99
6
48
18
02
-91
56
21
8
45
.0
12
.7
37
.2
2.0
1
55
3
Me
lgu
nd
1
99
7
49
38
04
-92
19
27
1
20
1.8
4
.7
11
.3
3.2
1
56
3
Me
rcu
tio
2
00
0
48
48
58
-91
07
00
1
47
3.1
4
.9
15
.3
1.8
1
38
2
Me
ta
19
98
5
03
05
9-8
72
22
0
30
79
.7
1.7
1
1.6
2
.5
11
70
Mim
inis
ka
2
00
1
51
33
11
-88
35
55
6
19
5.7
2
.8
17
.2
2.8
1
14
8
Min
n
19
99
4
82
04
2-9
15
41
4
46
3.8
5
.9
39
.7
3.1
1
54
8
Min
nita
ki
20
01
4
95
82
6-9
15
83
1
18
08
7.9
1
7.8
4
8.5
2
.6
15
36
Mo
un
t 2
00
0
49
00
47
-92
10
47
1
08
1.3
1
2.3
3
7.2
3
.2
14
29
Na
ma
ka
n
20
00
4
82
70
0-9
23
50
0
51
50
.0
15
.8
45
.3
2.6
1
62
4
Ne
sto
n
20
01
5
01
63
3-9
03
05
7
50
5.6
2
.1
11
.0
3.5
1
34
5
Nip
igo
n4
.1
99
7,
19
98
, 1
99
9,
20
00
&
20
01
49
50
00
-88
30
00
4
48
06
0.0
5
4.9
1
37
.0
6.5
1
20
5
No
rth
ern
Lig
ht
19
98
4
81
50
4-9
04
04
5
68
69
.8
8.4
3
9.7
3
.4
13
59
Ob
on
ga
1
99
9
49
58
58
-89
18
26
3
57
8.4
1
7.4
7
1.7
2
.7
12
27
of
the
Wo
od
s4
.1
99
7 &
20
00
4
91
45
9-9
44
50
2
24
67
18
.0
7.9
6
8.8
7
.2
16
41
Old
Ma
n
20
00
4
90
20
4-9
10
21
1
47
3.2
1
3.5
2
6.5
2
.3
13
75
On
am
an
1
99
8
50
00
17
-87
26
09
1
08
38
.0
2.2
1
8.9
1
.4
11
53
Ott
er
19
98
5
10
55
6-9
31
74
3
97
9.0
3
.8
10
.7
5.8
1
39
6
Otu
ka
ma
mo
an
1
99
8
48
57
42
-92
51
48
5
16
5.8
1
6.9
5
3.6
3
.7
16
16
Pa
rtri
dg
e
20
01
4
94
30
9-8
72
30
9
65
9.5
5
.1
30
.5
2.8
1
14
8
Pa
tric
ia
20
00
4
90
44
8-9
22
34
9
95
.1
5.9
1
1.0
4
.2
13
87
Pe
ka
go
nin
g
19
98
& 1
99
9
49
09
00
-92
11
27
1
31
9.6
1
0.6
3
8.4
4
.7
15
66
NW
- 5
Ta
ble
NW
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Pe
lica
n
20
01
5
00
70
4-9
15
80
7
23
42
.0
10
.4
35
.1
1.8
1
51
8
Pe
rch
1
99
8
48
44
49
-91
50
41
7
03
.5
5.9
2
4.7
1
.8
15
02
Pe
ttit
20
00
4
85
70
7-9
21
60
2
11
97
.1
10
.6
25
.0
3.8
1
44
8
Pic
kw
ick
19
99
4
85
93
8-9
30
51
1
49
2.1
8
.2
31
.1
3.7
1
60
9
Pip
esto
ne
2
00
0
49
05
28
-93
33
15
3
89
0.7
1
8.6
5
3.4
4
.3
16
40
Po
ny
20
00
4
85
40
0-9
35
20
0
29
.5
4.9
1
2.2
2
.0
16
47
Qu
etico
1
99
7
48
33
52
-91
56
35
4
26
1.3
1
3.1
6
1.0
3
.4
15
25
Ra
iny
4.
19
97
, 1
99
8 &
1
99
94
84
20
0-9
31
00
0
70
15
0.0
9
.3
49
.1
2.7
1
64
6
Ra
wn
Re
se
rvo
ir
20
00
4
84
70
4-9
13
20
8
21
2.8
4
.2
16
.0
2.6
1
38
3
Re
d
20
00
5
10
23
4-9
35
65
4
17
67
6.7
1
4.6
4
2.7
2
.3
14
50
Ric
ha
rdso
n
20
00
4
90
15
7-9
10
70
5
14
8.4
5
.1
9.3
3
.5
13
75
Ro
bin
so
n
19
99
4
81
14
6-9
13
94
0
42
0.9
1
2.7
3
5.1
6
.6
15
29
Ru
gb
y
19
97
4
95
72
8-9
25
74
8
98
4.4
3
.4
7.6
1
.1
15
56
Sa
kw
ite
2
00
1
49
01
49
-92
54
00
3
79
.3
8.8
2
4.5
2
.8
16
12
Sa
nd
Po
int
20
00
4
82
23
6-9
22
80
5
35
96
.0
11
.6
56
.1
3.0
1
61
7
Sa
nd
sto
ne
1
99
4
48
14
07
-90
14
08
9
35
.2
7.3
2
3.8
4
.5
12
83
Sa
pa
we
2
00
0
48
46
31
-91
20
32
5
03
.9
5.1
1
1.3
5
.2
13
84
Sa
va
nn
e1
99
6,
19
97
&
20
01
48
50
00
-90
06
00
3
64
.0
2.6
4
.3
1.5
1
24
4
Se
pa
ratio
n
19
96
& 1
99
7
50
14
45
-94
23
43
4
89
5.3
6
.9
35
.7
1.5
1
58
0
Se
ul
19
96
& 1
99
7
20
19
30
-92
29
13
1
40
94
3.3
9
.5
47
.2
1.5
1
50
2
Sh
oa
l 1
99
8 &
20
01
4
93
30
0-9
50
10
0
25
90
0.0
8
.9
35
.6
3.6
1
61
4
Sm
oo
thro
ck
20
01
5
03
13
1-8
92
64
7
98
24
.5
5.4
4
7.2
2
.7
11
91
St.
Jo
se
ph
1
99
9
51
04
36
-90
43
25
5
15
40
.8
5.1
2
6.2
2
.6
13
01
Ste
el
20
00
4
91
21
1-8
65
04
3
15
82
.5
16
.2
64
.1
3.7
1
09
7
Ste
ep
Ro
ck
19
95
4
84
80
0-9
14
00
0
61
6.0
3
.6
14
.5
3.4
1
47
6
Str
aw
1
99
9
49
07
59
-93
23
31
4
27
.4
4.6
1
1.0
2
.7
16
31
NW
- 6
Ta
ble
NW
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Stu
rge
on
1
99
5
49
58
29
-90
46
07
2
14
12
.7
13
.6
93
.0
4.0
1
40
0
Su
ca
n
20
00
4
90
65
9-9
32
44
2
17
6.1
3
.2
16
.0
3.2
1
49
9
Su
rpri
se
2
00
0
48
22
00
-92
45
00
2
47
.3
3.0
9
.0
1.5
1
49
9
Syd
ne
y
20
00
5
03
91
0-9
42
63
4
58
68
.2
19
.7
70
.1
7.9
1
50
7
Th
ree
Fin
ger
20
01
4
84
33
3-8
61
90
3
24
.9
4.1
1
0.7
3
.2
11
29
To
ron
to
20
00
5
02
11
1-8
74
91
7
98
2.8
1
0.0
4
0.3
3
.1
11
57
Tro
ut
19
98
5
11
15
5-9
31
83
3
34
80
6.9
1
3.7
4
7.3
5
.8
13
88
Tw
ist
20
00
4
90
21
5-9
30
42
6
10
9.3
6
.0
21
.4
4.1
1
50
0
Tyre
ll 2
00
0
48
49
53
-91
25
12
1
17
.4
3.2
9
.3
2.7
1
38
3
Un
ion
2
00
0
48
49
00
-90
58
00
2
36
.3
3.9
1
6.0
2
.0
13
81
Vis
ta
20
00
4
90
34
9-9
30
44
2
85
.0
4.9
2
1.6
3
.3
13
79
Wa
ba
bim
iga
1
99
7
50
20
38
-86
22
25
1
99
8.2
5
.1
18
.7
2.4
1
21
0
Wa
big
oo
n
20
01
4
94
41
7-9
24
30
7
99
22
.9
6.1
1
4.6
0
.9
15
85
Wa
pe
si
19
98
5
03
32
7-9
22
04
3
23
66
.4
2.0
9
.1
1.8
1
45
4
Wa
tco
mb
1
99
6
49
51
40
-91
18
06
1
05
4.5
7
.0
15
.6
3.9
1
47
8
Wa
wa
ng
1
99
8
49
25
35
-90
33
07
2
02
1.5
6
.5
26
.5
2.3
1
33
7
Wh
ale
n
20
00
4
90
35
2-9
31
43
2
29
1.4
3
.6
9.2
2
.0
15
00
Wh
ite
Ott
er
19
99
4
90
64
0-9
15
26
3
82
49
.0
22
.1
56
.4
4.8
1
50
8
Wh
ite
fish
19
95
, 1
99
6 &
2
00
14
81
30
2-8
95
95
5
30
15
.0
2.0
6
.4
1.5
1
29
3
Wh
ite
rock
19
98
4
95
22
0-9
11
45
4
77
6.9
3
.8
9.5
2
.6
14
68
Wh
ite
wa
ter
19
98
5
04
83
5-8
91
13
5
10
52
9.9
4
.6
22
.9
3.2
1
15
4
Wild
go
ose
1
99
9
49
44
05
-87
09
50
1
73
7.7
3
.6
15
.6
4.3
1
13
4
Win
nip
eg
Riv
er
19
97
& 1
99
8
50
14
00
-95
09
00
1
71
40
.0
12
.0
No
da
ta
3.0
1
60
5
Win
teri
ng
2
00
1
49
26
14
-87
16
22
1
66
5.2
4
.8
36
.6
3.2
1
11
1
Wo
lse
ley
19
97
4
82
60
0-9
20
50
0
13
38
.0
12
.6
40
.0
2.6
1
56
2
Yo
un
g
19
98
4
95
10
3-9
11
32
6
80
4.7
5
.1
11
.9
2.9
1
47
1
NW
- 7
Ta
ble
NW
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Ziz
an
ia
19
99
5
03
10
0-9
33
10
0
52
3.0
9
.9
19
.0
2.5
1
50
0
1.
Latitu
de a
nd longitude in d
egre
es,
min
ute
s,
and s
econds.
2.
GD
D>
5 is g
row
ing-d
egre
e-d
ays a
bove 5
o C
els
ius.
3.
Com
panio
n L
ake is d
ivid
ed in
to U
pper
and L
ow
er
Com
pa
nio
n L
ake (
i.e., s
epara
te F
ISH
NE
T p
roje
cts
).
4.
Data
not
inclu
ded in t
he lake c
hara
cte
ristics s
um
mary
sta
tistics b
ecause o
nly
a s
mall
port
ion o
f th
e e
ntire
wate
rbody w
as s
am
ple
d.
Ch
ara
cte
ris
tic
s o
f n
ort
hw
es
t re
gio
n l
ak
es
sa
mp
led
us
ing
fa
ll w
all
eye
in
de
x n
ett
ing
sta
nd
ard
.
S
urf
ace
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
Me
an
4
22
1.5
7
.7
26
.5
3.1
1
41
4
Min
imu
m
19
.5
1.4
3
.0
0.8
1
09
7
Ma
xim
um
1
40
94
3.3
3
6.8
1
17
.0
11
.7
16
47
Me
dia
n
97
9.0
6
.0
20
.2
2.9
1
44
8
5%
Pe
rce
ntile
9
5.1
2
.0
6.4
1
.5
11
48
25
% Q
ua
rtile
3
89
.1
3.9
1
2.2
2
.3
13
30
75
% Q
ua
rtile
3
01
4.4
9
.5
35
.6
3.6
1
51
3
95
% P
erc
en
tile
1
80
87
.9
18
.6
64
.1
5.6
1
62
2
Sa
mp
le S
ize
1
55
1
54
1
54
1
55
1
56
NW - 8
Table NW-2. Walleye relative abundance and average size for northwest region waterbodies sampled using the fall walleye index netting standard 1993 to 2001.
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
Abram 14.2 0.20 12 12 384 628
Agimak 16.3 0.19 9 9 268 352
Alph 1.2 0.26 12 8 478 1506
Anders 1.8 0.34 6 5 634 3125
Ara 26.0 0.11 10 10 384 584
Ball (19973.
) 11.2 0.49 10 9 357 479
Ball (1998) 13.1 0.52 12 11 403 932
Bar 2.2 1.03 4 3 486 1308
Barr 1.3 0.50 4. 3 563 1903
Beak 3.1 0.55 6 5 302 427
Beaverhouse 0.9 0.27 15 6 531 1714
Bending 9.8 0.22 12 12 308 332
Boffin 3.2 0.46 8 6 494 1424
Botsford 20.4 0.19 12 12 344 482
Bradshaw 3.7 0.34 4 4 418 755
Brennan 3.6 0.19 24 20 350 458
Brewer 5.8 0.14 4 4 327 485
Broad 26.4 0.20 4 4 453 1104
Burditt 9.4 0.27 17 16 411 800
Burrows 13.4 0.13 15 15 315 406
Calm 5.5 0.46 13 11 313 360
Chill 3.4 0.35 4 4 405 692
Churchill 29.1 0.11 12 12 364 553
Cloud 17.8 0.18 5 5 569 2525
Upper Companion 2.2 0.39 6 5 390 873
Lower Companion 11.3 0.39 6 6 350 513
Crooked Pine 1.2 0.28 17 7 525 1732
Cuttle 6.5 0.40 9 8 350 448
Darby 5.3 0.75 4 4 391 610
des Mille Lacs (19933.
) 15.3 0.21 22 22 336 487
des Mille Lacs (19943.
) 18.8 0.13 24 24 318 467
des Mille Lacs (1999) 17.2 0.10 38 38 366 643
Despair 7.1 0.35 8 8 375 562
Devious 7.5 0.15 12 12 303 345
Dimple 8.4 0.30 6 6 367 628
Dog 20.4 0.19 25 24 370 699
Dore 1.6 0.21 12 10 358 714
Dovetail 10.3 0.32 8 8 386 648
Drum 1.6 0.39 4 3 424 1030
Eagle 9.2 0.15 51 47 394 813
Elbow 6.4 0.33 9 8 277 314
Eltrut 15.7 0.12 12 12 316 316
Elva 7.0 0.25 7 7 337 519
NW - 9
Table NW-2. (continued)
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
Expanse 20.4 0.15 10 10 391 786
Factor 15.4 0.39 7 7 390 729
Favourable 18.0 0.09 20 20 378 665
Finlayson 10.0 0.53 11 9 318 401
Footprint 11.1 0.21 8 8 347 603
Gammon 3.1 0.20 18 16 517 1626
Gamsby 11.5 0.20 10 10 311 356
Garden 7.0 0.17 12 12 327 414
Geikie 7.0 0.14 10 10 335 479
Georgia 10.9 0.18 14 14 370 655
Harris 9.8 0.16 11 11 341 477
Henderson 9.6 0.08 12 12 450 857
Hicks 5.0 0.23 12 11 334 437
Husband 12.1 0.32 4 4 381 785
Hutchison 6.7 0.39 10 9 320 566
Icarus 11.2 0.29 13 13 378 824
Ice 8.5 0.13 8 8 401 720
Iron Range 24.2 0.23 8 8 384 631
Jac Saga 0.4 0.44 3 1 572 1863
Jackfish 6.7 0.44 9 8 418 1127
Jones 25.0 0.12 3 3 348 452
Jutten 11.4 0.25 11 11 340 457
Kaopskikamak 5.9 0.27 10 10 389 636
Kapkichi 14.8 0.15 16 16 305 361
Kawawiag 6.0 0.32 6 6 382 751
Keckush 3.4 0.69 4 3 289 356
Keikewabik 19.2 0.17 12 12 324 341
Kenogamisis 23.7 0.25 8 8 308 383
Kenorain 16.3 0.23 8 8 497 1627
Kenozhe 2.6 0.30 4 4 374 601
Kilburn 11.6 0.42 11 10 403 724
Kirk 3.1 0.37 4 4 390 703
Klotz 17.2 0.27 9 9 494 1372
Kukukus 18.8 0.20 12 12 406 824
Lac la Croix 9.8 0.56 4 4 411 883
Light 1.4 0.68 4 2 348 511
Little Eye 4.2 0.05 4 4 381 803
Little Sandford 14.9 0.22 12 12 305 344
Little Trout 22.1 0.15 11 11 373 675
Little Turtle 2.2 0.22 12 10 312 307
Longlegged 13.9 0.18 16 16 423 904
Lount (19973.
) 3.3 0.47 10 8 378 605
Lount (1998) 6.0 0.39 12 11 384 724
Mainville 16.3 0.14 12 12 325 395
NW - 10
Table NW-2. (continued)
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
Makokibatan 13.7 0.13 14 14 366 677
Manion 6.1 0.29 12 11 376 635
Manomin 9.2 0.30 4 4 323 335
Marmion 14.0 0.37 7 7 360 633
McAree 6.2 0.28 15 13 343 499
Melgund 16.8 0.15 17 17 325 433
Mercutio 8.8 0.27 10 10 382 723
Meta 20.0 0.08 6 6 371 578
Miminiska 20.6 0.14 9 9 407 684
Minn 3.7 0.42 8 6 384 658
Minnitaki 19.0 0.23 24 23 382 669
Mount 14.5 0.27 10 10 386 576
Namakan 5.4 0.18 25 23 318 354
Neston 8.3 0.17 12 12 345 454
Nipigon (19973.
)
Ombabika Bay23.4 0.28 24 23 294 302
Nipigon (19983.
)
Ombabika Bay 11.4 0.16 18 16 286 277
Nipigon (1999)
Ombabika Bay8.2 0.44 18 15 351 514
Nipigon (2000)
Wabinosh Bay 8.8 0.36 28 23 300 296
Nipigon (2001)
Gull Bay (GB)5.5 0.17 50 43 332 451
Nipigon (2001)
Chief Bay (CB)1.0 0.16 24 13 374 622
Northern Light 14.9 0.30 12 12 324 811
Obonga 8.0 0.19 20 20 349 476
of the Woods (1997) 10.4 0.11 60 60 259 248
of the Woods (2000)
Sabaskong Bay 7.7 0.09 50 50 339 472
Old Man 7.0 0.34 6 6 357 525
Onaman 14.9 0.17 14 14 400 840
Otter 13.9 0.32 5 5 353 545
Otukamamoan 12.9 0.27 19 19 354 474
Partridge 21.6 0.17 10 10 335 396
Patricia 18.9 0.11 4 4 392 565
Pekagoning (19983.
) 17.7 0.25 13 13 327 395
Pekagoning (1999) 21.1 0.16 13 13 344 436
Pelican 17.6 0.38 12 11 356 499
Perch 8.8 0.27 8 8 320 403
Pettit 10.6 0.37 10 10 384 620
Pickwick 10.7 0.22 12 12 314 308
Pipestone 5.1 0.28 20 17 484 1332
Pony 0.3 0.26 3 1 435 750
NW - 11
Table NW-2. (continued)
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
Quetico 1.5 0.22 12 9 530 1596
Rainy (1997)
North Arm 9.4 0.29 20 19 436 1125
Rainy (1998)
Redgut Bay 6.5 0.16 12 12 289 287
Rainy (1999)
South Arm 10.0 0.18 20 20 364 557
Rawn Reservoir 7.1 0.79 6 5 321 413
Red 29.4 0.15 39 38 350 664
Richardson 12.4 0.32 4 4 340 719
Robinson 4.1 0.30 16 13 479 1150
Rugby 15.5 0.19 14 14 296 372
Sakwite 10.6 0.36 11 10 352 454
Sand Point 8.1 0.18 20 19 365 597
Sandstone 11.8 0.62 8 7 351 630
Sapawe 14.0 0.20 6 6 309 351
Savanne (19963.
) 8.0 0.16 12 12 403 678
Savanne (19973.
) 16.9 0.16 12 12 384 671
Savanne (2001) 10.5 0.12 12 12 402 599
Separation (19963.
) 9.0 0.28 12 11 363 567
Separation (1997) 4.8 0.37 12 10 338 482
Seul (19963.
) 14.1 0.11 64 64 363 650
Seul (1997) 16.2 0.10 62 62 359 673
Shoal (19983.
) 4.8 0.22 20 18 424 844
Shoal (2001) 4.1 0.13 60 51 398 1207
Smoothrock 12.2 0.14 24 24 340 455
St. Joseph 25.6 0.09 37 37 389 680
Steel 4.9 0.20 28 26 332 359
Steep Rock 2.9 0.32 12 9 370 850
Straw 12.8 0.13 12 12 356 507
Sturgeon 14.9 0.31 6 6 414 715
Sucan 16.3 0.25 4 4 332 489
Surprise 4.3 0.25 6 6 330 394
Sydney 18.1 0.21 12 12 374 637
Three Finger 18.5 0.25 6 6 323 331
Toronto 9.0 0.41 14 12 365 410
Trout 8.9 0.22 23 23 367 634
Twist 0 - 4 0 - -
Tyrell 5.1 0.32 4 4 426 822
Union 4.6 0.45 6 5 436 982
Vista 3.0 0.63 4 3 459 1241
Wababimiga 1.0 0.14 24 16 426 1387
Wabigoon 3.2 0.11 61 52 349 602
Wapesi 22.1 0.08 11 11 360 485
NW - 12
Table NW-2. (continued)
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
Watcomb 17.7 0.15 6 6 349 594
Wawang 3.5 0.36 16 11 404 1111
Whalen 8.5 0.21 6 6 369 550
White Otter 12.0 0.31 12 12 258 261
Whitefish (19953.
) 30.9 0.06 10 10 355 698
Whitefish (19963.
) 30.0 0.17 12 12 361 597
Whitefish (2001) 33.4 0.19 12 12 408 756
Whiterock 1.6 0.20 12 9 504 1382
Whitewater 17.4 0.18 12 12 376 573
Wildgoose 23.7 0.13 8 8 367 509
Winnipeg River (19973.
) 10.5 0.15 40 40 328 487
Winnipeg River (1998) 10.9 0.14 34 34 328 437
Wintering 11.1 0.25 15 14 340 484
Wolseley 3.4 0.24 14 14 409 782
Young 17.6 0.21 9 9 415 1099
Zizania 23.7 0.13 6 6 396 702
1. Average catch is the geometric mean number of walleye caught net-1
.2. Average size is the arithmetic mean of total length in mm and wet weight in grams. 3. Data not used to calculate relative abundance or average size benchmarks.
Northwest region walleye relative abundance benchmarks.
Average Size AverageCatch Total
LengthWeight
Mean 10.7 376 705
Minimum 0 258 248
Maximum 33.4 634 3125
Median 9.8 367 606
5% Percentile 1.3 302 331
25% Quartile 5.1 335 454
75% Quartile 15.5 401 784
95% Percentile 23.7 504 1596
Sample Size 167 166
NW - 13
Table NW-3a. Annual female walleye mortality (%) for northwest region waterbodies sampled using the fall walleye index netting standard. “Observed mortality” is a direct estimate based on age structure of female fish and was calculated from age 5 onwards using Robson and Chapman’s estimator. “Estimated mortality” is an indirect estimate based on age structure of the population regardless of sex using the empirical formula: Female mortality = 0.952(Combined sex mortality) (n=246, R2=0.77, standard error of the estimate=6.493).
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Manomin No data 7.6% Three Finger 19% 19.0%
Wababimiga 7% 8.6% Jackfish 18% 19.0%
Alph 7% 9.5% Henderson (1997) 21% 19.0%
Crooked Pine 10% 10.5% Savanne (1997) 22% 20.0%
Manion 9% 10.5% Otukamamoan 19% 20.0%
Cuttle 7% 11.4% Seul (1997) 23% 20.0%
Anders No data 11.4% Wapesi 25% 20.0%
Dore No data 12.4% Gammon 21% 20.0%
Elva No data 12.4% Robinson 19% 20.0%
Vista No data 12.4% Old Man 19% 20.0%
White Otter 13% 13.3% Quetico No data 20.0%
Miminiska 14% 13.3% Winnipeg River (1998) 20% 20.0%
Trout 15% 13.3% Burditt 17% 20.9%
Husband No data 14.3% Otter 24% 20.9%
Kawawiag 13% 14.3% Expanse 19% 20.9%
Northern Light 14% 15.2% Kilburn 21% 20.9%
Ara 17% 15.2% Kaopskikamak 21% 20.9%
Onaman No data 15.2% Dimple 15% 20.9%
Pipestone 16% 15.2% Wildgoose 22% 21.9%
Young 15% 15.2% Boffin 23% 21.9%
Meta 16% 15.2% Steep Rock 28% 21.9%
Beaverhouse No data 16.2% Mainville 25% 21.9%
Georgia 18% 16.2% Shoal (2001) 21% 21.9%
Richardson No data 16.2% Separation (1997) No data 22.8%
Rainy (1998) 11% 16.2% Zizania 23% 22.8%
Keckush No data 16.2% Cloud 24% 22.8%
Iron Range 15% 16.2% Lount (1997) No data 22.8%
Little Trout 18% 16.2% Mercutio 19% 22.8%
Pekagoning 14% 16.2% Longlegged 24% 22.8%
Hicks 19% 16.2% Factor 17% 22.8%
St. Joseph 21% 17.1% Nipigon (1998) 28% 22.8%
Chill No data 17.1% Eagle 25% 22.8%
Ice 17% 17.1% Union 23% 22.8%
Favourable 16% 17.1% Kapkichi 23% 22.8%
Upper Companion 18% 17.1% Lount (1998) No data 23.8%
Churchill 19% 18.1% Footprint 24% 23.8%
Hutchison 18% 18.1% Klotz 27% 23.8%
Little Eye No data 18.1% Icarus 25% 23.8%
Devious 17% 18.1% Kenogamisis 19% 23.8%
Whitewater 18% 18.1% Sandstone No data 23.8%
Wawang 16% 18.1% Whitefish (1995) 23% 24.8%
Red 19% 19.0% Sydney 22% 24.8%
NW - 14
Table NW-3a. (continued)
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Savanne (2001) 19% 19.0% Neston 30% 24.8%
Ball (1998) 22% 24.8% Botsford 43% 34.3%
Broad 25% 24.8% Obonga 30% 34.3%
Kukukus 30% 25.7% Bending 40% 34.3%
Wolseley 21% 25.7% Wintering 34% 34.3%
Seul (1996) 29% 25.7% Calm 35% 34.3%
Minnitaki 30% 25.7% Geikie 37% 34.3%
Makokibatan 23% 25.7% Despair 32% 35.2%
Nipigon (1997) 29% 26.7% des Mille Lacs (1994) 36% 36.2%
Whitefish (2001) 27% 26.7% Shoal (1998) 13% 36.2%
Harris 33% 26.7% Gamsby 38% 36.2%
Savanne (1996) 26% 26.7% Whitefish (1996) 35% 36.2%
des Mille Lacs (1999) 27% 26.7% Agimak 33% 37.1%
Lower Companion 25% 27.6% Kenorain 29% 37.1%
Whiterock 29% 27.6% Minn No data 38.1%
Garden 29% 27.6% Abram 43% 38.1%
Smoothrock 31% 27.6% Rugby No data 39.0%
Finlayson 27% 27.6% Pettit 61% 40.0%
Brennan 27% 27.6% Straw 33% 40.9%
Perch 24% 28.6% Sakwite 45% 40.9%
Sucan No data 28.6% Pelican 49% 41.9%
Bar No data 28.6% Tyrell 44% 41.9%
Sand Point 37% 28.6% Watcomb No data 41.9%
Marmion 30% 28.6% Namakan 53% 43.8%
Whalen 27% 29.5% Sapawe 44% 43.8%
Wabigoon 30% 29.5% Rainy (1999) 45% 44.7%
Steel 39% 29.5% Nipigon (1999) 53% 44.7%
Separation (1996) No data 29.5% Dovetail 42% 45.7%
Ball 1997 No data 30.5% Patricia 73% 46.7%
of the Woods (2000) 35% 30.5% Brewer No data 47.6%
Rawn Reservoir 46% 30.5% Pickwick No data 50.5%
Rainy (1997) 33% 30.5% Mount 44% 51.4%
des Mille Lacs (1993) 33% 31.4% Lac la Croix 38% 52.4%
Beak No data 31.4% Sturgeon 59% 52.4%
Toronto 31% 31.4% Bradshaw No data 53.3%
Pekagoning (1999) 32% 31.4% Nipigon (2001) (CB) 22% 57.1%
Keikewabik 34% 32.4% Eltrut 60% 58.1%
McAree 23% 32.4% Darby No data 60.0%
Kirk No data 33.3% Jones 67% 60.0%
Jutten 28% 33.3% Little Sandford 58% 60.9%
Burrows 35% 33.3% Nipigon (2001) (GB) 71% 71.4%
Dog 33% 33.3% Partridge 78% 80.9%
NW - 15
Northwest region female walleye mortality benchmarks.
Observed Mortality
EstimatedMortality
Mean 28% 27.5%
Minimum 7% 7.6%
Maximum 78% 80.9%
Median 25% 24.8%
5% Percentile 13% 12.4%
25% Quartile 19% 19.0%
75% Quartile 33% 33.3%
95% Percentile 59% 52.4%
Sample Size 138 168
NW - 16
Table NW-3b. Annual male walleye mortality (%) for northwest region waterbodies sampled using the fall walleye index netting standard. “Observed mortality” is a direct estimate based on age structure of male fish and was calculated from age 5 onwards using Robson and Chapman’s estimator. “Estimated mortality” is an indirect estimate based on age structure of the population regardless of sex using the empirical formula: Male mortality = 0.945(Combined sex mortality) + 2.844 (n=242, R2=0.65, standard error of the estimate=7.477).
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Manomin No data 10.4% Three Finger 20% 21.8%
Wababimiga 9% 11.4% Jackfish 29% 21.8%
Alph No data 12.3% Henderson 19% 21.8%
Crooked Pine 11% 13.2% Savanne (1997) 20% 22.7%
Manion 6% 13.2% Otukamamoan 25% 22.7%
Cuttle 15% 14.2% Seul (1997) 18% 22.7%
Anders No data 14.2% Wapesi 20% 22.7%
Dore No data 15.1% Gammon 19% 22.7%
Elva No data 15.1% Robinson 21% 22.7%
Vista No data 15.1% Old Man 20% 22.7%
White Otter 14% 16.1% Quetico No data 22.7%
Miminiska 14% 16.1% Winnipeg River (1998) 22% 22.7%
Trout 13% 16.1% Burditt 27% 23.7%
Husband No data 17.0% Otter 16% 23.7%
Kawawiag 14% 17.0% Expanse 27% 23.7%
Northern Light 19% 18.0% Kilburn 22% 23.7%
Ara 15% 18.0% Kaopskikamak 18% 23.7%
Onaman No data 18.0% Dimple 33% 23.7%
Pipestone 16% 18.0% Wildgoose 25% 24.6%
Young 17% 18.0% Boffin 20% 24.6^
Meta 16% 18.0% Steep Rock 18% 24.6%
Beaverhouse 18% 18.9% Mainville 22% 24.6%
Georgia 15% 18.9% Shoal (2001) 30% 24.6%
Richardson No data 18.9% Separation (1997) No data 25.5%
Rainy (1998) 19% 18.9% Zizania 24% 25.5%
Keckush No data 18.9% Cloud 24% 25.5%
Iron Range 20% 18.9% Lount (1997) No data 25.5%
Little Trout 16% 18.9% Mercutio 34% 25.5%
Pekagoning (1998) 31% 18.9% Longlegged 24% 25.5%
Hicks 14% 18.9% Factor 26% 25.5%
St. Joseph 16% 19.9% Nipigon (1998) 19% 25.5%
Chill No data 19.9% Eagle 24% 25.5%
Ice 16% 19.9% Union 22% 25.5%
Favourable 19% 19.9% Kapkichi 25% 25.5%
Upper Companion No data 19.9% Lount (1998) No data 26.5%
Churchill 19% 20.8% Footprint 21% 26.5%
Hutchison 18% 20.8% Klotz 22% 26.5%
Little Eye No data 20.8% Icarus 25% 26.5%
Devious No data 20.8% Kenogamisis 33% 26.5%
Whitewater 19% 20.8% Sandstone No data 26.5%
Wawang 24% 20.8% Whitefish (1995) 34% 27.4%
Red 23% 21.8% Sydney 32% 27.4%
NW - 17
Table NW-3b. (continued)
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Savanne (2001) 22% 21.8% Neston 23% 27.4%
Ball (1998) 29% 27.4% Botsford 31% 36.9%
Broad 26% 27.4% Obonga No data 36.9%
Kukukus 25% 28.4% Bending 39% 36.9%
Wolseley 38% 28.4% Wintering 38% 36.9%
Seul (1996) 25% 28.4% Calm 34% 36.9%
Minnitaki 24% 28.4% Geikie 31% 36.9%
Makokibatan 39% 28.4% Despair 42% 37.8%
Nipigon (1997) 21% 29.3% des Mille Lacs (1994) 41% 38.8%
Whitefish (2001) 29% 29.3% Shoal (1998) 75% 38.8%
Harris 24% 29.3% Gamsby 36% 38.8%
Savanne (1996) 30% 29.3% Whitefish (1996) 41% 38.8%
des Mille Lacs (1999) 29% 29.3% Agimak 66% 39.7%
Lower Companion 27% 30.3% Kenorain 71% 39.7%
Whiterock No data 30.3% Minn No data 40.7%
Garden 29% 30.3% Abram 38% 40.7%
Smoothrock 26% 30.3% Rugby No data 41.6%
Finlayson 32% 30.3% Pettit No data 42.6%
Brennan 30% 30.3% Straw 56% 43.5%
Perch 41% 31.2% Sakwite 40% 43.5%
Sucan No data 31.2% Pelican 38% 44.5%
Bar 28% 31.2% Tyrell No data 44.5%
Sand Point 25% 31.2% Watcomb No data 44.5%
Marmion 30% 31.2% Namakan 41% 46.4%
Whalen No data 32.2% Sapawe 43% 46.4%
Wabigoon 32% 32.2% Rainy (1999) 49% 47.3%
Steel 25% 32.2% Nipigon (1999) 37% 47.3%
Separation (1996) No data 32.2% Dovetail 62% 48.2%
Ball 1997 No data 33.1% Patricia 30% 49.2%
of the Woods (2000) 26% 33.1% Brewer No data 50.1%
Rawn Reservoir 38% 33.1% Pickwick 47% 53.0%
Rainy (1997) 30% 33.1% Mount 60% 53.9%
des Mille Lacs (1993) 33% 34.1% Lac la Croix No data 54.9%
Beak 50% 34.1% Sturgeon 48% 54.9%
Toronto 37% 34.1% Bradshaw 43% 55.8%
Pekagoning (1999) 33% 34.1% Nipigon (2001) (CB) No data 59.6%
Keikewabik 34% 35.0% Eltrut 62% 60.5%
McAree 44% 35.0% Darby 7% 62.4%
Kirk No data 35.9% Jones 82% 62.4%
Jutten 50% 35.9% Little Sandford 68% 63.4%
Burrows 35% 35.9% Nipigon (2001) (GB) No data 73.8%
Dog 40% 35.9% Partridge No data 83.2%
NW - 18
Northwest region male walleye mortality benchmarks.
Observed Mortality
EstimatedMortality
Mean 30% 30.1%
Minimum 6% 10.4%
Maximum 82% 83.2%
Median 26% 27.4%
5% Percentile 14% 15.1%
25% Quartile 20% 21.8%
75% Quartile 36% 35.9%
95% Percentile 62% 54.9%
Sample Size 132 168
NW - 19
Table NW-4a. Predicted female walleye weight-at-total length for northwest region waterbodies sampled using the fall walleye index netting standard. A geometric mean predictive regression model was fitted to logarithmically transformed (base 10) individual observations (only
lakes with 30 fish sampled). Weight in grams and total length in millimetres.
Weight-length relation1. Calculated weight at standard total lengthsLake
x10-6 350 400 450 500 550 Condition2.
Favourable 2.71 55.41 435 624 859 1142 1479 1.01
Elbow 2.75 51.86 514 742 1026 1371 1782 1.49
Kenogamisis 2.75 43.27 429 619 856 1144 1487 1.27
Cuttle 2.80 27.73 368 535 745 1000 1306 1.00
Kilburn 2.85 21.27 379 554 775 1047 1373 1.01
Savanne (2001) 2.86 20.33 384 562 788 1065 1398 0.96
Savanne (1997) 2.89 16.75 377 555 779 1057 1392 0.98
Onaman 2.92 18.45 495 731 1031 1403 1853 1.21
Finlayson 2.94 12.66 382 566 800 1090 1442 1.08
Partridge 2.95 11.85 379 562 796 1086 1438 1.01
Little Trout 2.96 12.19 413 614 870 1188 1576 1.07
Cloud 2.99 14.04 568 846 1204 1649 2193 1.34
Churchill 2.99 9.15 370 552 784 1075 1429 1.01
Sydney 2.99 9.97 403 601 855 1171 1557 1.04
Calm 3.01 8.75 398 595 848 1164 1551 1.07
Bending 3.01 7.88 358 535 763 1048 1396 0.98
White Otter 3.03 7.74 396 593 847 1166 1556 1.08
Dovetail 3.04 7.11 385 578 827 1140 1523 1.00
Klotz 3.05 7.70 442 665 952 1313 1756 1.11
Pelican 3.05 6.48 372 560 801 1105 1478 1.01
Henderson 3.05 6.52 375 563 806 1112 1487 0.95
Northern Light 3.07 7.49 484 729 1047 1447 1938 1.28
Wawang 3.08 6.47 443 669 961 1330 1783 1.18
Straw 3.08 5.61 384 580 833 1153 1546 1.03
Keikewabik 3.09 4.72 343 518 745 1032 1386 0.93
Perch 3.09 5.15 374 565 813 1126 1512 1.00
Marmion 3.10 5.11 394 595 858 1189 1598 1.06
Seul (1996) 3.11 5.14 420 636 9178 1273 1712 1.13
Savanne (1996) 3.12 4.47 387 587 848 1178 1586 1.02
Longlegged 3.12 4.51 391 592 855 1188 1600 1.02
Miminiska 3.12 4.10 355 539 778 1080 1455 0.94
Nipigon (2001) (GB) 3.13 4.76 437 664 960 1335 1799 1.15
Rawn Reservoir 3.13 4.05 372 565 817 1136 1530 1.02
Nipigon (2000) 3.14 4.18 407 619 896 1247 1682 1.09
Botsford 3.14 3.62 352 536 776 1080 1457 0.97
Three Finger 3.14 3.71 361 549 795 1107 1493 0.96
Neston 3.14 3.94 384 583 844 1176 1586 1.00
Harris 3.14 3.71 361 549 795 1107 1493 1.00
Nipigon (1997) 3.14 4.18 407 619 896 1247 1682 1.11
Eltrut 3.14 3.69 359 546 791 1101 1485 1.00
Steel 3.15 3.35 346 527 763 1064 1436 0.93
Kaopskikamak 3.15 3.72 384 585 848 1181 1595 1.03
NW - 20
Table NW-4a. (continued)
Weight-length relation1. Calculated weight at standard total lengthsLake
x10-6 350 400 450 500 550 Condition2.
Nipigon (1999) 3.15 4.18 432 657 952 1327 1792 1.14
Gammon 3.15 4.05 418 637 923 1286 1736 1.06
Pekagoning (1998) 3.16 3.11 340 519 753 1051 1420 0.91
Pekagoning (1999) 3.16 3.43 375 573 831 1159 1566 1.01
Pipestone 3.17 3.43 398 608 883 1233 1668 1.08
Gamsby 3.17 2.99 347 530 770 1075 1454 0.94
Geikie 3.17 3.25 377 576 837 1168 1581 1.00
Sakwite 3.17 3.05 354 541 785 1097 1483 0.96
Namakan 3.18 3.03 373 570 829 1159 1570 1.02
Sand Point 3.19 2.78 363 555 809 1132 1534 1.00
Sapawe 3.20 2.82 390 598 872 1222 1657 1.04
Hutchison 3.20 2.81 389 596 869 1217 1651 1.06
Iron Range 3.20 2.71 375 575 838 1174 1593 1.01
Brennan 3.20 2.55 353 541 789 1105 1499 0.96
Whitewater 3.20 2.82 390 598 872 1222 1657 1.02
Kapkichi 3.20 2.75 380 583 850 1191 1616 1.00
Manion 3.21 2.69 395 606 884 1240 1684 1.09
Watcomb 3.21 2.48 364 559 815 1143 1552 1.00
Mount 3.21 2.50 367 563 822 1152 1565 0.98
Lower Companion 3.21 2.74 402 617 901 1263 1715 1.07
St. Joseph 3.21 2.46 361 554 809 1134 1540 0.99
Smoothrock 3.21 2.52 370 568 828 1162 1577 0.98
Footprint 3.21 2.57 377 579 845 1185 1609 1.04
Wabigoon 3.21 2.45 359 552 805 1129 1534 0.99
Factor 3.22 2.37 369 567 828 1163 1580 0.96
Wintering 3.22 2.34 364 560 818 1148 1560 1.01
Meta 3.22 2.37 369 567 828 1163 1580 1.02
McAree 3.23 2.31 381 586 858 1206 1641 1.03
Zizania 3.23 2.39 394 607 888 1248 1697 1.03
Nipigon (1998) 3.23 2.61 431 663 969 1362 1854 1.14
Red 3.23 2.25 371 571 836 1175 1598 1.02
Shoal (2001) 3.24 2.47 432 666 975 1372 1868 1.16
des Mille Lacs (1993) 3.24 2.32 406 625 916 1289 1755 1.13
Trout 3.24 2.03 355 547 802 1128 1536 0.96
Minnitaki 3.24 2.06 360 555 813 1144 1558 0.98
Pickwick 3.24 2.06 360 555 813 1144 1558 0.95
Mercutio 3.24 2.14 374 577 845 1189 1619 1.05
Burrows 3.25 2.10 389 601 881 1241 1692 1.04
Shoal (1998) 3.25 2.04 378 584 856 1206 1644 1.01
Hicks 3.26 1.80 354 547 803 1132 1545 0.96
Eagle 3.26 1.95 383 593 870 1227 1673 1.02
Abram 3.27 1.72 359 555 816 1151 1572 0.95
Rainy (1999) 3.27 1.81 377 584 858 1211 1654 1.01
of the Woods (2000) 3.27 1.84 384 594 873 1232 1682 1.02
Obonga 3.28 1.65 365 565 832 1175 1606 0.97
NW - 21
Table NW-4a. (continued)
Weight-length relation1. Calculated weight at standard total lengthsLake
x10-6 350 400 450 500 550 Condition2.
Otukamamoan 3.28 1.51 334 517 761 1075 1470 0.93
Jutten 3.28 1.66 367 569 837 1182 1616 0.97
Garden 3.29 1.64 384 597 879 1243 1701 1.02
Ice 3.29 1.66 389 604 890 1258 1721 1.06
Kukukus 3.29 1.53 359 557 820 1160 1587 1.00
Winnipeg River (1998) 3.29 1.54 361 560 825 1167 1597 0.97
des Mille Lacs (1994) 3.29 1.64 384 597 879 1243 1701 1.03
Patricia 3.29 1.47 345 535 788 1114 1524 0.92
Wildgoose 3.29 1.49 349 542 798 1129 1545 0.95
Robinson 3.31 1.35 356 554 817 1159 1588 0.97
Pettit 3.31 1.40 369 574 848 1201 1647 1.02
Toronto 3.31 1.36 358 558 824 1167 1600 0.98
Jones 3.31 1.50 395 615 908 1287 1765 1.04
Whitefish (1996) 3.32 1.29 360 562 830 1178 1617 0.99
des Mille Lacs (1999) 3.32 1.44 402 627 927 1315 1805 1.09
Expanse 3.33 1.30 385 601 889 1263 1735 1.07
Despair 3.33 1.26 373 582 862 1224 1682 1.00
Broad 3.34 1.24 390 609 902 1282 1763 1.06
Whitefish (1995) 3.35 1.09 363 568 843 1199 1651 0.98
Wapesi 3.35 1.07 356 558 827 1177 1620 0.98
Ball (1998) 3.35 1.32 440 688 1021 1453 1999 1.21
Seul (1997) 3.35 1.12 373 584 866 1232 1696 1.06
Jackfish 3.36 1.12 396 620 920 1311 1806 1.10
Dimple 3.38 0.90 357 561 836 1193 1647 0.97
Little Sandford 3.38 0.89 353 555 827 1180 1629 0.94
Makokibatan 3.39 0.85 358 563 839 1199 1657 1.01
Icarus 3.40 0.91 406 640 955 1366 1889 1.13
Young 3.42 0.74 371 587 877 1258 1743 1.07
Dog 3.43 0.86 458 724 1084 1556 2157 1.28
Devious 3.44 0.64 361 572 857 1232 1710 0.94
Rainy (1997) 3.46 0.62 393 624 939 1352 1880 1.17
Ara 3.48 0.46 328 522 787 1135 1582 0.96
Georgia 3.50 0.45 361 576 870 1258 1756 0.97
Kenorain 3.61 0.23 351 569 871 1274 1797 1.13
Agimak 3.67 0.16 347 567 874 1286 1825 0.92
Sturgeon 4.04 0.01 190 325 524 801 1178 0.90
1. Two-tailed Student t-tests were employed to test significance of slope and intercept ( o: or = 0). 2. Condition calculated as the average of the individual ratios of observed weight-to-predicted weight from northwest region
median regression model (see below).
NW - 22
Northwest region female walleye weight-at-total length benchmarks.
Weight-length relation Calculated weight at standard total lengths
x10-6 350 400 450 500 550
Mean 3.20 5.10 382 585 853 1195 1621
Minimum 2.71 0.01 190 325 524 801 1178
Maximum 4.04 55.41 568 846 1204 1649 2193
Median 3.21 2.55 375 575 844 1178 1598
5% Percentile 2.89 0.64 346 530 763 1057 1398
25% Quartile 3.12 1.53 360 555 809 1134 1534
75% Quartile 3.29 4.47 394 604 879 1247 1701
95% Percentile 3.44 18.45 442 669 975 1371 1868
Sample Size 123
NW - 23
Table NW-4b. Predicted male walleye weight-at-total length for northwest region waterbodies sampled using the fall walleye index netting standard. A geometric mean predictive regression model was fitted to logarithmically transformed (base 10) individual
observations (only lakes with 30 fish sampled). Weight in grams and total length in millimetres.
Weight-length relation1. Calculated weight at standard total lengthsLake
x10-6 350 400 450 500 550 Condition2.
Favourable 2.63 87.12 428 607 828 1092 1404 1.08
Little Trout 2.81 31.78 448 652 907 1220 1594 1.21
Savanne (2001) 2.89 16.78 378 556 781 1059 1395 1.02
Finlayson 2.93 14.06 400 592 835 1138 1504 1.14
Whitefish (1995) 2.96 11.04 374 556 788 1076 1427 1.03
Eltrut 2.97 10.30 370 551 781 1069 1418 1.05
Partridge 2.98 9.96 380 565 803 1099 1461 1.05
Rawn Reservoir 3.00 8.55 367 547 779 1069 1423 1.05
Perch 3.00 8.62 370 552 785 1078 1434 1.05
Sydney 3.01 8.86 403 602 858 1179 1570 1.07
Northern Light 3.01 10.70 486 727 1036 1423 1896 1.28
Zizania 3.02 8.08 389 583 832 1144 1525 1.03
McAree 3.03 7.55 386 578 826 1137 1518 1.08
Henderson 3.03 7.33 375 561 802 1104 1474 0.99
Calm 3.03 7.78 398 596 852 1172 1564 1.08
Kilburn 3.04 6.90 374 561 803 1106 1478 1.02
Wawang 3.04 8.24 447 670 959 1321 1765 1.18
White Otter 3.05 6.35 365 548 785 1083 1448 1.05
Cuttle 3.06 6.42 391 589 844 1165 1560 1.08
Marmion 3.07 5.98 386 582 836 1155 1547 1.09
Devious 3.07 4.93 319 480 689 952 1276 0.89
Seul (1996) 3.09 5.93 431 651 936 1297 1741 1.16
Dovetail 3.09 5.36 389 588 846 1172 1574 1.06
Three Finger 3.10 4.60 354 536 772 1070 1438 0.98
Pekagoning (1998) 3.12 3.90 338 512 740 1028 1384 0.93
Whalen 3.12 4.52 391 594 857 1191 1604 1.04
Steel 3.12 4.07 352 535 772 1072 1444 0.95
Nipigon (1997) 3.12 4.72 409 620 895 1244 1674 1.13
Savanne (1997) 3.12 4.45 385 585 844 1173 1579 1.04
Smoothrock 3.13 4.01 368 559 809 1124 1515 1.02
Mount 3.13 4.00 367 558 807 1122 1511 0.98
Pekagoning (1999) 3.13 4.10 376 572 827 1150 1549 1.01
Nipigon (2001) (GB) 3.14 4.42 430 654 947 1319 1779 1.18
Nipigon (2000) 3.15 3.82 394 601 870 1213 1638 1.11
Whitewater 3.15 3.63 375 571 827 1153 1556 1.03
Minnitaki 3.16 3.40 372 568 823 1149 1552 1.04
Miminiska 3.16 3.45 378 576 836 1166 1575 0.98
Garden 3.16 3.45 378 576 836 1166 1575 1.02
Klotz 3.17 3.78 439 670 973 1359 1838 1.18
Straw 3.18 3.13 385 589 857 1197 1621 1.07
Iron Range 3.18 3.08 379 580 843 1178 1596 1.03
NW - 24
Table NW-4b. (continued)
Weight-length relation1. Calculated weight at standard total lengthsLake
x10-6 350 400 450 500 550 Condition2.
Wolseley 3.18 3.20 394 602 876 1224 1658 1.08
Abram 3.18 2.99 368 563 818 1144 1549 0.99
Hutchison 3.19 2.93 382 585 852 1193 1617 1.07
Namakan 3.19 2.87 375 573 835 1168 1584 1.01
Kapkichi 3.19 2.89 377 577 841 1177 1595 1.04
Longlegged 3.19 2.94 384 587 855 1197 1622 1.06
Wildgoose 3.20 2.59 358 549 801 1122 1522 0.97
Red 3.20 2.77 383 588 857 1200 1628 1.04
Whitefish (2001) 3.21 2.55 374 574 838 1176 1596 0.99
of the Woods (2000) 3.21 2.61 383 588 858 1203 1634 1.04
Onaman 3.21 2.90 425 653 953 1337 1815 1.16
Meta 3.21 2.60 381 586 855 1199 1628 1.03
Mercutio 3.21 2.59 380 583 851 1194 1621 1.02
Botsford 3.22 2.27 353 543 793 1114 1514 0.98
Harris 3.22 2.29 356 548 800 1123 1527 0.99
Nipigon (1999) 3.22 2.73 425 653 954 1339 1820 1.16
Factor 3.23 2.19 361 556 813 1143 1555 1.00
Neston 3.23 2.35 388 597 873 1227 1669 1.04
Savanne (1996) 3.23 2.26 373 574 839 1180 1605 1.03
Burrows 3.23 2.41 398 612 895 1258 1712 1.05
Lower Companion 3.24 2.23 390 601 880 1239 1687 1.08
Wabigoon 3.24 2.08 364 561 821 1155 1573 1.01
Pelican 3.24 2.05 359 553 809 1139 1551 1.01
Pickwick 3.24 2.05 359 553 809 1139 1551 0.97
Trout 3.24 2.06 360 555 813 1144 1558 1.00
Pipestone 3.24 2.29 401 617 904 1272 1732 1.12
Sakwite 3.25 2.00 371 572 839 1182 1611 1.01
Geikie 3.26 1.93 380 587 861 1214 1656 1.04
des Mille Lacs (1994) 3.26 1.96 385 596 874 1233 1682 1.07
Shoal (1998) 3.26 1.95 383 593 870 1227 1673 1.04
Churchill 3.28 1.69 374 579 852 1204 1645 1.02
Kukukus 3.28 1.74 385 596 877 1239 1694 1.05
Sapawe 3.28 1.80 398 617 907 1282 1753 1.07
Kaopskikamak 3.28 1.77 391 606 892 1261 1723 1.04
Jutten 3.28 1.66 367 569 837 1182 1616 0.98
Footprint 3.28 1.82 402 623 918 1296 1772 1.08
Gamsby 3.28 1.65 365 565 832 1175 1606 0.96
Makokibatan 3.28 1.74 385 596 877 1239 1694 1.02
Hicks 3.28 1.65 365 565 832 1175 1606 0.98
Pettit 3.29 1.58 370 575 847 1197 1639 1.01
Shoal (2001) 3.29 1.72 403 626 922 1304 1784 1.13
Eagle 3.29 1.60 375 582 857 1213 1659 1.04
Wintering 3.30 1.48 368 572 843 1194 1635 0.98
St. Joseph 3.30 1.50 373 579 854 1210 1657 1.02
NW - 25
Table NW-4b. (continued)
Weight-length relation1. Calculated weight at standard total lengthsLake
x10-6 350 400 450 500 550 Condition2.
des Mille Lacs (1999) 3.31 1.50 395 615 908 1287 1765 1.11
Gammon 3.31 1.45 382 595 878 1244 1706 1.09
Rainy (1999) 3.31 1.45 382 595 878 1244 1706 1.05
Toronto 3.32 1.34 374 583 863 1224 1679 1.00
Elva 3.34 1.12 352 550 815 1158 1592 0.95
Otukamamoan 3.34 1.13 355 555 822 1169 1607 0.97
Obonga 3.34 1.16 364 569 844 1200 1649 0.99
Ice 3.34 1.26 396 618 916 1303 1791 1.10
Brennan 3.35 1.10 366 573 850 1210 1666 0.98
Seul (1997) 3.37 1.05 393 617 917 1308 1804 1.08
Winnipeg River (1998) 3.37 1.01 378 593 882 1258 1735 1.00
Little Sandford 3.38 0.88 349 549 817 1167 1610 0.94
Keikewabik 3.38 0.87 346 543 808 1154 1592 0.96
des Mille Lacs (1993) 3.39 0.94 396 622 928 1326 1832 1.08
Bending 3.41 0.78 369 582 870 1246 1725 0.97
Watcomb 3.43 0.68 362 572 857 1230 1706 1.04
Lount (1998) 3.43 0.77 410 648 971 1393 1932 1.10
Expanse 3.44 0.71 401 634 951 1367 1897 1.10
Rainy (1998) 3.45 0.58 347 550 826 1188 1651 0.94
Broad 3.45 0.63 377 598 897 1291 1793 1.12
Jones 3.48 0.54 385 613 924 1333 1857 1.04
Whitefish (1996) 3.48 0.52 371 590 890 1284 1788 1.01
Wapesi 3.51 0.45 383 612 925 1338 1870 1.03
Young 3.53 0.40 382 613 929 1347 1886 1.10
Kenorain 3.54 0.38 385 618 938 1362 1908 1.04
Icarus 3.64 0.23 419 681 1046 1535 2171 1.17
Sturgeon 3.99 0.02 283 482 772 1175 1718 0.91
1. Two-tailed Student t-tests were employed to test significance of slope and intercept ( o: or = 0). 2. Condition calculated as the average of the individual ratios of observed weight-to-predicted weight from northwest region
median regression model (see below).
Northwest region male walleye weight-at-total length benchmarks.
Weight-length relation Calculated weight at standard total lengths
x10-6 350 400 450 500 550
Mean 3.22 4.37 381 586 856 1203 1636
Minimum 2.63 0.02 283 480 689 952 1276
Maximum 3.99 87.12 486 727 1046 1535 2171
Median 3.22 2.38 380 583 850 1194 1621
5% Percentile 2.97 0.52 349 543 779 1070 1423
25% Quartile 3.12 1.50 368 562 822 1150 1554
75% Quartile 3.30 4.26 391 602 886 1245 1715
95% Percentile 3.48 10.70 430 653 954 1359 1886
Sample Size 112
NW - 26
Table NW-5a. Average female walleye total length-at-age (millimetres) data for northwest region waterbodies sampled using the fall walleye index netting method. Only
mean values with a minimum sample size 4 are included.
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Alph 516
Kenorain 453 600 558 606
Lac la Croix 467
Cloud 622
Whitewater 328 402 417 436 454
Savanne (2001) 356 358 420 426 457
Minn 501
Whiterock 496
Broad 451 439 484 605
Savanne (1996) 370 390 457 467 481
Klotz 491 523 553 578
Favourable 432 448 474 491
Sturgeon 426 480
Boffin 504 557
Miminiska 277 308 350 394 408 471
Makokibatan 284 347 409 492 512 539
Marmion 294 301 370 455 570
Toronto 312 356 383 419 441 482
Meta 317 444 462
Old Man 320 373 471
Savanne (1997) 324 342 398 405 457 468 476
Ara 338 343 390 459 450 463 491
Patricia 345 393 476
Manomin 347 374
Rugby 356 388 407 430 461
Onaman 378 424 476 501 513
Sandstone 392 449
Union 395
Lount (1998) 401 454
Husband 409
Tyrell 412 535
Wolseley 437
Bar 459
Robinson 468 517
Neston 195 263 315 372 417 460 470
Geikie 208 270 327 411
Jutten 215 283 369 406 433 477 532
des Mille Lacs (1993) 218 266 321 383 454 479 529 605
Harris 230 293 509
Abram 239 297 370 438 479
Rainy (1998) 239 304 346
Jones 247 339 359 400
Botsford 248 307 373 421 481 500 522
NW - 27
Table NW-5a. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
McAree 250 336 477 550
Perch 251 272 333 335
Brennan 251 282 309 366 389 437
Sakwite 253 312 368 407 420 493
Smoothrock 254 297 354 399 420 456 493
Kapkichi 254 301 336 371 427 473
Wapesi 255 310 352 384 431 452 465
Hicks 256 299 340 481
Pelican 258 306 364 424 460 607
Wabigoon 263 336 389 436 546
Steel 271 298 354 403 418 460
Namakan 272 316 357 420 431
Dovetail 274 364 440 523 544
Partridge 281 355 370 402
Sand Point 283 394 442 502
Three Finger 284 327 363
Expanse 284 342 393 433 505 515 524 552
Keikewabik 285 326 354 391 423 461
Rainy (1999) 290 345 418 447 539
Bending 294 338 335 377 447
Kukukus 297 372 421 462 513 610 618
Chill 304
Ball (1998) 306 378 435 478 579
Cuttle 309 340 387
Jackfish 321 384 460 532 568
Kaopskikamak 321 428
Watcomb 323 358 379 469 499
Despair 326 370 417 480 517
Sucan 326 375 450
Zizania 328 371 441 472
Footprint 329 394 518
Whalen 332
Elva 332 414 503
Richardson 335 563
Dog 337 397 467 545 553 637
Darby 341 454
Manion 342 427
Gammon 354 585
Ice 362 422
Rainy (1997) 381 479 513 529 542 573 609
Pipestone 407 478 543 660
Red 183 264 326 370 413 481 516 553 572
Garden 187 241 290 304 351 360 397 453
des Mille Lacs (1994) 189 274 314 356 442
Wintering 189 273 325 370 390 473
NW - 28
Table NW-5a. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Sapawe 190 265 327 401
Minnitaki 192 255 316 375 445 495 528 593
Kilburn 192 266 286 351 428 458 551
Calm 193 245 304 364 399 433
Burrows 194 257 321 343 387 435 484
Mercutio 197 343 379 480 504
Finlayson 198 281 345 354 399 446
Rawn Reservoir 199 285 368
Beak 205 325
Eltrut 210 256 353 385
St. Joseph 211 268 325 373 399 427 475 497 508
Little Sandford 212 282 369 418 467
Sydney 214 277 350 387 462 483
Gamsby 214 294 353 404 435
Nipigon (2000) 216 245 362 430
des Mille Lacs (1999) 216 265 32 381 435 453 532 556 580
Icarus 216 311 471 544 650
Eagle 217 309 393 454 492 543 497 588 540
Factor 220 336 438
Georgia 221 280 343 388 421 423 489
Seul (1997) 223 292 355 393 436 490 507 518 559 548
Mainville 224 318 351 332
Churchill 227 290 341 362 345 453 482 498 502
Whitefish (1996) 228 316 451 476 593
Winnipeg River (1998) 229 281 334 376 391 367 469 505 528 530
Little Trout 229 291 350 394 503 509 574
Dimple 229 309 462
Nipigon (2001) (CB) 230 390
Kenogamisis 231 268 319 356 397 452 400
Devious 231 306 393
Otter 232 298 384 429 534
Obonga 232 307 381 425 485 563
Keckush 235
Seul (1995) 237 305 356 408 425 474 474 517 543 549
of the Woods (2000) 237 308 367 418 488 558
White Otter 237 315 384
Agimak 238 322 335 352
Nipigon (2001) (GB) 238 326 434 473
Nipigon (1997) 240 338 405 451
Lower Companion 241 369
Trout 244 309 361 445 566
Pekagoning (1998) 244 312 384 433 575
Northern Light 249 330 415 563 586
Whitefish (2001) 250 323 390 439 471 502 544 601
Mount 250 328 418 483 474
NW - 29
Table NW-5a. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Nipigon (1998) 250 332 397 445
Straw 251 312 360 408 434
Pettit 251 360 427 476 491 542
Otukamamoan 252 308 383 442 467 555 532 564
Steep Rock 253
Hutchison 254 346 486 572
Whitefish (1995) 255 410 437 580 612
Longlegged 255 309 370 488 524 564 624
Pickwick 256 333 382 460
Elbow 261
Kawawiag 262 478 641
Pekagoning (1999) 263 333 387 439 499 611
Henderson 268 437 488 490 521
Nipigon (1999) 273 345 400 488 502
Iron Range 275 440
Wildgoose 279 343 390
Wababimiga 280
Upper Companion 282 365
Shoal (2001) 282 403 503 549 587 656 631 639
Burditt 294 368 451 513 557
Young 296 371 503 610
Wawang 305 471 522 611 661
Shoal (1998) 310 395 528
Crooked Pine 312 545
Northwest region female walleye total length-at-age benchmarks.
Average total length (mm) at age
1 2 3 4 5 6 7 8 9 10
Mean 237 298 360 404 431 469 490 527 534 542
Minimum 183 195 263 301 332 360 397 400 436 454
Maximum 312 407 503 549 587 600 656 661 660 641
Median 235 301 356 391 434 473 484 522 529 548
5% Percentile 189 239 277 315 351 385 405 417 437 457
25% Quartile 216 265 319 355 390 420 452 470 474 491
75% Quartile 254 328 393 444 472 504 528 580 579 580
95% Percentile 296 369 471 522 523 568 578 650 639 624
Sample Size 73 108 118 112 88 59 51 35 33 35
NW - 30
Table NW-5b. Average male walleye total length-at-age (millimetres) data for northwest region waterbodies sampled using the fall walleye index netting method. Only
mean values with a minimum sample size 4 are included.
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Savanne (2001) 348 377 390 389 415
Cloud 544
Brewer 394
Klotz 454 500 510
Whiterock 471
Jones 344 380
Dore 460
Bar 499
Henderson 409 454 474 497
Alph 488
Savanne (1996) 367 376 421 447
Miminiska 246 314 327 388 423
Perch 271 298 357
Whitewater 281 316 356 388 383 411 423
Wapesi 284 356 367 382 407 403 413
Savanne (1997) 312 375 382 410 412 416
Agimak 314 321 346
Keikewabik 321 345 374 397 411
Three Finger 321 337 373 384
Toronto 322 329 366 387
Patricia 322 375
Ara 326 360 382 417 409 420 427
Expanse 330 382 409 446 455 496
Manomin 330
Kenorain 333 380 473
Watcomb 347 383 433
Ball (1998) 367 428 439 503 525 496
Lac la Croix 371 377 431
Despair 374 442
Sturgeon 375 416 426
Union 375
Broad 409 413 419 460 508 518
Bradshaw 442
Neston 207 255 323 378 418 420
Marmion 210 282 328 372
Abram 218 295 408 403 454
Jutten 219 280 344 395 402 424
Geikie 223 287 335
des Mille Lacs (1994) 225 266 319 351 396 406 405
Makokibatan 227 273 342 389
Pelican 235 291 370 410 454
Harris 237 293 412 455 478
Brennan 241 268 299 368 379 376 393
NW - 31
Table NW-5b. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Eltrut 245 328 342 363
Smoothrock 245 292 355 382 400 424 462
Rainy (1998) 247 298 363
Nipigon (2000) 248 354 407
Botsford 249 304 362 397 430
Kapkichi 249 305 338 400 416
Kilburn 251 301 346 389 385 456
Sakwite 252 309 343 380 399
Minnitaki 256 309 362 414 454 464 520
McAree 259 288 344 383 423 444
Steel 260 284 367 371 402 393 431
Meta 266 349 423 428 414
Dovetail 272 353 417 456
Sydney 276 341 394 418 437 402
des Mille Lacs (1999) 278 319 380 393 371 376 429 408 444
Bending 284 325 308 417
Rainy (1999) 287 344 388 442 474
Namakan 288 275 350 371 434
Otukamamoan 294 377 401 455
Kukukus 298 357 395 441 462 472 496 507 482
Sand Point 302 371 439 455
Little Turtle 302 359
Dimple 304 446
Rugby 305 332 375 351 391 406
Sandstone 307 382 418
Lount (1998) 308 370 401
Kaopskikamak 308 394
Cuttle 319 340 366
Nipigon (2001) (CB) 322 484
Whalen 325 380
Zizania 328 365 406 433 468
Onaman 330 382 413 442 454 475 476
Husband 331 400
Jackfish 332 386
Darby 335 377
Dog 338 386 434 481 478
Manion 342 405
Footprint 344 363
Ice 357 406
Rainy (1997) 357 461 468 526 515 587
Minn 364 415 469
Gammon 366 473 534
Wolseley 368 436
Robinson 380 443 483 515
Tyrell 382
NW - 32
Table NW-5b. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Vista 392
Pipestone 399 451 531
des Mille Lacs (1993) 158 231 260 311 381 443 480 472
Hicks 188 249 317 342
Garden 189 239 279 312 357 367 406
Red 189 261 321 368 394 421 450 507 508
Sapawe 190 274 317 375 408
Old Man 192 328 373
Wintering 193 266 323 370 376 399 421 468
Finlayson 196 284 339 432
Favourable 198 321 365 385 415 410 434 470
Burrows 198 253 316 335 352 369
Partridge 201 280 346 359 383
Calm 202 243 297 376
Mercutio 204 342 379 473 455
Rawn Reservoir 205 289 354
St. Joseph 206 268 318 372 389 391 439 445 466
Wabigoon 210 268 323 373 392 478 495
Little Sandford 210 280 359 400 427 449
Gamsby 212 298 345 391 393 418
Factor 215 307 415 478 494 500 527
Little Trout 217 291 361 386 404 469 472 486 487
Eagle 219 301 373 424 444 476 527
Kenogamisis 220 275 311 351 341 384 405
Beak 222
Georgia 227 292 372
Whitefish (1996) 227 309 402 485
Icarus 228 302 437 494 564
Mainville 229 284 307 341 455
Winnipeg River (1998) 230 282 330 368 464 457 465
Trout 230 315 359 415 501
Churchill 231 289 343 367 363 413 425 426 439
Seul (1996) 233 303 346 374 410 432 428 460 450 475
Obonga 235 291 366 400 452
Nipigon (1997) 235 317 387 373
White Otter 236 307 380
Devious 236 308 358
of the Woods (2000) 236 309 362 401 488
Keckush 241
Steep Rock 241 416
Richardson 241 331 479
Seul (1997) 242 315 374 413 417 448 465 483 488 499
Elva 242 315 405
Straw 244 315 369 393 413
NW - 33
Table NW-5b. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Otter 245 313 366
Pekagoning (1998) 246 309 372 405
Elbow 247
Nipigon (2001) (GB) 247 326 433 451
Whitefish (2001) 247 327 383 417 433 449 465 485
Nipigon (1998) 247 343
Lower Companion 247 365 424
Sucan 248 333 360
Longlegged 252 313 374 398 407 424 456 465 484
Pickwick 256 331 377 421 441
Hutchison 256 352 505
Northern Light 257 348 407
Pettit 259 357 432
Kawawiag 260
Pekagoning (1999) 261 333 391 429
Whitefish (1995) 262 385 409 454
Mount 267 345 392 440 442 474
Wababimiga 272
Wildgoose 272 340 372 394
Iron Range 273 296 408
Nipigon (1999) 273 336 403 430 442 476
Shoal (2001) 278 391 477 518 553 568
Burditt 288 380 428 457 498 527
Young 290 375 425 471 519
Shoal (1998) 306 396 485
Wawang 307 386 489 540 551
Crooked Pine 309 471 576
Northwest region male walleye total length-at-age benchmarks.
Average total length (mm) at age
1 2 3 4 5 6 7 8 9 10
Mean 236 301 350 388 402 432 442 462 457 461
Minimum 158 207 246 298 327 357 367 383 393 384
Maximum 309 399 477 518 553 534 568 587 527 527
Median 236 303 353 380 394 432 444 455 465 467
5% Percentile 189 225 273 314 344 369 376 405 403 413
25% Quartile 212 266 314 350 372 391 402 411 420 425
75% Quartile 256 332 380 416 427 468 469 483 486 492
95% Percentile 290 382 437 484 478 515 527 576 507 520
Sample Size 69 115 123 114 74 58 49 33 25 32
NW - 34
Table NW-6a. Average female walleye weight-at-age (grams) data for northwest region waterbodies sampled using the fall walleye index netting method. Only mean
values with a minimum sample size 4 are included.
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Alph 1655
Sturgeon 648 1029
Klotz 1248 1516 1856 2123
Cloud 3193
Broad 847 939 1224 2471
Whitewater 326 610 729 832 917
Savanne (2001) 389 448 656 691 811
Kenorain 1017 2405 1850 2682
Minn 1145
Lac la Croix 1097
Savanne (1996) 460 562 898 934 1004
Favourable 792 833 1026 1061
Whiterock 1170
Boffin 1335 1901
Miminiska 177 254 376 532 619 891
Makokibatan 189 383 620 1125 1377 1655
Toronto 238 398 479 674 759 1040
Marmion 242 256 471 920 1890
Old Man 280 420 1006
Ara 290 319 470 870 815 943 1037
Meta 292 750 937
Savanne (1997) 293 383 573 613 828 927 944
Patricia 334 486 947
Manomin 347 428
Rugby 438 569 688 900 1045
Sandstone 443 908
Union 545
Husband 623
Lount (1998) 648 1000
Tyrell 656 1513
Onaman 664 816 1235 1361 1485
Wolseley 810
Robinson 907 1303
Bar 1026
Neston 60 153 274 438 645 882 1031
Geikie 73 166 340 637
Jutten 77 181 468 584 738 1008 1493
des Mille Lacs (1993) 91 172 333 514 1007 1151 1597 2475
Rainy (1998) 106 233 338
Harris 108 208 1203
Abram 109 211 429 772 1046
Brennan 119 184 235 410 517 753
Jones 121 371 439 625
NW - 35
Table NW-6a. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Botsford 126 241 461 662 1093 1208 1359
Wapesi 128 251 389 509 757 899 984
Hicks 130 218 335 1000
Sakwite 132 254 425 618 602 1076
Smoothrock 136 222 384 579 682 903 1162
Kapkichi 136 232 334 459 721 1048
McAree 138 340 1245 1730
Perch 138 174 312 348
Wabigoon 149 347 539 743 1475
Steel 159 210 361 549 617 868
Namakan 167 294 431 673 774
Pelican 173 246 422 722 917 2168
Keikewabik 185 288 362 498 641 858
Three Finger 190 287 407
Sand Point 191 608 811 1264
Dovetail 192 446 780 1309 1451
Expanse 204 373 556 788 1370 1467 1438 1746
Partridge 204 407 459 594
Bending 211 343 303 439 824
Kukukus 219 442 680 972 1302 2228 2302
Chill 224
Rainy (1999) 230 380 753 909 1531
Sucan 264 459 816
Cuttle 268 323 543
Ball (1998) 274 589 945 1261 2386
Watcomb 282 409 520 943 1163
Despair 291 457 688 1056 1340
Elva 299 638 1345
Kaopskikamak 304 744
Jackfish 305 523 1079 1605 1985
Richardson 312 2050
Zizania 312 463 827 1047
Footprint 317 583 1381
Whalen 318
Darby 334 936
Manion 364 717
Dog 374 705 1363 2177 2350 3180
Gammon 421 2073
Ice 433 746
Rainy (1997) 565 1239 1484 1801 1873 2305 2851
Pipestone 655 1106 1638 3122
Red 47 163 308 463 669 1126 1486 1751 1941
Garden 48 113 220 254 410 403 664 895
Wintering 49 159 286 429 514 973
Burrows 57 142 292 372 537 801 1205
NW - 36
Table NW-6a. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Mercutio 58 365 495 1041 1211
Sapawe 58 158 311 637
Minnitaki 59 137 280 471 812 1213 1432 2193
des Mille Lacs (1994) 60 177 280 414 940
Calm 62 160 270 442 573 878
Little Sandford 65 184 429 663 928
Rawn Reservoir 67 200 432
Beak 68 303
Georgia 71 171 373 52 652 655 1134
Eltrut 73 143 384 512
Gamsby 73 200 358 559 697
St. Joseph 77 162 290 437 553 669 1016 1234 1267
Finlayson 77 190 381 392 580 838
Agimak 79 305 318 355
Icarus 80 292 1218 1817 3600
Factor 80 352 760
des Mille Lacs (1999) 85 173 395 536 827 846 1720 1863 2382
Eagle 85 254 540 894 1161 1663 1393 2238 1814
Sydney 90 193 404 555 920 1171
Dimple 90 248 890
Whitefish (1996) 91 262 865 1175 2298
Nipigon (2000) 93 135 454 799
Mainville 94 304 384 338
Devious 94 210 658
Seul 1997 95 217 412 620 809 1203 1401 1536 1919 1826
Winnipeg River (1998) 99 191 333 471 524 423 953 1200 1553 1488
Kilburn 100 154 204 383 747 914 1492
Obonga 100 237 485 694 1106 1706
Churchill 101 189 325 423 371 873 1006 1184 1185
of the Woods (2000) 110 260 446 712 1194 1792
Trout 111 245 396 780 1785
Keckush 114
Nipigon (2001) (CB) 115 522
Otukamamoan 117 231 472 799 915 1512 1358 1735
Little Trout 118 245 395 547 1300 1214 1788
Pekagoning (1998) 120 244 448 696 1762
White Otter 120 274 524
Lower Companion 125 465
Mount 127 303 665 1014 973
Nipigon (2001) (GB) 129 344 849 1210
Pettit 129 413 709 1092 1236 1520
Whitefish (1995) 130 610 758 2045 2438
Pickwick 130 307 475 900
Nipigon (1997) 130 379 661 934
Longlegged 132 273 438 1089 1391 1775 2350
NW - 37
Table NW-6a. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Whitefish (2001) 132 275 519 756 975 1163 1527 2149
Seul (1996) 136 294 465 685 819 1154 1217 1517 1802 1643
Kawawiag 140 1019 2488
Kenogamisis 140 218 321 441 615 919 650
Straw 141 274 436 618 786
Nipigon (1998) 148 362 638 932
Hutchison 149 365 1170 2000
Henderson 152 729 1051 1089 1200
Pekagoning (1999) 158 325 519 793 1175 2189
Wildgoose 176 337 485
Iron Range 179 780
Northern Light 184 413 799 2055 2444
Burditt 194 434 799 1235 1714
Wababimiga 198
Nipigon (1999) 198 411 682 1264 1426
Upper Companion 200 455
Steep Rock 211
Shoal (2001) 211 651 1401 1829 2399 3312 2925 3421
Otter 216 421 477 725 1434
Elbow 225
Young 225 440 1325 2613
Shoal (1998) 250 555 1454
Crooked Pine 256 1738
Wawang 272 1031 1708 2614 3225
Northwest region female walleye weight-at-age benchmarks.
Average weight (g) at age
1 2 3 4 5 6 7 8 9 10
Mean 123 252 462 684 810 1072 1250 1620 1691 1633
Minimum 47 60 153 235 338 403 610 650 753 811
Maximum 272 655 1401 1829 2399 2405 3312 3600 3421 2488
Median 115 230 410 551 765 1007 1203 1434 1597 1655
5% Percentile 57 108 177 280 376 498 619 729 832 891
25% Quartile 80 160 288 390 530 669 870 1000 1040 1134
75% Quartile 148 312 583 837 996 1302 1451 2045 2228 2149
95% Percentile 225 455 1026 1638 1513 2073 2350 3225 3180 2475
Sample Size 73 108 118 112 88 59 51 35 33 35
NW - 38
Table NW-6b. Average male walleye weight-at-age (grams) data for northwest region waterbodies sampled using the fall walleye index netting method. Only mean
values with a minimum sample size 4 are included.
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Klotz 1003 1354 1523
Brewer 677
Cloud 2201
Whiterock 1025
Savanne (2001) 392 500 547 550 651
Jones 371 504
Alph 1332
Savanne (1996) 455 485 694 824
Henderson 607 888 959 1129
Dore 1063
Bar 1341
Miminiska 114 299 296 503 678
Wapesi 173 416 439 509 618 608 664
Perch 180 244 404
Whitewater 199 279 438 541 494 668 719
Savanne (1997) 232 509 517 650 655 682
Agimak 250 260 364
Ara 258 383 454 605 558 625 669
Keikewabik 266 350 436 542 662
Three Finger 274 327 428 505
Patricia 275 454
Toronto 282 309 437 534
Manomin 289
Kenorain 320 506 1150
Expanse 345 536 697 917 972 1342
Watcomb 350 517 779
Sturgeon 430 604 642
Union 475
Despair 487 816
Ball (1998) 512 901 1555 1731 1925 1607
Lac la Croix 623 643 893
Broad 697 755 724 1080 1293 1671
Bradshaw 814
Neston 72 139 298 488 701 637
Abram 77 230 593 590 882
Jutten 81 182 348 534 596 696
des Mille Lacs (1994) 84 159 299 426 606 688 688
Makokibatan 94 176 396 543
Harris 104 208 547 842 1031
Pelican 105 198 456 604 939
Marmion 105 204 304 476
Brennan 110 147 212 429 464 479 516
Geikie 111 242 329
NW - 39
Table NW-6b. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Rainy (1998) 119 217 392
Botsford 122 238 415 546 743
Eltrut 128 299 350 441
Kapkichi 132 255 348 588 695
Smoothrock 135 214 399 475 560 694 871
Sakwite 137 252 351 489 591
Minnitaki 139 264 447 687 894 941 1399
Nipigon (2000) 139 430 694
Kilburn 143 226 359 577 531 848
Steel 149 179 370 405 566 505 690
Meta 153 404 723 700 657
McAree 167 216 376 580 746 901
Dovetail 188 394 663 952
Bending 191 279 223 698
Sydney 195 361 586 715 763 713
Otukamamoan 212 470 566 920
Namakan 223 175 396 443 727
Kukukus 223 402 573 833 949 1070 1191 1363 1143
des Mille Lacs (1999) 225 310 533 600 489 501 856 726 917
Dimple 228 802
Rainy (1999) 229 389 604 878 1037
Little Turtle 249 398
Sand Point 260 481 1348 905
Kaopskikamak 264 581
Nipigon (2001) (CB) 273 1114
Lount (1998) 274 466 648
Sandstone 278 541 754
Cuttle 285 368 461
Whalen 297 500
Husband 310 618
Zizania 321 439 577 794 975
Rugby 322 456 544 525 600 684
Darby 330 520
Onaman 343 605 752 986 1041 1141 1153
Jackfish 357 544
Manion 364 614
Dog 365 613 895 1469 1522
Footprint 381 440
Ice 420 665
Rainy (1997) 435 1062 1122 1786 1596 2435
Gammon 454 1069 1608
Wolseley 468 810
Minn 502 760 783
Tyrell 530
Vista 550
NW - 40
Table NW-6b. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Robinson 590 865 985 1250
Pipestone 631 928 1795
des Mille Lacs (1993) 26 99 163 285 519 862 1114 1075
Hicks 51 123 258 327
Wintering 52 146 286 423 473 580 735 1020
Garden 54 117 185 250 417 450 593
Sapawe 55 178 280 497 785
Georgia 58 205 445
Old Man 60 290 462
Burrows 63 134 280 332 407 498
Red 65 154 300 466 558 724 905 1223 1337
Little Sandford 65 184 390 555 692 801
St. Joseph 69 154 265 446 532 547 803 860 980
Mercutio 71 352 491 1020 880
Calm 71 146 256 495
Gamsby 71 212 345 510 526 670
Finlayson 75 229 363 758
Wabigoon 78 163 310 472 566 1022 1224
Partridge 79 200 382 407 512
Factor 79 237 652 1022 1120 1191 1408
Eagle 79 240 467 729 847 1103 1559
Whitefish (1996) 86 240 593 1200
Rawn Reservoir 90 208 397
Beak 93
Trout 94 263 390 628 1240
Icarus 95 248 949 1529 2278
Devious 97 209 347
Winnipeg River (1998) 99 186 326 463 1058 880 1023
Churchill 99 194 338 435 401 665 715 684 814
Favourable 100 332 478 521 679 693 787 965
Richardson 100 298 1194
Mainville 103 159 248 353 842
Obonga 105 200 434 574 908
Elva 108 244 573
of the Woods (2000) 111 261 432 596 1141
White Otter 115 254 490
Seul (1997) 116 275 515 675 728 915 1095 1186 1239 1438
Otter 117 230 428
Nipigon (1997) 119 306 578 514
Keckush 120
Pekagoning (1998) 120 236 407 575
Sucan 122 592 417
Whitefish (2001) 124 289 504 635 753 844 942 1074
Straw 127 279 432 565 735
Longlegged 129 314 470 579 625 703 941 1002 1095
NW - 41
Table NW-6b. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Pickwick 130 309 467 663 743
Seul (1996) 131 285 446 488 692 876 839 1100 1115 1073
Lower Companion 134 453 800
Hutchison 138 393 1328
Nipigon (1998) 141 381
Pettit 141 406 735
Whitefish (1995) 143 493 627 888
Nipigon (2001) (GB) 145 347 857 1026
Kawawiag 148
Little Trout 153 227 434 541 600 1021 1050 1204 1144
Pekagoning (1999) 153 322 530 713
Wildgoose 157 331 439 535
Kenogamisis 168 207 306 435 406 563 669
Iron Range 172 235 630
Wababimiga 178
Mount 189 360 512 762 758 932
Shoal (2001) 194 627 1177 1544 1968 1815
Nipigon (1999) 197 384 669 870 917 1168
Elbow 198
Young 207 446 766 1107 1460
Northern Light 209 464 758
Burditt 210 430 679 846 1216 1515
Steep Rock 217 883
Shoal (1998) 243 555 1125
Crooked Pine 253 1088 1920
Wawang 279 573 1242 1625 1756
Northwest region male walleye weight-at-age benchmarks.
Average weight (g) at age
1 2 3 4 5 6 7 8 9 10
Mean 122 265 425 583 662 819 907 1074 982 984
Minimum 26 72 114 212 296 417 450 494 516 505
Maximum 279 631 1177 1544 1968 1795 1815 2435 1607 1671
Median 116 237 390 516 585 752 880 888 1023 972
5% Percentile 54 99 175 279 371 441 501 558 608 637
25% Quartile 79 154 258 392 473 547 605 693 695 686
75% Quartile 148 331 515 694 743 952 1070 1186 1200 1148
95% Percentile 217 555 857 1122 1348 1596 1625 2278 1559 1438
Sample Size 69 115 123 114 74 58 49 33 25 32
NW - 42
Table NW-7a. Empirical female walleye von Bertalanffy growth equation for northwest region waterbodies
sampled using the fall walleye index netting standard. L defines the asymptotic or maximum total length, K is the Brody growth coefficient and defines growth rate towards the maximum, and to shifts the growth curve along the axis to allow for apparent non-zero
body length at age zero (to=-1). Omega ( ) is early growth and corresponds to the growth rate near to. h is pre-maturation growth (i.e., growth to 350mm total length for combined sexes).
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Jones 32 542 0.173 94.02 57.85
Eltrut 81 565 0.174 98.18 57.34
Keikewabik 98 591 0.158 93.57 70.00
Nipigon (1997) 183 594 0.260 154.44 105.71
Steel 89 602 0.168 101.20 70.22
Savanne (2001) 78 603 0.154 92.42 No data
Whitewater 100 606 0.132 79.99 60.18
Nipigon (1998) 127 607 0.246 149.14 107.69
Nipigon (2001) (GB) 153 613 0.327 200.27 108.24
Sapawe 43 614 0.264 162.28 79.27
Savanne (1997) 122 617 0.144 88.99 No data
Calm 92 922 0.180 112.08 73.85
Three Finger 54 623 0.136 84.57 71.01
Neston 55 624 0.150 93.36 60.93
Savanne (1996) 73 630 0.141 88.98 No data
Nipigon (1999) 180 635 0.239 151.73 111.11
Wapesi 100 636 0.138 87.53 69.42
Partridge 111 640 0.187 119.56 88.12
Mount 79 642 0.207 132.82 109.48
Namakan 50 656 0.190 124.24 69.22
Rawn Reservoir 39 659 0.221 145.68 90.78
Rugby 54 661 0.153 101.24 85.07
Watcomb 43 661 0.219 144.78 87.50
Harris 53 661 0.189 124.70 78.42
Hicks 34 663 0.139 92.01 No data
Toronto 75 663 0.135 89.66 67.81
Ara 89 666 0.132 87.89 72.41
Pettit 86 668 0.233 155.92 119.77
Miminiska 125 670 0.121 80.84 56.41
Churchill 159 671 0.137 91.76 78.92
Gamsby 64 673 0.198 133.08 87.50
Henderson 53 674 0.146 98.20 No data
Sakwite 92 675 0.161 108.65 72.36
Despair 36 675 0.185 125.00 98.20
Little Sandford 103 684 0.210 143.59 91.82
Sturgeon 47 686 0.195 134.03 94.18
Botsford 132 686 0.164 112.39 74.39
Otter 42 686 0.162 110.89 96.86
Mainville 92 687 0.133 91.24 No data
Patricia 50 688 0.208 142.86 82.16
NW - 43
Table NW-7a. (continued)
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Bending 55 689 0.154 105.84 64.23
St. Joseph 547 690 0.132 90.83 76.94
Lower Companion 38 694 0.228 157.96 122.30
Georgia 81 696 0.133 92.41 83.44
Meta 44 696 0.115 80.37 66.42
Favourable 112 697 0.116 80.82 60.29
Rainy (1999) 88 697 0.171 119.35 83.94
Smoothrock 163 699 0.133 93.16 70.92
Burrows 127 701 0.142 99.31 66.26
Pelican 119 707 0.149 105.11 71.25
Brennan 63 709 0.109 77.39 61.13
Abram 102 710 0.158 112.34 76.09
Straw 86 716 0.171 122.32 94.30
Garden 86 717 0.102 73.49 54.37
Sucan 24 721 0.187 134.49 98.44
Trout 165 722 0.155 111.72 91.90
Kapkichi 118 724 0.110 79.47 64.67
Pekagoning (1999) 147 725 0.205 148.49 105.95
Little Trout 110 726 0.144 104.09 89.88
Dovetail 43 726 0.169 123.03 89.34
Kaopskikamak 38 727 0.182 132.58 103.28
Wolseley 27 731 0.182 133.23 115.37
Jutten 72 733 0.132 96.36 70.97
Finlayson 108 735 0.145 106.42 68.25
Longlegged 113 736 0.145 106.77 96.95
Perch 44 739 0.121 89.08 No data
of the Woods (2000) 182 740 0.176 130.14 93.78
Minnitaki 322 740 0.148 109.69 74.90
Obonga 161 742 0.157 116.62 95.34
Geikie 44 744 0.136 100.90 66.41
White Otter 75 749 0.137 102.73 98.79
Otukamamoan 166 753 0.147 110.90 94.53
Kenogamisis 96 753 0.104 78.58 71.51
Rainy (1997) 148 756 0.177 133.66 120.35
Devious 39 760 0.147 111.43 94.47
Hutchison 48 762 0.154 117.63 No data
Makokibatan 122 765 0.135 103.16 68.73
Dog 92 766 0.172 131.53 104.29
des Mille Lacs (1994) 126 767 0.120 91.92 59.13
Icarus 87 767 0.208 159.36 105.60
McAree 36 768 0.141 107.99 60.87
Sand Point 57 768 0.148 113.58 83.49
Factor 41 768 0.145 111.27 106.83
Seul (1996) 514 774 0.126 97.64 84.97
Whitefish (2001) 177 778 0.150 116.80 102.84
des Mille Lacs (1993) 102 781 0.114 88.97 64.50
NW - 44
Table NW-7a. (continued)
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Kukukus 129 782 0.150 117.40 92.36
Kilburn 84 783 0.120 93.52 69.52
Wintering 88 788 0.110 86.29 76.67
Dimple 31 789 0.152 119.63 98.59
Seul (1997) 431 790 0.127 100.06 89.11
Sandstone 29 790 0.183 144.64 96.47
Sydney 124 793 0.131 103.45 85.14
Broad 33 795 0.125 98.97 No data
Red 474 796 0.123 98.18 76.46
Pekagoning (1998) 169 797 0.149 118.99 97.71
Whitefish (1996) 162 798 0.134 106.94 No data
des Mille Lacs (1999) 227 799 0.121 96.67 76.22
Footprint 39 800 0.145 115.98 103.31
Wildgoose 72 801 0.158 126.93 109.17
Expanse 92 804 0.116 93.20 84.22
Onaman 65 806 0.118 95.05 98.30
Winnipeg River (1998) 209 807 0.106 85.34 78.19
Iron Range 78 810 0.145 117.23 No data
Klotz 78 812 0.158 128.68 No data
Whitefish (1995) 92 815 0.141 114.80 No data
Boffin 29 818 0.128 104.39 No data
Eagle 366 826 0.132 109.43 97.08
Robinson 47 830 0.163 135.42 131.25
Marmion 58 834 0.109 91.12 62.12
Zizania 74 839 0.134 112.38 98.97
Wabigoon 128 839 0.124 104.07 79.19
Mercutio 50 843 0.122 102.53 82.61
Pipestone 96 853 0.158 134.68 No data
Cloud 31 853 0.143 122.24 No data
Ball (1998) 40 863 0.124 106.48 93.94
Burditt 98 871 0.145 125.99 129.45
Jackfish 51 874 0.137 119.39 100.87
Northern Light 65 874 0.125 108.71 108.57
Young 63 881 0.131 115.60 127.66
Kenorain 77 884 0.135 119.73 No data
Shoal (2001) 132 888 0.156 138.06 132.76
Wawang 71 907 0.143 129.27 133.15
Gammon 32 912 0.141 128.80 No data
NW - 45
Northwest region female walleye von Bertalanffy growth equation benchmarks.
L (mm) K (mm year –1) h (mm year–1)1.
Mean 733 0.155 112.14 89.37
Minimum 542 0.102 73.49 54.37
Maximum 912 0.327 200.27 140.65
Median 732 0.145 109.56 89.24
5% Percentile 606 0.110 80.82 60.29
25% Quartile 674 0.132 93.80 72.41
75% Quartile 794 0.171 125.50 103.28
95% Percentile 874 0.228 151.73 127.66
Sample Size 124 131
1. Pre-maturation growth benchmarks are calculated from all northwest lakes.
NW - 46
Table NW-7b. Empirical male walleye von Bertalanffy growth equation for northwest region waterbodies
sampled using the fall walleye index netting standard. L defines the asymptotic or maximum total length, K is the Brody growth coefficient and defines growth rate towards the maximum, and to shifts the growth curve along the axis to allow for apparent non-zero body
length at age zero (to=-1). Omega ( ) is early growth and corresponds to the growth rate near to. h is pre-maturation growth (i.e., growth to 350mm total length for combined sexes).
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Wildgoose 120 485 0.248 120.12 109.17
Eltrut 77 486 0.200 97.17 57.34
Perch 32 491 0.198 97.22 No data
Jones 32 491 0.206 101.01 57.85
Three Finger 41 501 0.180 89.95 71.01
Nipigon (1997) 155 506 0.270 136.76 105.71
Kenogamisis 114 508 0.179 90.86 71.51
Sucan 32 515 0.214 110.42 98.44
Burrows 90 524 0.189 98.69 66.26
Rugby 61 531 0.193 102.53 85.07
Little Sandford 101 535 0.243 129.75 91.82
Jutten 80 538 0.198 106.20 70.97
Kapkichi 66 539 0.158 85.10 64.67
Ara 140 539 0.173 92.97 72.41
Keikewabik 80 539 0.163 87.73 70.00
Wapesi 101 539 0.156 83.68 69.42
Savanne (2001) 56 542 0.175 94.60 No data
Sapawe 49 547 0.214 117.10 79.27
Nipigon (1998) 121 547 0.234 127.56 107.69
Bending 31 551 0.204 112.60 64.23
Rainy (1998) 32 553 0.135 74.34 70.00
Nipigon (1999) 146 557 0.224 124.96 111.11
Brennan 54 557 0.145 80.67 61.13
Toronto 44 560 0.159 89.15 67.81
Savanne (1997) 79 561 0.142 79.29 No data
Meta 57 563 0.155 87.07 66.42
Straw 65 564 0.213 119.98 94.30
Pickwick 93 564 0.238 134.10 103.46
Neston 59 565 0.155 87.20 60.93
Rawn Reservoir 37 565 0.254 143.22 90.78
Pekagoning (1998) 126 566 0.262 148.06 97.71
Namakan 87 566 0.96 110.88 69.22
Sandstone 71 567 0.320 181.19 96.47
Mount 93 573 0.228 130.45 109.48
Makokibatan 71 574 0.194 111.17 68.73
Churchill 152 577 0.150 86.60 78.92
Geikie 31 577 0.203 116.68 66.41
Old Man 29 578 0.152 87.86 77.42
Savanne (1996) 34 580 0.143 82.99 No data
Miminiska 67 581 0.143 83.17 56.41
Whitefish (1996) 101 582 0.188 109.58 No data
NW - 47
Table NW-7b. (continued)
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Gamsby 59 585 0.176 102.68 87.50
Iron Range 133 586 0.216 126.44 No data
Mainville 94 588 0.169 99.23 No data
Despair 26 589 0.188 110.98 98.20
des Mille Lacs (1994) 95 589 0.143 84.32 59.13
Cuttle 41 589 0.182 107.19 81.82
Otukamamoan 112 592 0.186 109.86 94.53
Dovetail 37 592 0.207 122.62 89.34
Mercutio 58 593 0.195 115.90 82.61
Hicks 32 594 0.143 85.09 No data
Whitefish (2001) 199 595 0.200 118.59 102.84
Calm 41 595 0.171 101.47 73.85
Rainy (1999) 126 602 0.187 112.46 83.94
Longlegged 129 602 0.168 101.30 96.95
Finlayson 78 605 0.175 105.67 68.25
Sydney 122 606 0.176 106.90 85.14
Smoothrock 168 608 0.151 91.64 70.92
Whitefish (1995) 95 609 0.176 107.27 No data
Onaman 137 611 0.158 96.63 98.30
Garden 97 612 0.125 76.49 54.37
des Mille Lacs (1993) 80 616 0.148 91.01 64.50
Botsford 121 617 0.164 101.35 74.39
Wintering 114 618 0.150 92.40 76.67
Sakwite 57 620 0.154 95.13 72.36
Seul (1996) 506 620 0.156 96.47 84.97
Sturgeon 51 625 0.201 125.46 94.18
Shoal (1998) 77 627 0.283 177.53 140.65
Lower Companion 51 627 0.204 128.01 122.30
of the Woods (2000) 147 628 0.195 122.24 93.78
Husband 27 628 0.186 117.11 107.02
Little Trout 123 629 0.159 100.13 89.88
Wolseley 31 631 0.245 154.18 115.37
Footprint 30 632 0.172 108.52 103.31
Ice 33 634 0.178 112.59 No data
Harris 61 635 0.163 103.21 78.42
Expanse 54 636 0.151 96.22 84.22
Lount (1998) 32 636 0.146 92.57 94.89
Georgia 42 637 0.154 97.82 83.44
Sand Point 99 639 0.166 105.62 83.49
Pelican 81 640 0.173 110.80 71.25
McAree 53 642 0.154 99.05 60.87
Zizania 71 642 0.163 104.25 98.97
Watcomb 34 643 0.169 108.26 87.50
Favourable 169 645 0.121 78.24 60.29
Winnipeg River (1998) 178 647 0.139 90.12 78.19
Seul (1997) 353 650 0.152 98.98 89.11
NW - 48
Table NW-7b. (continued)
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Steel 65 652 0.118 76.90 70.22
Minnitaki 272 653 0.166 108.13 74.90
Factor 78 654 0.170 111.07 106.83
Dog 92 655 0.173 113.28 104.29
Kilburn 103 656 0.132 86.35 69.52
Trout 134 657 0.171 112.47 91.90
Kenorain 68 657 0.160 104.98 No data
Hutchison 46 658 0.196 129.12 No data
Jackfish 29 659 0.185 122.04 100.87
Whitewater 176 662 0.105 69.67 60.18
Henderson 65 663 0.137 90.67 No data
St. Joseph 439 663 0.123 81.52 76.94
Broad 72 664 0.157 104.17 No data
Northern Light 94 668 0.178 118.54 108.57
Klotz 111 671 0.190 127.36 No data
Eagle 291 674 0.166 111.70 97.08
Kukukus 110 674 0.154 103.74 92.36
Red 323 678 0.137 92.76 76.46
des Mille Lacs (1999) 226 679 0.110 74.73 76.22
Abram 73 680 0.151 102.57 76.09
Marmion 55 682 0.126 86.10 62.12
Ball (1998) 51 683 0.152 103.93 93.94
Icarus 130 683 0.190 129.83 105.60
Burditt 94 688 0.177 121.30 129.45
Rainy (1997) 132 703 0.206 144.76 120.35
Wabigoon 135 706 0.149 104.84 79.19
Cloud 63 708 0.158 112.02 No data
Wawang 48 717 0.169 120.66 133.15
Young 80 720 0.155 111.38 127.66
Pipestone 81 722 0.165 118.70 No data
Robinson 51 730 0.164 119.68 131.25
Shoal (2001) 115 741 0.196 145.38 132.76
Crooked Pine 28 784 0.147 115.16 No data
Gammon 42 812 0.134 109.07 No data
NW - 49
Northwest region male walleye von Bertalanffy growth equation benchmarks.
L (mm) K (mm year –1) h (mm year–1)1.
Mean 613 0.176 106.82 89.37
Minimum 485 0.105 69.67 54.37
Maximum 812 0.320 181.19 140.65
Median 612 0.170 105.62 89.24
5% Percentile 508 0.126 79.29 60.29
25% Quartile 566 0.152 92.57 72.41
75% Quartile 655 0.195 117.11 103.28
95% Percentile 717 0.245 143.22 127.66
Sample Size 121 131
1. Pre-maturation growth benchmarks are calculated from all northwest lakes.
NW - 50
Table NW-8a. Female walleye age and size-at-maturity (total length in millimetres) schedules for northwest region waterbodies sampled using the fall walleye index netting standard. A logistic regression model was fitted to logarithmically transformed (base 10) maturity ogive (as percents).
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Darby 2.01 0.59 3.44 340.3 280.9 399.7
Upper Companion 2.03 1.19 2.87 367.0 287.8 446.4
Shoal (2001) 2.17 0.50 3.84 498.0 468.3 527.8
Lower Companion 2.23 1.56 2.90 396.2 355.4 437.1
Pipestone 2.34 0.98 3.70 451.2 430.0 472.3
Pettit 2.56 2.07 3.05 402.5 384.2 420.9
Alph 2.57 1.49 3.64 435.7 418.8 452.6
Footprint 2.64 1.60 3.67 370.4 309.0 431.8
Union 2.65 1.85 3.46 358.0 293.7 422.4
Richardson 2.68 1.59 3.76 418.9 396.5 441.2
Mount 2.68 2.01 3.34 403.2 385.9 420.4
Nipigon (2001) (GB) 2.77 1.78 3.77 403.8 321.6 486.0
Gammon 2.81 1.45 4.18 481.6 459.7 503.4
Rainy (1997) 2.84 2.07 3.60 460.4 417.9 502.9
Whalen 2.87 2.44 3.30 365.0 348.5 381.5
Crooked Pine 2.90 1.72 4.08 518.7 492.7 544.6
Tyrell 2.90 2.36 3.45 406.4 360.5 452.3
Icarus 2.95 2.60 3.29 467.2 434.4 500.1
Kawawiag 3.00 2.40 3.60 466.5 444.1 488.9
Sucan 3.02 2.00 4.03 373.9 337.2 410.6
Despair 3.05 1.65 4.46 379.7 345.0 414.3
Nipigon (2000) 3.07 2.35 3.80 367.6 325.5 409.7
Factor 3.10 2.13 4.07 451.7 410.0 493.5
White Otter 3.11 2.33 3.88 427.9 399.9 455.9
Wildgoose 3.12 2.37 3.86 407.7 363.2 452.2
Iron Range 3.17 2.17 4.18 376.4 334.8 418.0
Sapawe 3.23 2.89 3.57 359.3 326.6 392.0
Devious 3.23 2.13 4.33 396.7 343.4 449.9
Ice 3.31 2.44 4.18 444.1 405.8 482.3
Wawang 3.33 2.18 4.47 473.0 426.8 519.3
Burditt 3.37 2.28 4.46 473.4 442.7 504.0
Pickwick 3.39 2.34 4.44 401.2 360.3 442.0
Kaopskikamak 3.41 2.37 4.45 443.9 414.4 473.4
Jackfish 3.42 2.29 4.54 426.4 397.0 455.8
Straw 3.48 2.75 4.22 375.0 355.5 394.5
Nipigon (2001) (CB) 3.53 2.31 4.75 452.0 429.3 474.7
Lac la Croix 3.58 1.48 5.68 379.3 359.1 399.6
Whitefish (2001) 3.64 2.25 5.02 426.4 380.9 471.8
Three Finger 3.64 2.97 4.32 354.1 333.6 374.6
Rawn Reservoir 3.65 3.12 4.18 449.7 422.9 476.5
Northern Light 3.72 1.80 5.64 489.8 460.0 519.5
Nipigon (1999) 3.76 2.77 4.76 456.5 406.6 506.4
NW - 51
Table NW-8a. (continued)
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Husband 3.80 2.81 4.79 479.4 461.1 497.8
Whitefish (1995) 3.80 1.85 5.74 416.5 351.8 481.2
Dimple 3.84 3.17 4.51 451.2 429.1 473.3
Henderson 3.87 2.19 5.54 424.7 391.5 457.8
Onaman 3.91 2.82 5.00 436.2 408.4 463.9
Zizania 3.93 2.96 4.90 421.3 374.3 468.4
Sandstone 3.97 3.08 4.86 456.7 368.1 545.3
Manion 4.00 3.02 4.98 492.4 468.5 516.3
Young 4.02 3.13 4.90 500.8 477.0 524.5
Shoal (1998) 4.05 3.58 4.52 531.2 495.0 567.4
Hicks 4.06 2.62 5.51 387.7 346.7 428.7
Nipigon (1998) 4.07 3.71 4.42 454.4 433.4 475.4
Pekagoning (1999) 4.13 3.67 4.59 451.2 427.1 475.4
Eagle 4.15 2.90 5.41 462.3 386.4 538.2
Little Sandford 4.21 3.26 5.16 422.7 394.7 450.7
Gamsby 4.24 3.09 5.39 427.6 360.2 495.0
Pekagoning (1998) 4.25 3.41 5.10 458.4 431.6 485.2
Otukamamoan 4.30 3.34 5.26 468.3 429.4 507.3
Obonga 4.48 3.86 5.09 459.0 436.7 481.3
Longlegged 4.50 4.05 4.95 437.9 410.8 465.0
Harris 4.51 4.18 4.84 458.9 436.3 481.5
Whiterock 4.55 2.92 6.18 500.1 485.1 515.1
Kenogamisis 4.58 2.74 6.43 368.6 330.7 406.6
Kenorain 4.61 3.45 5.76 482.0 440.4 523.5
Rainy (1999) 4.61 3.22 6.01 450.0 418.0 482.0
Sturgeon 4.63 3.93 5.34 461.5 445.1 477.9
of the Woods (2000) 4.71 3.57 5.85 461.3 405.5 517.1
Mercutio 4.72 3.71 5.72 445.4 401.5 489.2
Whitefish (1996) 4.72 2.97 6.46 429.7 368.5 490.9
Sand Point 4.75 3.57 5.93 460.8 437.4 484.2
Nipigon (1997) 4.79 3.76 5.82 482.6 454.1 511.2
Watcomb 4.83 4.09 5.56 439.1 409.1 469.1
Hutchison 4.84 4.05 5.63 480.3 453.6 506.9
Dovetail 4.92 4.50 5.35 429.0 411.2 446.9
Savanne (2001) 4.92 3.89 5.95 383.6 340.2 427.1
Old Man 5.01 4.46 5.56 420.5 401.8 439.2
Otter 5.01 4.42 5.59 473.6 452.3 494.8
Cuttle 5.04 3.35 6.73 397.5 354.9 440.0
Wabigoon 5.10 3.13 7.08 441.9 349.4 534.4
Kukukus 5.11 4.28 5.94 477.7 445.4 509.9
Seul (1997) 5.11 3.55 6.68 440.5 376.7 504.3
Seul (1996) 5.16 2.52 7.80 435.4 351.4 519.4
Expanse 5.28 3.91 6.65 448.0 411.4 484.7
Sydney 5.29 4.39 6.19 467.3 437.0 497.5
Mainville 5.40 4.26 6.54 392.9 350.6 435.2
NW - 52
Table NW-8a. (continued)
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Botsford 5.40 3.72 7.07 430.7 380.2 481.2
Burrows 5.45 4.71 6.18 413.8 368.2 459.3
Broad 5.52 2.39 8.65 482.9 441.8 524.0
Calm 5.54 4.95 6.13 450.1 423.9 476.3
Minnitaki 5.62 4.58 6.65 480.9 439.9 521.9
Eltrut 5.77 4.94 6.60 408.9 294.8 522.9
Namakan 5.81 3.79 7.84 451.1 367.8 534.3
Wintering 5.84 4.85 6.82 446.0 397.0 495.0
Trout 5.93 4.83 7.04 497.4 463.5 531.4
Little Trout 6.06 4.96 7.16 476.5 436.6 516.4
Finlayson 6.11 5.35 6.87 482.5 445.0 519.9
Geikie 6.16 4.46 7.85 453.7 428.8 478.7
Pelican 6.17 4.62 7.72 471.5 406.0 537.0
Sakwite 6.20 5.64 6.75 446.1 398.9 493.4
Jones 6.31 4.21 8.41 439.9 320.4 559.4
Smoothrock 6.31 4.30 8.31 427.5 379.6 475.4
Abram 6.38 4.40 8.35 465.0 397.2 532.7
Georgia 6.38 4.42 8.35 428.4 375.9 480.9
Red 6.39 4.45 8.34 488.3 410.8 565.8
Churchill 6.46 4.49 8.43 430.8 369.6 492.1
Kilburn 6.48 4.58 8.37 464.2 436.3 492.2
des Mille Lacs (1994) 6.49 5.61 7.36 483.7 454.1 513.4
Wapesi 6.58 4.89 8.27 412.0 373.4 450.5
Partridge 6.60 4.41 8.78 478.8 433.9 523.8
Meta 6.61 5.44 7.77 421.2 355.5 486.9
Savanne (1996) 6.71 5.77 7.65 408.7 383.4 434.1
Toronto 6.73 4.44 9.02 428.6 379.8 477.5
St. Joseph 6.81 4.82 8.81 443.4 389.6 497.2
Ara 6.85 5.96 7.73 418.8 399.3 438.2
Keikewabik 6.86 5.21 8.51 419.3 360.0 478.6
des Mille Lacs (1999) 6.88 4.64 9.13 493.8 456.7 531.0
Savanne (1997) 6.93 3.94 9.91 416.7 356.8 476.6
Jutten 6.94 6.08 7.79 476.9 446.9 507.0
Kapkichi 6.97 5.15 8.79 437.6 397.5 477.6
Steel 7.07 6.61 7.54 495.0 433.7 556.2
Marmion 7.11 6.35 7.88 443.3 420.9 465.7
Makokibatan 7.29 5.79 8.79 514.7 472.0 557.6
Neston 7.32 6.50 8.14 466.7 420.6 512.9
Winnipeg River (1998) 7.43 5.10 9.76 484.7 406.4 563.0
Miminiska 8.02 4.85 11.19 426.3 354.1 498.5
Whitewater 8.54 5.83 11.24 414.2 355.9 472.6
des Mille Lac (1993) 8.89 6.66 11.12 542.6 424.6 660.0
Brennan 9.30 8.41 10.20 510.2 485.5 535.0
Garden 10.19 7.96 12.42 492.1 454.4 529.7
NW - 53
Northwest region female walleye maturity benchmarks.
Maturity by age (years) Maturity by size (mm)
50% 10% 90% 50% 10% 90%
Mean 4.78 3.56 6.00 441.4 399.4 483.5
Minimum 2.01 0.50 2.87 340.3 280.9 374.6
Maximum 10.19 8.41 12.42 542.6 495.0 660.6
Median 4.61 3.45 5.63 444.1 405.5 482.3
5% Percentile 2.57 1.49 3.44 367.6 321.6 406.6
25% Quartile 3.41 2.36 4.44 416.7 360.3 452.6
75% Quartile 6.16 4.49 7.65 471.5 436.3 515.1
95% Percentile 7.32 6.08 9.76 500.1 468.5 556.2
Sample Size 131
NW - 54
Table NW-8b. Male walleye age and size-at-maturity (total length in millimetres) schedules for northwest region waterbodies sampled using the fall walleye index netting standard. A logistic regression model was fitted to logarithmically transformed (base 10) maturity ogive (as percents).
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Lower Companion 1.50 0.76 2.25 313.7 263.1 364.3
Burditt 1.57 0.31 2.83 361.2 292.7 429.7
Shoal (1998) 1.69 1.08 2.31 374.1 355.8 392.4
Wawang 1.74 1.01 2.46 371.1 351.5 390.7
Manion 1.76 0.50 3.01 345.8 328.2 363.4
Kawawiag 1.78 0.81 2.75 360.9 337.1 384.7
Wildgoose 1.85 0.71 2.98 316.4 252.2 380.6
Jackfish 1.85 1.07 2.62 337.6 316.5 358.7
Straw 1.89 1.08 2.71 306.4 284.9 327.9
Kaopskikamak 1.93 1.02 2.85 340.8 326.8 354.8
Shoal (2001) 2.00 0.86 3.14 401.0 339.0 463.0
Footprint 2.02 0.96 3.08 317.6 223.8 411.5
Pettit 2.05 1.80 2.30 367.9 319.9 415.7
Mount 2.07 1.19 2.94 348.8 307.3 390.2
Richardson 2.09 1.38 2.79 342.8 326.6 359.0
Young 2.10 0.68 3.52 389.9 367.6 412.3
Longlegged 2.13 0.58 3.67 338.1 310.2 365.9
Whalen 2.17 0.70 3.63 306.9 170.1 443.8
Union 2.24 1.33 3.16 336.7 319.5 353.9
Cuttle 2.28 1.46 3.11 325.5 309.7 341.3
Hutchison 2.28 1.64 2.92 361.0 342.1 379.9
Pickwick 2.43 1.48 3.39 342.9 314.3 371.5
Nipigon (2001) (GB) 2.44 1.93 2.96 378.8 328.9 428.7
Despair 2.46 1.73 3.19 329.0 312.3 345.7
White Otter 2.49 1.79 3.19 356.1 338.6 373.7
Factor 2.51 1.64 3.38 374.4 357.9 390.9
Rainy (1997) 2.52 1.44 3.61 399.1 352.5 445.7
Devious 2.55 1.58 3.52 339.6 310.7 368.5
Otukamamoan 2.59 1.84 3.34 340.7 314.3 367.0
Broad 2.59 0.54 4.65 365.7 327.8 403.6
Whitefish (1996) 2.59 1.54 3.64 345.8 308.5 383.0
Icarus 2.64 1.98 3.29 386.1 291.3 480.8
Eagle 2.70 1.16 4.25 377.1 335.8 418.5
Keikewabik 2.73 0.75 4.72 319.8 277.9 361.7
Whitefish (1995) 2.74 1.85 3.63 318.7 301.8 335.6
Northern Light 2.77 1.86 3.68 395.5 362.0 428.9
Nipigon (1999) 2.79 1.77 3.81 381.5 353.1 409.9
Georgia 2.81 1.88 3.73 335.8 321.5 350.2
of the Woods (2000) 2.83 1.96 3.70 349.4 314.2 384.5
Sturgeon 2.86 2.29 3.44 347.3 329.0 365.6
Dimple 2.86 2.44 3.29 379.5 368.9 390.1
Seul (1997) 2.88 0.75 5.01 365.4 313.8 417.0
NW - 55
Table NW-8b. (continued)
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Watcomb 2.88 2.07 3.69 333.0 273.1 392.8
Rawn Reservoir 2.91 2.54 3.27 345.1 331.7 358.6
Little Sandford 2.92 2.42 3.43 369.6 335.5 403.7
Elva 2.94 2.62 3.26 399.7 385.2 414.2
Pekagoning (1998) 2.96 2.55 3.37 374.6 351.2 397.9
Iron Range 2.97 2.11 3.82 333.6 305.1 362.0
Husband 2.98 2.38 3.58 417.2 352.4 481.8
Obonga 3.03 1.84 4.23 365.0 329.6 400.3
Pekagoning (1999) 3.05 2.32 3.78 385.3 360.0 410.6
Manomin 3.05 2.61 3.49 333.4 318.0 348.8
Mainville 3.06 2.30 3.82 301.3 268.8 333.9
Nipigon (2001) (CB) 3.06 2.87 3.25 400.6 387.0 414.2
Sandstone 3.07 2.21 3.93 366.1 302.1 430.1
Gamsby 3.12 2.33 3.92 338.7 317.6 359.8
Nipigon (2000) 3.17 2.42 3.91 371.6 324.1 419.1
Sapawe 3.20 2.51 3.89 324.1 301.9 346.3
Expanse 3.29 1.90 4.68 356.5 311.9 401.1
Kenogamisis 3.29 2.66 3.92 328.9 316.5 341.3
Dovetail 3.43 2.37 4.48 371.5 343.0 400.0
Kukukus 3.44 2.41 4.48 377.3 353.5 401.2
Nipigon (1998) 3.54 2.44 4.63 385.3 369.9 400.7
Savanne (1997) 3.61 2.46 4.77 330.5 304.7 356.3
Eltrut 3.64 2.97 4.31 295.2 251.6 338.7
Savanne (2001) 3.65 2.09 5.21 322.8 297.0 348.7
Sakwite 3.66 2.66 4.67 331.7 294.4 369.0
Mercutio 3.70 2.52 4.88 376.3 324.7 428.0
Wapesi 3.71 1.91 5.52 322.5 273.2 371.9
Wabigoon 3.73 1.94 5.53 363.1 302.5 423.7
Sydney 3.74 2.26 5.21 362.5 306.8 418.2
Chill 3.81 2.54 5.07 436.8 423.9 449.7
Harris 3.82 3.20 4.45 373.9 354.8 392.9
Seul (1996) 3.85 2.17 5.53 362.5 305.7 419.3
Churchill 3.86 2.28 5.44 345.8 275.4 416.2
Pelican 3.86 3.11 4.60 356.7 311.6 401.8
Old Man 3.87 3.33 4.42 360.6 325.0 396.3
Trout 3.89 2.69 5.09 425.1 402.8 447.4
Hicks 3.89 3.09 4.69 338.0 312.2 363.8
Rainy (1999) 3.90 2.75 5.05 373.6 338.3 408.8
Partridge 3.92 2.64 5.19 359.4 331.4 387.5
Nipigon (1997) 3.94 2.75 5.13 415.5 372.5 458.6
Burrows 3.94 2.76 5.12 336.2 300.8 371.5
des Mille Lacs (1999) 3.98 1.66 6.30 355.6 249.9 461.2
Kilburn 4.00 2.60 5.40 347.2 319.3 375.2
Botsford 4.06 3.28 4.85 360.8 319.1 402.6
Wintering 4.08 1.73 6.43 355.8 280.6 430.9
NW - 56
Table NW-8b. (continued)
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Kenorain 4.10 3.16 5.05 340.7 267.0 414.4
Calm 4.12 3.57 4.67 384.4 358.2 410.7
Meta 4.14 3.18 5.10 355.6 329.8 381.5
Otter 4.16 3.64 4.68 424.2 409.0 439.4
Jones 4.18 3.58 4.78 280.3 268.8 291.7
des Mille Lacs (1993) 4.28 3.32 5.24 340.6 294.6 386.6
Sand Point 4.32 1.91 6.74 394.2 340.0 448.4
Ara 4.42 1.92 6.91 361.0 326.0 395.9
Namakan 4.42 2.58 6.25 364.8 327.2 402.3
Minnitaki 4.44 2.91 5.96 386.5 340.6 432.3
Little Trout 4.51 2.97 6.04 408.2 354.1 452.4
Kapkichi 4.54 3.47 5.61 345.1 332.1 358.0
Agimak 4.61 3.73 5.49 341.1 331.9 350.2
des Mille Lacs (1994) 4.62 3.59 5.66 331.4 303.2 359.6
Wababimiga 4.64 3.60 5.67 439.9 354.0 525.8
Neston 4.65 3.06 6.25 329.0 292.0 366.0
Abram 4.66 2.03 7.29 392.2 324.5 459.9
Jutten 4.73 3.93 5.53 373.2 356.3 390.1
Savanne (1996) 4.74 4.14 5.34 347.7 319.8 375.6
Smoothrock 4.78 3.69 5.87 364.3 328.5 400.1
Marmion 4.94 4.18 5.70 364.8 343.0 386.6
Toronto 4.95 4.48 5.42 372.2 346.6 397.9
Rainy (1998) 4.99 3.29 6.69 371.2 353.5 388.8
Winnipeg River (1998) 5.04 1.71 8.38 376.1 325.4 426.8
Brennan 5.07 4.34 5.81 341.9 323.9 360.0
Perch 5.09 3.64 6.54 358.7 331.6 385.9
St. Joseph 5.11 2.75 7.48 383.8 327.3 440.3
Finlayson 5.11 4.53 5.69 375.5 358.2 392.9
Steel 5.28 4.42 6.15 363.2 305.3 421.1
Red 5.87 2.58 9.17 387.5 334.8 440.2
Makokibatan 6.09 4.57 7.62 441.9 371.6 512.3
Garden 6.73 5.00 8.46 366.2 319.2 413.2
Whitewater 6.89 4.74 9.05 371.4 336.9 405.9
Miminiska 7.71 5.41 10.01 386.8 354.2 419.4
NW - 57
Northwest region male walleye maturity benchmarks.
Maturity by age (years) Maturity by size (mm)
50% 10% 90% 50% 10% 90%
Mean 3.49 2.36 4.62 359.9 323.1 396.6
Minimum 1.50 0.31 2.25 280.3 170.1 291.7
Maximum 7.71 5.41 10.01 441.9 423.9 525.8
Median 3.43 2.33 4.48 361.0 325.0 395.9
5% Percentile 1.85 0.71 2.75 316.4 267.0 341.3
25% Quartile 2.59 1.71 3.43 339.6 305.7 366.0
75% Quartile 4.16 2.97 5.49 376.1 343.0 418.5
95% Percentile 5.11 4.42 7.48 415.5 371.6 459.9
Sample Size 121
NW
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27
.51
0
.77
1
2.2
5
56
.39
2
6.6
8
17
.84
2
3.7
2
30
.00
3
8.0
1
86
Ea
gle
3
9.2
2
1.5
5
5.8
8
63
.12
4
3.5
1
8.0
2
33
.88
4
8.9
0
55
.14
8
8
Alp
h
39
.54
1
.67
3
1.5
6
47
.07
3
9.1
2
32
.70
3
5.4
9
44
.40
4
6.3
2
10
Ga
mm
on
4
6.2
9
1.2
9
36
.41
5
7.1
4
45
.63
5
5.0
5
43
.06
5
1.2
9
55
.05
2
0
Me
ta
46
.48
1
.92
3
0.2
6
66
.74
4
6.3
5
31
.08
4
1.6
5
51
.56
6
1.9
8
24
Lo
ng
leg
ge
d
50
.29
1
.36
4
1.1
0
59
.58
5
0.7
6
41
.53
5
0.7
6
54
.09
5
8.2
1
17
Ara
5
2.0
0
1.0
4
33
.42
7
6.3
9
50
.95
4
1.6
9
46
.52
5
7.3
1
63
.16
6
2
On
am
an
5
3.2
9
1.3
0
35
.79
7
0.6
0
54
.06
3
9.5
7
49
.37
5
7.9
3
65
.43
3
9
Wa
ba
bim
iga
5
3.5
0
2.4
7
47
.94
5
9.5
0
53
.29
4
8.4
8
50
.65
5
6.1
4
58
.83
4
Syd
ne
y
55
.36
2
.12
4
2.1
8
69
.99
5
4.2
9
44
.39
4
9.6
1
62
.71
6
7.4
9
16
Exp
an
se
5
6.8
0
3.9
7
21
.37
8
93
.49
5
5.6
6
34
.05
5
0.8
3
61
.72
7
9.5
7
16
Se
ul
(1997)
56
.95
1
.72
3
2.9
6
19
9.6
0
56
.53
4
0.7
4
49
.04
6
1.9
0
68
.94
9
8
Kilb
urn
6
9.9
0
10
.00
4
2.8
1
22
4.0
8
59
.38
4
6.0
9
53
.64
6
5.5
8
11
7.8
6
17
No
rth
we
st
reg
ion
wa
lleye
re
lati
ve
fe
cu
nd
ity b
en
ch
ma
rks
.
Ave
rag
es
M
ean
Min
imu
m
Maxim
um
Media
n
5%
P
erc
en
tile
25
%Q
ua
rtile
75
%Q
ua
rtile
95
%P
erc
en
tile
Re
gio
n A
ve
rag
e1
.4
1.3
3
26
.18
7
2.3
0
40
.44
2
9.3
2
36
.29
4
5.1
7
54
.55
Re
gio
n A
ve
rag
e2
.4
0.6
6
24
.98
7
3.0
2
39
.73
2
8.2
6
35
.49
4
4.5
6
54
.32
1.
All
lakes c
om
bin
ed (
n=
19).
2.
Only
lakes w
ith
sam
ple
siz
e
10 (
n=
18).
NW
- 5
9
Ta
ble
NW
-10
. W
alle
ye
fe
cu
nd
ity-t
ota
l le
ng
th r
ela
tio
nsh
ips f
or
no
rth
we
st
reg
ion
wa
terb
od
ies s
am
ple
d u
sin
g t
he
fa
ll w
alle
ye
in
de
x
ne
ttin
g
sta
nd
ard
. A
g
eo
me
tric
m
ea
n
pre
dic
tive
re
gre
ssio
n
mo
de
l w
as
fitte
d
to
log
ari
thm
ically
tra
nsfo
rme
d (
Na
pie
ria
n lo
ga
rith
ms)
ind
ivid
ua
l o
bse
rva
tio
ns.
To
tal le
ngth
in
mill
ime
tre
s.
Fe
cu
nd
ity-l
en
gth
re
latio
n1
.C
alc
ula
ted
fe
cu
nd
ity a
t sta
nd
ard
to
tal le
ng
th (
mm
)
La
ke
x 1
0-6
R2
n
40
0
45
0
50
0
55
0
60
0
65
0
Ga
mm
on
2
.63
5
14
7.4
5
0.8
6
20
3
54
47
4
83
06
6
37
15
8
18
50
1
02
87
9
12
69
63
Syd
ne
y
2.8
3
15
81
.84
0
.86
1
6
35
71
7
49
81
7
67
08
8
87
81
8
11
22
89
1
40
78
1
Min
nita
ki
2.8
6
63
7.0
1
0.5
6
86
1
75
06
2
45
15
3
31
32
4
35
10
5
57
98
7
01
46
Se
ul
(1997)
3.0
9
32
6.9
9
0.8
1
98
3
60
55
5
18
88
7
18
61
9
64
79
1
26
24
9
16
16
85
Pe
lica
n
3.2
8
42
.43
2
0.7
1
25
1
41
49
2
08
11
2
93
88
4
01
56
5
33
98
6
94
05
On
am
an
3
.42
4
1.0
48
0
.89
3
9
32
90
9
49
24
4
70
62
1
97
85
4
13
17
91
1
73
31
6
Ara
3
.49
2
2.2
02
0
.78
6
2
27
54
1
41
56
6
60
06
9
83
81
2
11
35
98
1
50
26
4
Wa
big
oo
n
3.6
0
4.3
61
0
.93
3
1
10
00
0
15
27
5
22
31
5
31
44
1
42
99
7
57
34
4
Lo
ng
leg
ge
d
3.6
1
10
.69
4
0.8
6
17
2
60
95
3
99
11
5
83
68
8
23
20
1
12
67
7
15
03
99
Bo
tsfo
rd
3.6
2
4.8
90
0
.71
3
3
12
78
0
19
57
3
28
65
9
40
46
4
55
44
1
74
07
0
Exp
an
se
3
.66
8
.71
7
0.6
6
16
2
94
85
4
53
87
6
67
58
9
46
44
1
30
16
1
17
44
96
Me
ta
3.6
8
6.6
61
0
.75
2
4
24
52
0
37
80
8
55
69
3
79
06
3
10
88
67
1
46
11
4
Ab
ram
3
.72
2
.57
4
0.7
1
24
1
24
97
1
93
75
2
86
81
4
08
98
5
65
43
7
61
73
Ma
ko
kib
ata
n
3.8
2
1.2
40
0
.64
3
2
10
66
5
16
72
1
25
00
2
35
97
5
50
15
1
68
07
7
Mim
inis
ka
3
.89
0
.59
8
0.6
4
60
8
16
7
12
92
0
19
47
6
28
23
1
39
62
0
54
11
4
Alp
h
4.2
4
0.1
94
0
.96
1
0
20
77
7
34
23
2
53
50
7
80
14
6
11
58
98
1
62
72
1
Ea
gle
5
.82
0
.00
00
05
7
0.6
7
88
8
12
1
16
12
6
29
78
8
51
89
5
86
14
5
13
73
09
1. T
wo-t
aile
d S
tudent
t-te
sts
were
em
plo
yed t
o t
est
sig
nific
ance o
f slo
pe a
nd inte
rcept
(o:
or
= 0
).
NW
- 6
0
No
rth
we
st
reg
ion
wa
lleye
fe
cu
nd
ity-t
ota
l le
ng
th b
en
ch
ma
rks
.
F
ecu
nd
ity-l
en
gth
re
latio
n
Ca
lcu
late
d fe
cu
nd
ity a
t sta
nd
ard
to
tal le
ng
th (
mm
)
x 1
0-6
R2
n
40
0
45
0
50
0
55
0
60
0
65
0
Me
an
3
.60
4
61
.70
0
.76
4
0
21
32
0
31
97
0
46
12
5
64
50
4
87
91
2
11
72
58
Min
um
um
2
.63
0
.00
00
05
7
0.5
6
10
8
12
1
12
92
1
19
47
7
28
23
2
39
62
0
54
11
5
Ma
xim
um
5
.82
5
14
7.4
5
0.9
6
98
3
60
56
5
18
89
7
18
62
9
78
54
1
31
79
2
17
44
97
Me
dia
n
3.6
1
8.7
20
0
.75
3
1
20
77
7
34
23
2
53
50
7
79
06
4
10
28
79
1
37
31
0
5%
Pe
rce
ntile
2
.63
0
.00
00
05
7
0.5
6
10
8
12
1
12
92
1
19
47
7
28
23
2
39
62
0
54
11
5
25
% Q
ua
rtile
3
.28
2
.57
0
0.6
7
20
1
24
97
1
93
76
2
86
82
4
04
65
5
54
42
7
01
47
75
% Q
ua
rtile
3
.72
4
2.4
30
0
.86
6
0
29
48
6
45
38
8
63
71
6
83
81
3
11
35
98
1
50
40
0
95
% P
erc
en
tile
5
.82
5
14
7.4
5
0.9
6
98
3
60
56
5
18
89
7
18
62
9
78
54
1
31
79
2
17
44
97
Sa
mp
le S
ize
1
7
NW
- 6
1
Ta
ble
NW
-11
a.
Ma
ture
fe
ma
le w
alle
ye
fa
ll g
on
ad
oso
ma
tic i
nd
ex (
GS
I is
ova
ry w
eig
ht
exp
resse
d a
s a
pe
rce
nt
of
the
to
tal
we
igh
t m
inus
the
ova
ry w
eig
ht)
, vis
ce
ral fa
t in
de
x (
VF
I is
fa
t w
eig
ht
exp
resse
d a
s a
pe
rce
nt
of
the
to
tal w
eig
ht)
, a
nd
re
pro
du
ctive
in
de
x
(R i
s 3
h/L
) d
ata
fo
r n
ort
hw
est
reg
ion
wa
terb
od
ies s
am
ple
d u
sin
g t
he
fa
ll w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
. O
nly
me
an
va
lue
s w
ith
a m
inim
um
sa
mp
le s
ize
3
are
in
clu
de
d.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Mim
inis
ka
1
.54
%
0.2
5%
4
.39
%
64
0
.28
%
0%
1
.83
%
64
0
.25
Ma
ko
kib
ata
n
2.0
5%
0
.66
%
2.9
6%
3
3
4.3
9%
0
%
8.1
9%
3
3
0.2
7
Sh
oa
l(1
998)
2.5
7%
2
.07
%
4.4
2%
1
4
3.4
4%
2
.31
%
4.7
3%
8
0
.57
Alp
h
2.8
0%
1
.61
%
4.7
4%
1
0
5.1
8%
1
.98
%
7.5
9%
1
0
No
da
ta
Ara
4
.12
%
2.6
7%
7
.00
%
62
1
.82
%
0%
4
.18
%
62
0
.33
Wa
ba
bim
iga
5
.80
%
5.6
6%
6
.02
%
4
3.5
8%
3
.20
%
3.7
3%
4
N
o d
ata
No
rth
we
st
ma
ture
fe
ma
le g
on
ad
os
om
ati
c,
vis
ce
ral
fat,
an
d r
ep
rod
uc
tiv
e i
nd
ice
s b
en
ch
ma
rks
.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at In
de
x
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Re
gio
n A
ve
rag
e1
.3
.15
%
2.1
5%
4
.92
%
32
3
.11
%
1.2
5%
5
.04
%
30
0
.36
Re
gio
n A
ve
rag
e2
.2
.62
%
1.4
5%
4
.70
%
37
3
.02
%
0.8
6%
5
.31
%
35
0
.36
1.
All
lakes c
om
bin
ed (
n=
6).
2.
O
nly
la
ke
s w
ith
sam
ple
siz
e
10 (
n=
5).
NW
- 6
2
Ta
ble
NW
-11
b.
Ma
ture
ma
le w
alle
ye
fa
ll g
on
ad
oso
ma
tic in
de
x (
GS
I is
te
ste
s w
eig
ht
exp
resse
d a
s a
pe
rce
nt
of
the
to
tal w
eig
ht
min
us t
he
te
ste
s w
eig
ht)
, vis
ce
ral fa
t in
de
x (
VF
I is
fa
t w
eig
ht
exp
resse
d a
s a
pe
rce
nt
of
the
to
tal w
eig
ht)
, a
nd
re
pro
du
ctive
in
de
x (
R
is 3
h/L
) d
ata
fo
r n
ort
hw
est
reg
ion
wa
terb
od
ies s
am
ple
d u
sin
g t
he
fa
ll w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
. O
nly
me
an
va
lue
s
with
a m
inim
um
sa
mp
le s
ize
3
are
in
clu
de
d.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Mim
inis
ka
1
.73
%
0.4
9%
3
.50
%
38
0
.66
%
0%
1
2.9
7%
3
8
0.2
9
Ma
ko
kib
ata
n
1.7
5%
0
.58
%
3.3
2%
1
2
1.5
8%
0
%
3.4
2%
1
2
0.3
6
Sh
oa
l(1
998)
3.1
0%
0
.10
%
5.3
2%
5
9
2.3
9%
1
.03
%
4.5
7%
3
8
0.6
7
Alp
h
3.4
5%
0
.93
%
5.4
5%
1
1
3.1
5%
1
.26
%
4.6
3%
1
1
No
da
ta
Ara
3
.58
%
0.2
5%
6
.62
%
11
1
0.9
2%
0
%
3.4
4%
1
11
0
.40
Wa
ba
bim
iga
5
.03
%
4.3
0%
6
.35
%
11
3
.68
%
2.9
5%
4
.79
%
11
N
o d
ata
No
rth
we
st
ma
ture
ma
le g
on
ad
os
om
ati
c,
vis
ce
ral
fat,
an
d r
ep
rod
uc
tiv
e i
nd
ice
s b
en
ch
ma
rks
.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at In
de
x
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Re
gio
n A
ve
rag
e1
.3
.11
%
1.1
1%
5
.09
%
40
2
.06
%
0.8
7%
5
.64
%
37
0
.43
Re
gio
n A
ve
rag
e2
.3
.11
%
1.1
1%
5
.09
%
40
2
.06
%
0.8
7%
5
.64
%
37
0
.43
1.
All
lakes c
om
bin
ed (
n=
6).
2.
O
nly
la
ke
s w
ith
sam
ple
siz
e
10 (
n=
6).
NW
- 6
3
Ta
ble
NW
-12
. W
alle
ye
life
his
tory
pa
ram
ete
rs f
or
On
tari
o w
ate
rbo
die
s s
am
ple
d u
sin
g th
e w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
199
3 t
o 2
00
1.
Pe
rce
ntile
sL
ife
His
tory
Pa
ram
ete
r (S
am
ple
Siz
e)
Me
an
(±9
5%
C.I
.)1
.S
.D.
(C.V
.)2
.M
inim
um
to
Maxim
um
M
ed
ian
5%
2
5%
7
5%
9
5%
Fe
ma
le a
du
lt m
ort
alit
y (
A%
) (n
=2
96
)2
8(2
7-3
0)
13
.6(4
8%
)7
to
89
2
5
12
1
9
34
5
7
Ma
le a
du
lt m
ort
alit
y (
A%
) (n
=2
95
)3
0(2
9-3
2)
12
.8(4
2%
)6
to
82
2
8
14
2
1
38
5
1
Fe
ma
le r
ela
tive
co
nd
itio
n
(n=
24
1)
1.0
3(1
.02
-1.0
4)
0.0
79
(8%
)0
.90
to
1.4
9
1.0
2
0.9
4
0.9
8
1.0
7
1.1
6
Ma
le r
ela
tive
co
nd
itio
n
(n=
26
5)
1.0
3(1
.02
-1.0
4)
0.0
90
(9%
)0
.87
to
1.4
8
1.0
2
0.9
2
0.9
7
1.0
7
1.1
8
Fe
ma
le a
sym
pto
tic le
ngth
(L
, m
m)
(n=
23
8)
74
4(7
33
-75
5)
84
.2(1
1%
)5
29
to
95
7
74
8
60
6
68
4
80
4
88
0
Ma
le a
sym
pto
tic le
ng
th (
L,
mm
) (n
=2
52
)6
18
(61
0-6
26
)6
4.7
(10
%)
48
5 t
o 8
14
6
16
5
08
5
72
6
54
7
38
Fe
ma
le B
rod
y c
oe
ffic
ien
t (K
, yr-1
)(n
=2
38
)0
.15
8(0
.15
4-0
.16
3)
0.0
37
7(2
4%
)0
.09
0 t
o 0
.32
7
0.1
52
0.1
10
0.1
32
0.1
80
0.2
30
Ma
le B
rod
y c
oe
ffic
ien
t (K
, yr-1
)(n
=2
52
)0
.18
0(0
.17
6-0
.18
5)
0.0
35
0(1
9%
)0
.10
5 t
o 0
.32
00
.17
60
.12
90
.15
5
0.2
00
0.2
44
Fe
ma
le O
me
ga
(,
mm
•yr-1
)(n
=2
38
)1
16
.40
(11
3.4
1-1
19
.38
)2
3.3
22
(20
%)
73
.49
to
20
0.2
7
11
4.4
5
80
.84
9
9.3
1
13
2.8
2
15
5.9
2
Ma
le O
me
ga
(,
mm
•yr-1
)(n
=2
52
)1
10
.57
(10
8.1
2-1
13
.02
)1
9.7
48
(18
%)
69
.45
to
18
1.1
9
10
9.4
1
80
.67
9
7.2
0
12
2.0
7
14
8.0
6
Pre
-ma
tura
tio
n g
row
th (
h,
mm
•yr-1
)(n
=2
78
)9
3.0
0(9
0.4
8-9
5.5
1)
21
.30
6(2
3%
)5
4.3
1 t
o 1
85.9
0
92
.07
6
1.1
3
76
.59
1
05
.95
1
29
.45
Fe
ma
le a
ge
-at-
50
% m
atu
rity
(ye
ars
) (n
=2
61
)4
.5(4
.3-4
.7)
1.5
1(3
4%
)2
.0 t
o 1
0.2
4
.2
2.5
3
.4
5.4
7
.1
Ma
le a
ge
-at-
50
% m
atu
rity
(ye
ars
) (n
=2
66
)3
.0(2
.9-3
.2)
1.1
0(3
7%
)0
.8 t
o 7
.7
2.9
1
.5
2.2
3
.7
4.9
NW
- 6
4
Ta
ble
NW
-12
. (c
on
tin
ued
)
Pe
rce
ntile
sL
ife
His
tory
Pa
ram
ete
r (S
am
ple
Siz
e)
Me
an
(±9
5%
C.I
.)1
.S
.D.
(C.V
.)2
.M
inim
um
to
Maxim
um
M
ed
ian
5%
2
5%
7
5%
9
5%
Fe
ma
le s
ize
-at-
50
% m
atu
rity
(m
m)
(n=
26
1)
44
1(4
36
-44
7)
42
.1(1
0%
)3
38
to
58
6
44
0
36
9
41
6
47
3
50
4
Ma
le s
ize
-at-
50
% m
atu
rity
(m
m)
(n=
26
6)
34
9(3
45
-35
3)
32
.9(9
%)
24
6 t
o 4
76
3
46
2
99
3
31
3
70
4
01
Fe
ma
le g
on
ad
oso
ma
tic in
de
x (
%)
(n=
15
1)
4.4
(4.2
-4.7
)1
.64
(37
%)
1.5
to
11
.1
4.2
2
.1
3.3
5
.4
7.5
Ma
le g
on
ad
oso
ma
tic ind
ex (
%)
(n=
15
5)
2.9
(2.8
-3.0
)0
.76
(26
%)
1.3
to
5.4
2
.8
1.8
2
.5
3.2
4
.5
Re
lative
fe
cu
nd
ity (
eg
gs•g
-1)
(n=
13
6)
49
.79
(47
.41
-52
.17
)1
4.0
49
(28
%)
18
.04
to
87
.98
5
0.4
7
26
.22
4
0.0
1
58
.69
7
2.6
5
Fe
ma
le v
isce
ral fa
t in
de
x (
%)
(n=
15
1)
1.9
(1.7
-2.1
)1
.29
(68
%)
0.1
to
6.8
1
.6
0.3
0
.8
2.7
4
.4
Ma
le v
isce
ral fa
t in
de
x (
%)
(n=
15
5)
1.4
(1.2
-1.5
)0
.98
(72
%)
0.1
to
5.3
1
.1
0.2
0
.6
1.8
3
.2
Re
lative
ab
un
da
nce
(n
um
be
r•n
et-1
)(n
=4
41
)7
.5(6
.9-8
.2)
7.1
3(9
5%
)0
to
36
.1
5.3
0
.4
2.0
1
0.9
2
2.5
1.
95%
Confidence inte
rval of
the m
ean.
2.
Sta
ndard
devia
tion o
f th
e m
ean (
S.D
.) a
nd c
oeffic
ient
of
variation o
f th
e m
ean e
xpre
ssed a
s a
perc
ent
(C.V
.).
Fig
ure
3. M
ap o
f nort
heaste
rn O
nta
rio s
how
ing the location o
f th
e 1
31
wate
rbodie
s for
whic
h fall
walle
ye index n
ettin
g d
ata
were
availa
ble
.
NE
- 1
Ta
ble
NE
-1.
Th
e n
ort
he
ast
reg
ion
wate
rbo
die
s s
am
ple
d u
sin
g t
he
fa
ll w
alle
ye
in
de
x n
ett
ing
sta
nda
rd 1
99
4 t
o 2
00
1.
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Ab
itib
i1
99
5,
19
96
&
20
01
48
39
30
-79
50
42
9
09
71
.7
3.5
1
5.2
0
.3
13
16
Ag
ne
w
19
98
& 1
99
9
46
21
19
-81
45
40
2
91
6.7
1
2.1
3
9.6
2
.9
16
04
Am
iko
ug
am
i 2
00
1
48
10
59
-80
04
49
3
59
.4
1.8
4
.9
1.7
1
37
9
An
jiga
mi
20
00
4
75
00
0-8
43
50
0
11
06
.9
12
.4
45
.1
3.5
1
26
2
Ba
reh
ea
d
19
98
4
90
24
7-8
55
43
8
21
6.4
5
.3
18
.1
1.8
1
23
0
Be
arh
ea
d
19
99
4
61
82
6-8
25
83
2
26
6.8
3
.6
7.0
4
.6
16
70
Bir
ch
1
99
6
46
18
08
-81
58
47
1
07
9.7
1
0.3
2
7.4
3
.5
16
13
Bis
co
tasi
19
99
& 2
00
0
47
17
58
-82
04
55
6
57
9.4
4
.3
24
.0
3.9
1
44
7
Bu
rnfie
ld
19
99
4
82
05
0-8
50
20
7
11
3.5
4
.2
14
.6
3.5
1
13
3
Bu
rntb
ush
2
00
0
49
36
00
-79
59
00
1
26
.3
1.0
2
.1
0.9
1
15
6
Ca
no
e
19
98
4
61
65
0-8
30
22
1
16
8.8
3
.4
10
.1
2.9
1
68
5
Ca
tara
ct
19
99
4
61
53
3-8
30
00
8
10
8.3
1
.7
4.9
3
.8
17
04
Cra
b’s
To
e
19
98
4
82
65
5-8
42
11
8
19
7.9
4
.8
9.8
3
.2
12
00
Cro
uch
1
99
8
48
26
55
-84
21
18
1
97
.9
4.8
9
.8
3.2
1
20
0
De
pa
rtu
re
19
98
4
91
43
6-8
14
74
8
20
9.3
6
.1
16
.0
1.5
1
32
6
De
sch
en
es
20
00
4
72
23
6-8
13
04
7
62
1.5
3
.6
11
.0
0.8
1
46
1
Do
g
19
99
4
81
81
1-8
40
65
8
51
84
.0
12
.6
74
.7
4.4
1
25
5
Du
bo
rne
2
00
0
46
15
00
-82
55
00
9
33
.2
8.4
3
3.5
4
.6
17
03
Du
nca
n
19
98
4
74
52
1-8
05
75
4
11
31
.1
7.5
4
0.2
3
.7
14
56
Ech
o
19
98
4
63
32
3-8
35
90
1
11
49
.3
13
.9
27
.5
2.0
1
66
8
Fa
wce
tt
19
98
4
73
20
0-8
10
70
0
No
da
ta
No
da
ta
No
da
ta
No
da
ta1
46
6
Fir
th
19
98
4
74
01
5-8
05
21
0
43
3.4
2
.0
3.1
2
.1
14
65
Fla
nd
ers
1
99
9
49
21
44
-85
22
29
2
38
.0
3.7
2
7.5
3
.4
12
54
Fu
sh
imi
19
98
4
94
92
2-8
35
30
5
12
30
.2
3.1
7
.6
1.5
1
22
9
Ga
rnh
am
2
00
1
49
00
33
-85
29
29
6
73
.4
7.3
3
3.0
3
.2
12
28
Ge
ne
va
1
99
9
46
45
52
-81
32
47
3
56
.1
6.3
2
5.3
4
.5
15
12
Go
wg
an
da
1
99
8
47
37
41
-80
47
15
1
04
6.1
8
.1
45
.1
2.1
1
46
5
NE
- 2
Ta
ble
NE
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Gra
nite
hill
2
00
1
49
05
11
-85
15
53
1
44
7.6
5
.5
15
.6
2.7
1
21
9
Gra
ssy
19
97
4
74
74
2-8
11
41
0
17
2.6
1
.6
22
.6
3.5
1
45
4
Gro
un
dh
og
2
00
1
48
06
21
-82
14
16
9
01
.3
2.5
1
9.5
1
.5
14
01
Ha
mm
er
19
99
4
82
41
1-8
50
54
8
45
8.3
3
.6
11
.9
3.7
1
13
5
Hig
hb
rush
2
00
1
47
45
10
-83
32
51
3
77
.5
6.1
1
8.4
3
.5
13
11
Ho
we
lls
19
99
4
95
62
4-8
15
84
9
58
.7
1.1
4
.3
4.0
1
25
4
Iva
nh
oe
1
99
9
48
04
58
-82
37
37
1
01
3.8
6
.9
24
.0
2.8
1
36
7
Jo
wse
y
19
99
4
82
22
9-8
14
32
6
18
7.8
1
.9
7.9
1
.8
13
83
Ju
mp
ing
Ca
rib
oo
1
99
9
46
52
57
-79
46
33
4
08
.7
8.9
3
3.6
5
.0
15
41
Ju
nctio
n
20
00
4
75
72
1-8
10
42
0
51
.8
2.0
5
.5
3.1
1
42
8
Ka
bin
aka
gam
i 1
99
8
48
54
11
-84
25
01
1
20
16
.7
3.0
1
5.3
1
.7
12
20
Ka
gia
no
2
00
0
49
16
00
-86
26
00
1
36
.0
4.8
4
2.0
2
.5
11
69
Ke
bskw
ashe
sh
i 1
99
9
47
23
23
-82
54
10
8
50
.3
0.7
4
.3
1.4
1
39
6
Ke
cil
19
98
4
61
53
3-8
21
74
8
42
5.9
1
2.1
2
6.2
5
.0
16
39
Ke
ne
tog
am
i 1
99
8
47
46
56
-81
38
20
1
58
.6
3.2
1
3.0
2
.1
14
41
Ke
no
ga
min
g
19
97
4
80
34
1-8
15
41
4
65
6.5
2
.8
15
.8
2.1
1
40
6
Ke
no
ga
mis
si
20
00
4
81
10
2-8
13
30
0
24
98
.6
4.4
2
4.0
2
.0
14
15
Kn
ife
1
99
9
48
18
07
-85
05
48
5
61
.1
3.6
1
8.0
2
.7
11
33
Ku
sh
og
1
99
7
47
55
10
-80
15
25
7
42
.5
7.9
3
6.6
2
.3
14
42
La
bitic
he
1
99
7
46
59
00
-82
14
00
1
44
.3
4.8
1
5.5
4
.0
14
44
La
c la
Ca
ve
1
99
9
46
22
00
-78
45
00
N
o d
ata
N
o d
ata
N
o d
ata
N
o d
ata
17
00
La
Mo
tte
2
00
0
47
44
16
-81
39
10
1
15
.2
2.7
1
1.5
3
.6
14
42
La
rde
r 1
99
7
48
05
08
-79
38
37
3
70
3.7
1
2.3
3
3.5
4
.3
14
01
Le
ft
20
00
4
81
25
3-8
14
60
4
56
.9
0.5
2
.0
1.5
1
39
9
Llo
yd
1
99
9
47
51
43
-80
59
42
8
19
.1
5.8
2
5.6
4
.9
14
26
Lo
ng
po
int
20
01
4
74
10
1-8
03
63
5
37
6.6
4
.3
17
.7
3.7
1
45
5
Lo
we
r S
teth
am
2
00
0
47
46
12
-81
34
07
6
8.8
5
.5
23
.0
3.8
1
45
4
Ma
cu
tag
on
2
00
1
49
06
11
-85
28
37
1
49
.0
5.3
1
8.5
3
.1
12
49
NE
- 3
Ta
ble
NE
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Ma
nito
u
20
00
4
54
63
9-8
15
90
1
10
46
0.7
1
5.1
4
9.1
8
.4
16
19
Ma
ria
n
20
00
4
64
61
8-7
94
70
7
27
4.0
4
.4
13
.7
2.9
1
55
7
Ma
tta
ga
mi
19
98
4
75
00
9-8
13
34
0
40
03
.1
8.3
7
6.2
3
.3
14
48
McA
rth
ur
20
00
4
81
22
2-8
11
25
7
26
1.9
3
.4
12
.8
3.1
1
40
7
McD
iarm
id
19
99
4
82
80
6-7
93
33
7
40
3.0
1
.2
1.5
0
.3
13
38
McP
ha
il R
ese
rvo
ir
20
00
4
75
60
0-8
45
10
0
32
2.0
4
.7
17
.1
4.1
1
25
7
Me
so
mik
en
da
2
00
0
47
38
53
-81
52
44
1
70
6.2
1
3.6
7
1.4
5
.3
14
45
Min
de
mo
ya
1
99
7 &
20
01
45
45
40
-82
12
14
3
86
9.7
7
.4
21
.0
2.9
1
65
8
Min
isin
akw
a
19
99
4
73
94
8-8
14
41
8
19
11
.1
3.3
1
8.3
1
.8
14
57
Min
istic
19
97
4
63
33
9-8
13
40
8
56
7.2
1
1.1
3
1.1
5
.3
15
77
Mis
sin
aib
i 1
99
9
48
21
24
-83
41
29
7
70
6.9
1
9.2
9
4.0
4
.2
12
86
Mis
tin
iko
n
19
99
4
75
55
3-8
04
41
3
13
57
.1
5.9
3
6.6
2
.9
14
25
Mo
ntr
ea
l R
ive
r 2
00
0
47
14
00
-84
39
00
N
o d
ata
N
o d
ata
N
o d
ata
N
o d
ata
14
38
Mo
ntr
ose
2
00
0
47
58
30
-81
03
58
2
50
.0
3.8
6
.1
2.4
1
42
3
Mo
un
tain
1
99
8
47
38
46
-80
13
39
5
12
.8
4.7
4
8.8
2
.3
14
84
Na
ba
kw
asi
19
98
& 2
00
04
73
24
4-8
12
62
3
88
4.0
4
.7
26
.8
2.5
1
47
2
Na
ga
ga
mi
19
99
4
92
51
9-8
50
23
1
53
62
.5
7.5
2
7.5
2
.7
12
48
Na
ga
ga
mis
is
19
99
4
92
83
8-8
43
92
6
23
31
.1
3.8
8
.1
3.0
1
22
3
Ne
vill
e
20
00
4
73
94
6-8
15
52
0
11
9.0
1
.9
9.9
1
.8
14
40
Nig
ht
Ha
wk
19
97
4
82
82
2-8
05
83
3
98
32
.2
1.7
4
.6
0.3
1
39
5
Nip
issin
g1
99
8,
19
99
, 2
00
0 &
20
01
46
17
00
-80
00
00
8
73
25
.0
4.5
5
2.0
2
.5
16
59
Nu
ge
nt
19
97
4
75
25
2-8
11
35
6
11
0.5
2
.3
4.9
3
.4
14
39
Ob
ush
ko
ng
2
00
1
47
42
26
-80
48
00
4
37
.4
2.4
4
.6
2.7
1
46
0
Oke
2
00
1
48
56
14
-81
52
43
3
54
.0
1.5
3
.1
1.1
1
33
5
On
ap
ing
1
99
9 &
20
00
47
04
00
-81
30
00
6
86
8.0
1
0.5
5
3.4
5
.1
14
96
Op
ee
pe
esw
ay
19
98
4
73
70
0-8
21
50
0
15
75
.0
4.7
1
4.0
2
.5
14
25
Op
ikin
imik
a
20
01
4
72
20
0-8
12
50
0
58
3.0
2
7.4
5
4.9
3
.0
14
65
NE
- 4
Ta
ble
NE
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Osca
r 2
00
0
46
18
12
-83
05
44
1
02
.7
6.3
2
0.8
6
.5
16
87
Pa
pa
ko
me
ka
2
00
0
48
15
52
-81
17
31
2
02
.3
9.8
4
5.8
2
.0
14
09
Piv
ab
iska
1
99
8
49
48
47
-83
42
25
7
98
.9
3.1
6
.7
1.5
1
22
9
Po
pe
ye
1
99
9
46
26
39
-82
32
42
5
6.4
7
.1
15
.2
4.6
1
55
4
Po
st
19
98
4
70
10
0-8
11
30
0
17
0.4
1
8.2
5
5.5
3
.0
15
07
Ra
bb
it
20
00
4
65
94
3-7
93
83
9
21
06
.0
14
.2
42
.7
5.8
1
52
6
Ra
msa
y
19
98
4
92
70
0-8
54
70
0
14
4.0
N
o d
ata
2
4.0
N
o d
ata
12
96
Ra
mse
y
19
96
4
62
85
7-8
05
70
1
87
4.4
9
.3
21
.3
3.8
1
69
5
Re
d C
ed
ar
20
01
4
64
24
5-7
95
61
4
24
21
.7
5.8
3
4.7
2
.9
15
80
Re
dw
ing
2
00
0
47
59
37
-81
06
11
5
6.3
3
.8
6.7
2
.7
14
25
Re
nn
ie
19
99
4
82
30
9-8
35
75
9
57
9.3
7
.5
29
.3
4.7
1
25
4
Ritch
ie
19
99
4
95
24
4-8
33
11
1
96
.9
3.9
1
2.0
1
.7
12
22
Ro
cky I
sla
nd
1
99
9
46
55
06
-83
01
22
5
66
4.1
1
0.2
7
9.9
4
.1
14
66
Ro
un
d1
99
4,
19
97
, 2
00
0 &
20
01
48
00
58
-80
02
17
1
21
3.0
1
7.2
3
6.0
1
.5
14
38
Sa
ga
na
sh
1
99
9
49
02
28
-82
34
46
1
94
9.3
3
.4
10
.7
1.6
1
33
1
Sco
tt (
Lo
we
r)
20
01
4
80
82
0-8
11
52
0
42
.9
9.5
3
2.0
3
.8
14
12
Se
pa
ratio
n
20
01
4
80
35
8-8
03
42
9
18
1.4
2
.9
7.9
1
.8
14
13
Sh
en
an
go
2
00
1
48
23
00
-82
43
00
4
81
.0
No
da
ta
15
.0
2.1
1
33
6
Sh
ikw
am
kw
a
20
00
4
80
53
9-8
40
83
3
55
9.8
1
3.7
5
5.8
2
.4
12
62
Sid
eb
urn
ed
2
00
1
47
44
38
-83
31
08
1
15
5.4
1
.5
8.0
3
.3
13
13
Sin
cla
ir
20
00
4
75
12
5-8
12
05
1
94
7.1
9
.9
48
.8
3.4
1
44
9
Sis
se
ne
y
19
99
& 2
00
14
75
15
9-8
04
01
2
36
5.8
9
.4
36
.6
3.2
1
45
1
Ske
leto
n
19
94
4
75
15
4-7
93
83
7
35
6.7
3
.1
7.0
0
.6
14
55
Sm
ith
(W
alfo
rd)
19
98
4
61
30
5-8
22
03
5
15
0.0
N
o d
ata
N
o d
ata
3
.75
N
o d
ata
So
mm
e
20
00
4
74
02
6-8
15
73
5
66
.5
2.9
9
.0
1.9
1
42
7
Stu
mp
y
20
01
4
73
43
8-8
04
52
4
39
6.0
3
.2
6.7
4
.3
14
63
Su
ga
r 1
99
9
46
14
05
-82
13
42
8
6.2
9
.6
23
.8
5.0
1
63
9
NE
- 5
Ta
ble
NE
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Tem
agam
i3.
20
01
4
70
02
3-8
00
44
6
20
97
1.7
1
8.2
1
10
.0
10
.7
15
38
Th
istle
1
99
9
46
40
26
-80
01
77
2
42
.4
5.6
1
8.5
4
.1
15
98
Tim
iska
min
g
19
98
4
70
75
3-7
92
64
7
29
48
4.3
3
5.7
2
13
.5
0.9
1
58
7
To
mik
o
20
01
4
63
14
0-7
94
91
7
18
19
.5
7.9
2
4.4
3
.8
16
23
Up
pe
r C
ran
be
rry
19
99
4
61
62
2-8
25
60
6
47
.7
2.2
5
.5
1.9
1
68
9
Ve
rmili
on
1
99
6
46
31
22
-81
23
28
1
07
8.6
4
.7
13
.1
3.7
1
63
5
Wa
ba
ton
gu
sh
i 1
99
8 &
20
00
48
27
55
-84
13
37
3
79
1.0
7
.2
53
.4
2.9
1
23
2
Wa
bo
ose
kon
2
00
0
49
23
00
-86
12
00
N
o d
ata
N
o d
ata
N
o d
ata
N
o d
ata
12
33
Wa
ka
mi
19
98
& 2
00
14
72
92
2-8
25
10
3
17
13
.0
3.7
1
0.0
4
.4
13
83
Wa
rd
20
00
4
75
62
3-8
10
20
6
53
.4
3.7
9
.5
3.8
1
41
8
Wa
wa
go
sh
e
19
99
4
81
20
4-7
93
62
5
20
1.5
4
.1
7.3
1
.7
13
59
We
ne
be
go
n
19
97
4
72
34
2-8
30
55
6
27
21
.4
3.6
1
2.8
2
.1
13
94
We
st
Ka
be
nu
ng
1
99
9
48
15
38
-85
02
45
1
55
7.0
6
.4
21
.3
4.4
1
12
6
Wh
ite
1
99
9
48
46
10
-85
38
08
5
88
8.5
8
.7
48
.8
2.7
1
21
0
Wh
ite
fish
2
00
0
48
02
23
-84
27
11
1
82
1.1
1
5.2
5
4.9
3
.1
12
35
Wh
ite
wa
ter
19
96
4
63
20
0-8
10
83
1
94
3.7
2
.2
11
.3
2.9
1
64
8
Wh
itso
n
19
96
4
63
50
0-8
05
80
0
43
7.0
7
.3
16
.0
3.6
1
66
0
Wic
kste
ed
2
00
0
46
45
43
-79
40
54
1
31
2.8
4
.1
12
.2
2.1
1
56
9
Wiz
ard
2
00
0
47
44
08
-81
46
28
1
31
.5
2.3
7
.0
1.4
1
44
3
Wo
lf
19
99
4
73
81
4-8
15
91
4
10
4.7
2
.1
8.5
1
.9
14
34
Wo
wu
n
19
99
4
91
00
0-8
54
50
0
26
1.4
4
.6
11
.9
2.3
1
28
2
1.
Latitu
de a
nd longitude in d
egre
es,
min
ute
s,
and s
econds.
2.
GD
D>
5 is g
row
ing-d
egre
e-d
ays a
bove 5
o C
els
ius.
3.
Data
not
inclu
ded in t
he lake c
hara
cte
ristics s
um
mary
sta
tistics b
ecause o
nly
a s
mall
port
ion o
f th
e e
ntire
wate
rbody w
as s
am
ple
d.
NE
- 6
Ch
ara
cte
ris
tic
s o
f n
ort
he
as
t re
gio
n l
ak
es
sa
mp
led
us
ing
fa
ll w
all
eye
in
de
x n
ett
ing
sta
nd
ard
.
S
urf
ace
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
Me
an
2
96
2.8
6
.3
24
.9
3.0
1
41
9
Min
imu
m
42
.9
0.5
1
.5
0.3
1
12
6
Ma
xim
um
9
09
71
.7
35
.7
21
3.5
8
.4
17
04
Me
dia
n
51
2.8
4
.7
18
.0
2.9
1
42
8
5%
Pe
rce
ntile
5
6.9
1
.5
4.3
0
.9
11
69
25
% Q
ua
rtile
1
81
.4
3.1
9
.6
2.0
1
30
4
75
% Q
ua
rtile
1
44
7.6
8
.1
33
.5
3.8
1
49
0
95
% P
erc
en
tile
7
70
6.9
1
5.1
5
5.8
5
.0
16
85
Sa
mp
le S
ize
1
25
1
22
1
24
1
24
1
28
NE - 7
Table NE-2. Walleye relative abundance and average size for northeast region waterbodies sampled using the fall walleye index netting standard 1994 to 2001.
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
Abitibi (19953.
) 2.7 0.15 35 31 354 539
Abitibi (19963.
) 3.8 0.14 39 36 332 420
Abitibi (2001) 5.4 0.12 40 37 355 466
Agnew (19983.
) 1.5 0.19 12 10 309 367
Agnew (1999) 1.3 0.24 12 8 316 412
Amikougami 3.1 0.20 12 12 339 514
Anjigami 4.2 0.32 18 15 334 408
Barehead 0.3 0.12 12 4 617 3260
Bearhead 7.2 0.65 6 6 354 457
Birch 1.5 0.16 23 17 527 1807
Biscotasi (19993.
) 1.5 0.17 18 15 312 340
Biscotasi (2000) 2.2 0.10 38 33 372 585
Burnfield 4.9 0.36 6 6 306 424
Burntbush 6.2 0.23 12 12 356 461
Canoe 0 - 8 0 - -
Cataract 0.5 0.22 6 3 340 464
Crab’s Toe 8.3 0.13 12 12 308 292
Crouch 13.8 0.15 4 4 412 757
Departure 6.2 0.32 12 11 307 373
Deschenes 0.7 0.14 14 9 379 759
Dog 12.0 0.20 15 15 357 663
Duborne 3.1 0.30 13 11 360 883
Duncan 3.3 0.35 9 8 364 661
Echo 4.7 0.58 10 7 396 776
Fawcett 0.9 0.23 12 6 461 1072
Firth 3.5 0.22 6 6 408 783
Flanders 5.0 0.36 12 10 379 752
Fushimi 28.7 0.14 12 12 341 418
Garnham 16.2 0.38 12 11 360 514
Geneva 1.1 0.21 12 8 338 578
Gowganda 5.1 0.31 14 12 445 1288
Granitehill 25.8 0.20 7 7 396 735
Grassy 3.7 0.54 6 5 287 338
Groundhog 9.3 0.19 14 14 340 493
Hammer 7.5 0.23 12 12 336 436
Highbrush 17.2 0.08 12 12 380 655
Howells 20.1 0.20 6 6 351 475
Ivanhoe 19.7 0.27 10 10 365 673
Jowsey 14.4 0.14 12 12 343 459
Jumping Cariboo 4.2 0.35 12 11 298 280
Junction 4.6 0.24 6 6 356 557
Kabinakagami 17.4 0.11 18 18 316 364
Kagiano 6.4 0.29 18 17 326 400
NE - 8
Table NE-2. (continued)
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
Kebskwasheshi 23.5 0.12 9 9 352 563
Kecil 5.4 0.26 10 10 416 848
Kenetogami 2.6 0.30 12 9 376 656
Kenogaming 1.2 0.26 14 9 374 622
Kenogamissi 6.4 0.24 20 19 354 565
Knife 4.5 0.19 12 12 309 290
Kushog 8.2 0.21 12 12 330 396
Labitiche 1.6 0.26 8 7 439 999
Lac la Cave 5.8 0.25 18 17 357 546
LaMotte 0.2 0.09 12 3 533 1750
Larder 0.4 0.12 24 9 480 1748
Left 0 - 12 0 - -
Lloyd 0.1 0.06 12 1 725 4100
Longpoint 5.5 0.19 12 12 371 598
Lower Stetham 1.1 0.24 12 7 352 455
Macutagon 3.8 0.53 12 7 306 316
Manitou 2.0 0.27 10 7 429 1135
Marian 13.7 0.31 8 8 316 361
Mattagami 3.1 0.20 28 24 327 501
McArthur 6.7 0.28 12 11 300 262
McDiarmid 8.5 0.18 12 12 386 616
McPhail Reservoir 0.9 0.17 24 13 316 333
Mesomikenda 4.0 0.26 28 20 367 532
Mindemoya (19973.
) 23.6 0.27 10 10 319 332
Mindemoya (2001) 20.8 0.22 9 9 355 482
Minisinakwa 4.5 0.20 28 24 348 659
Ministic 2.9 0.26 12 11 365 567
Missinaibi 6.4 0.24 21 19 329 487
Mistinikon 2.7 0.33 12 11 374 759
Montreal River 0.2 0.11 24 3 311 264
Montrose 9.3 0.23 12 12 330 406
Mountain 8.5 0.12 6 6 322 374
Nabakwasi (19983.
) 5.4 0.29 14 13 357 464
Nabakwasi (2000) 1.2 0.21 14 9 300 359
Nagagami 21.7 0.18 12 12 342 489
Nagagamisis 10.9 0.40 12 11 305 343
Neville 4.8 0.28 12 11 289 325
Night Hawk 1.8 0.13 24 21 407 857
Nipissing (19983.
) 6.5 0.09 107 102 310 331
Nipissing (19993.
) 5.4 0.15 61 53 338 457
Nipissing (20003.
) 4.7 0.13 69 62 336 438
Nipissing (2001) 5.7 0.12 57 53 340 453
Nugent 2.8 0.30 12 10 331 546
Obushkong 7.2 0.18 12 12 392 599
NE - 9
Table NE-2. (continued)
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
Oke 4.0 0.20 12 11 297 274
Onaping (19993.
) 0.4 0.11 18 7 339 754
Onaping (2000) 1.1 0.12 22 16 340 534
Opeepeesway 6.9 0.18 17 17 314 378
Opikinimika 2.0 0.41 14 7 355 639
Oscar 0 - 7 0 - -
Papakomeka 3.5 0.31 12 10 348 506
Pivabiska 23.9 0.15 6 6 320 344
Popeye 8.0 0.42 6 6 433 1132
Post 5.3 0.23 8 8 384 589
Rabbit 2.4 0.22 26 19 381 622
Ramsay 0 - 12 0 - -
Ramsey 2.1 0.30 18 11 330 370
Red Cedar 2.2 0.32 14 10 357 661
Redwing 3.5 0.30 12 10 374 617
Rennie 6.3 0.18 12 12 370 674
Ritchie 4.6 0.32 12 10 291 263
Rocky Island 0.8 0.15 12 8 278 303
Round (19943.
) 16.0 1.03 6 5 402 899
Round (19973.
) 36.1 0.27 6 6 337 507
Round (20003.
) 6.5 0.12 12 12 347 597
Round (2001) 11.2 0.16 18 18 311 361
Saganash 15.9 0.15 12 12 371 644
Scott (Lower) 2.0 0.13 12 12 369 682
Separation 5.2 0.24 12 12 399 926
Shenango 10.8 0.14 16 16 416 795
Shikwamkwa 3.9 0.18 32 26 314 378
Sideburned 14.4 0.17 14 14 377 651
Sinclair 6.2 0.32 14 13 340 488
Sisseney (19993.
) 0.7 0.19 12 6 432 1155
Sisseney (2001) 5.9 0.15 12 12 365 550
Skeleton 1.9 0.20 20 15 310 304
Smith (Walford) 3.3 0.24 6 6 352 559
Somme 3.1 0.15 12 12 308 332
Stumpy 19.6 0.07 10 10 356 513
Sugar 6.4 0.39 6 6 374 859
Temagami 3.6 0.22 22 18 362 586
Thistle 3.7 0.34 12 10 333 423
Timiskaming 7.2 0.17 43 40 298 270
Tomiko 6.4 0.29 12 11 343 461
Upper Cranberry 0.3 0.22 6 2 514 1408
Vermilion 4.7 0.17 12 12 324 518
Wabatongushi (19983.
) 7.9 0.28 18 17 332 558
Wabatongushi (2000) 11.6 0.24 18 17 335 370
NE - 10
Table NE-2. (continued)
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
Waboosekon 4.7 0.35 12 12 363 561
Wakami (19983.
) 24.6 0.38 10 10 385 686
Wakami (2001) 22.5 0.22 8 8 339 636
Ward 0.5 0.13 12 6 553 1862
Wawagoshe 1.0 0.27 12 6 307 345
Wenebegon 11.1 0.12 22 22 357 534
West Kabenung 6.7 0.22 24 22 366 483
White 10.4 0.23 20 19 333 387
Whitefish 3.9 0.27 11 10 323 418
Whitewater 4.6 0.20 13 13 378 657
Whitson 0.5 0.13 16 8 367 585
Wicksteed 10.0 0.26 12 11 333 468
Wizard 3.8 0.22 12 12 311 492
Wolf 3.5 0.27 12 11 376 783
Wowun 6.9 0.38 12 11 376 670
1. Average catch is the geometric mean number of walleye caught net-1
.2. Average size is the arithmetic mean of total length in mm and wet weight in grams. 3. Data not used to calculate relative abundance or average size benchmarks.
Northeast region walleye relative abundance benchmarks.
Average Size AverageCatch Total
LengthWeight
Mean 6.4 362 641
Minimum 0 278 262
Maximum 28.7 725 4100
Median 4.6 354 533
5% Percentile 0.2 298 290
25% Quartile 2.2 326 406
75% Quartile 8.0 376 661
95% Percentile 20.8 480 1408
Sample Size 130 126
NE - 11
Table NE-3a. Annual female walleye mortality (%) for northeast region waterbodies sampled using the fall walleye index netting standard. “Observed mortality” is a direct estimate based on age structure of female fish and was calculated from age 5 onwards using Robson and Chapman’s estimator. “Estimated mortality” is an indirect estimate based on age structure of the population regardless of sex using the empirical formula: Female mortality = 0.952(Combined sex mortality) (n=246, R2=0.77, standard error of the estimate=6.493).
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Grassy No data 7.6% Burntbush 21% 20.0%
Sisseney (1999) No data 7.6% Nabakwasi (2000) 22% 20.0%
Wawagoshe No data 8.6% Mindemoya (2001) 21% 20.9%
Agnew (1998) No data 9.5% Biscotasi (1999) 11% 20.9%
Larder No data 12.4% Redwing 17% 20.9%
Geneva No data 12.4% Timiskaming 33% 21.9%
Agnew (1999) No data 12.4% Ritchie No data 21.9%
Round (2001) 11% 12.4% Deschenes No data 21.9%
Neville 14% 13.3% Saganash 25% 22.8%
Duncan 14% 13.3% Crouch 26% 22.8%
Gowganda 15% 13.3% Granitehill 23% 22.8%
Longpoint 14% 13.3% Missinaibi 20% 22.8%
Birch 12% 13.3% Ministic 20% 24.8%
Kebskwasheshi 17% 14.3% Mindemoya (1997) 22% 24.8%
Opikinimika No data 14.3% Onaping (2000) No data 24.8%
Popeye 12% 15.2% McDiarmid 25% 24.8%
Wizard 10% 15.2% Groundhog 24% 25.7%
Mistinikon 7% 15.2% Rocky Island No data 25.7%
Nugent No data 16.2% Marian 25% 25.7%
Kagiano 16% 16.2% Post No data 25.7%
Red Cedar 17% 16.2% Waboosekon 33% 25.7%
Obushkong 16% 16.2% Flanders No data 25.7%
Firth No data 16.2% Wakami (1998) 25% 25.7%
Dog 16% 17.1% Fawcett No data 26.7%
Rennie 21% 17.1% Oke 32% 26.7%
Night Hawk 14% 17.1% Labitiche No data 27.6%
Mattagami 15% 17.1% Temagami 26% 27.6%
Mountain No data 18.1% Wakami (2001) 34% 27.6%
Stumpy 20% 18.1% Biscotasi (2000) 36% 27.6%
Separation 19% 18.1% Wicksteed 27% 27.6%
Wabatongushi (1998) 17% 18.1% Kushog 21% 27.6%
Amikougami No data 19.0% Round (1994) 25% 28.6%
Howells 17% 19.0% Nagagami 34% 28.6%
Round (2000) 18% 19.0% Wenebegon 25% 28.6%
Junction No data 19.0% Macutagon 39% 29.5%
Kenogaming 14% 20.0% Nagagamisis 28% 29.5%
Vermilion 14% 20.0% Scott (Lower) No data 29.5%
Burnfield 11% 20.0% Garnham 26% 29.5%
Sinclair 19% 20.0% Ivanhoe 26% 30.5%
Montrose 21% 20.0% Duborne No data 30.5%
Minisinakwa 21% 20.0% Thistle 21% 30.5%
Departure 16% 20.0% Sideburned 27% 30.5%
NE - 12
Table NE-3a. (continued)
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Opeepeesway 20% 20.0% Shenango 34% 30.5%
Anjigami 29% 30.5% Sugar 36% 39.0%
Highbrush 28% 30.5% Pivabiska 45% 39.0%
Lac la Cave 40% 31.4% Bearhead 67% 40.0%
Crab’s Toe 37% 31.4% Wowun 33% 40.0%
Nabakwasi 38% 31.4% Nipissing (2000) 45% 40.9%
Jowsey 38% 31.4% Kecil 44% 40.9%
Mesomikenda 71% 31.4% McPhail Reservoir 29% 40.9%
Abitibi (2001) 30% 31.4% Hammer 43% 41.9%
Round (1997) 37% 32.4% Knife 35% 41.9%
Kabinakagami No data 32.4% Montreal River No data 42.8%
Kenogamissi 23% 33.3% Whitefish 45% 45.7%
West Kabenung 26% 33.3% Wabatongushi (2000) 45% 46.7%
Abitibi (1996) 22% 34.3% Papakomeka 40% 47.6%
Smith (Walford) No data 34.3% Nipissing (1999) 52% 48.6%
Sisseney (2001) 31% 36.2% Shikwamkwa 55% 49.5%
Abitibi (1995) 27% 36.2% Nipissing (1998) 57% 54.3%
Rabbit 32% 36.2% White 68% 55.2%
Tomiko 41% 37.1% McArthur 50% 57.1%
Fushimi 33% 37.1% Skeleton 54% 58.1%
Nipissing (2001) 40% 37.1% Echo No data 61.9%
Somme No data 38.1% Kenetogami No data 63.8%
Jumping Cariboo 36% 38.1% Whitewater 89% 90.4%
Northeast region female walleye mortality benchmarks.
Observed Mortality
EstimatedMortality
Mean 29% 27.9%
Minimum 7% 7.6%
Maximum 89% 90.4%
Median 26% 25.7%
5% Percentile 12% 12.4%
25% Quartile 19% 19.0%
75% Quartile 36% 33.3%
95% Percentile 55% 54.3%
Sample Size 101 130
NE - 13
Table NE-3b. Annual male walleye mortality (%) for northeast region waterbodies sampled using the fall walleye index netting standard. “Observed mortality” is a direct estimate based on age structure of male fish and was calculated from age 5 onwards using Robson and Chapman’s estimator. “Estimated mortality” is an indirect estimate based on age structure of the population regardless of sex using the empirical formula: Male mortality = 0.945(Combined sex mortality) + 2.844 (n=242, R2=0.65, standard error of the estimate=7.477).
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Grassy No data 10.4% Opeepeesway 19% 22.7%
Sisseney (1999) No data 10.4% Nabakwasi (2000) No data 22.7%
Wawagoshe No data 11.4% Mindemoya (2001) 23% 23.7%
Agnew (1998) No data 12.3% Biscotasi (1999) 26% 23.7%
Larder No data 15.1% Redwing 22% 23.7%
Geneva No data 15.1% Timiskaming 20% 24.6%
Agnew (1999) No data 15.1% Ritchie No data 24.6%
Round (2001) 15% 15.1% Deschenes No data 24.6%
Neville No data 16.1% Saganash 23% 25.5%
Duncan No data 16.1% Crouch 22% 25.5%
Gowganda 10% 16.1% Granitehill 25% 25.5%
Longpoint 13% 16.1% Missinaibi 29% 25.5%
Birch 14% 16.1% Ministic 24% 27.4%
Kebskwasheshi 13% 17.0% Mindemoya (1997) 27% 27.4%
Opikinimika No data 17.0% Onaping (2000) 29% 27.4%
Popeye 18% 18.0% McDiarmid 27% 27.4%
Wizard 40% 18.0% Groundhog 32% 28.4%
Mistinikon 25% 18.0% Rocky Island 15% 28.4%
Nugent No data 18.9% Marian 25% 28.4%
Kagiano 17% 18.9% Post 8% 28.4%
Red Cedar 15% 18.9% Waboosekon 17% 28.4%
Obushkong 19% 18.9% Flanders No data 28.4%
Firth 17% 18.9% Wakami (1998) 28% 28.4%
Dog 22% 19.9% Fawcett No data 29.3%
Rennie 15% 19.9% Oke 26% 29.3%
Night Hawk 30% 19.9% Labitiche No data 30.3%
Mattagami 18% 19.9% Temagami 33% 30.3%
Mountain No data 20.8% Wakami (2001) 21% 30.3%
Stumpy 18% 20.8% Biscotasi (2000) 26% 30.3%
Separation 20% 20.8% Wicksteed 29% 30.3%
Wabatongushi (1998) 23% 20.8% Kushog 32% 30.3%
Amikougami No data 21.8% Round (1994) 40% 31.2%
Howells 20% 21.8% Nagagami 26% 31.2%
Round (2000) 22% 21.8% Wenebegon 34% 31.2%
Junction 30% 21.8% Macutagon 26% 32.2%
Kenogaming 50% 22.7% Nagagamisis 34% 32.2%
Vermilion 50% 22.7% Scott (Lower) No data 32.2%
Burnfield 33% 22.7% Garnham 37% 32.2%
Sinclair 23% 22.7% Ivanhoe 40% 33.1%
Montrose 19% 22.7% Duborne No data 33.1%
Minisinakwa 21% 22.7% Thistle 38% 33.1%
Departure 27% 22.7% Sideburned 57% 33.1%
NE - 14
Table NE-3b. (continued)
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Burntbush 21% 22.7% Shenango 29% 33.1%
Anjigami 50% 33.1% Sugar 43% 41.6%
Highbrush 34% 33.1% Pivabiska 38% 41.6%
Lac la Cave 11% 34.1% Bearhead 25% 42.6%
Crab’s Toe 28% 34.1% Wowun No data 42.6%
Nabakwasi (1998) 29% 34.1% Nipissing (2000) 39% 43.5%
Jowsey 30% 34.1% Kecil 25% 43.5%
Mesomikenda 28% 34.1% McPhail Reservoir 46% 43.5%
Abitibi (2001) 35% 34.1% Hammer 40% 44.5%
Round (1997) 31% 35.0% Knife 50% 44.5%
Kabinakagami No data 35.0% Montreal River No data 45.4%
Kenogamissi 44% 35.9% Whitefish 45% 48.2%
West Kabenung 44% 35.9% Wabatongushi (2000) 54% 49.2%
Abitibi (1996) 43% 36.9% Papakomeka No data 50.1%
Smith (Walford) 36% 36.9% Nipissing (1999) 49% 51.1%
Sisseney (2001) 50% 38.8% Shikwamkwa 48% 52.0%
Abitibi (1995) 47% 38.8% Nipissing (1998) 57% 56.8%
Rabbit 41% 38.8% White 51% 57.7%
Tomiko 31% 39.7% McArthur 50% 59.6%
Fushimi 41% 39.7% Skeleton 66% 60.5%
Nipissing (2001) 35% 39.7% Echo No data 64.3%
Somme 40% 40.7% Kenetogami No data 66.2%
Jumping Cariboo 33% 40.7% Whitewater No data 92.7%
Northeast region male walleye mortality benchmarks.
Observed Mortality
EstimatedMortality
Mean 31% 30.6%
Minimum 8% 10.4%
Maximum 66% 92.7%
Median 29% 28.4%
5% Percentile 14% 15.1%
25% Quartile 22% 21.8%
75% Quartile 40% 35.9%
95% Percentile 50% 56.8%
Sample Size 102 130
NE - 15
Table NE-4a. Predicted female walleye weight-at-total length for northeast region waterbodies sampled using the fall walleye index netting standard. A geometric mean predictive regression model was fitted to logarithmically transformed (base 10) individual observations (only
lakes with 30 fish sampled). Weight in grams and total length in millimetres.
Weight-length relation1. Calculated weight at standard total lengthsLake
x10-6 350 400 450 500 550 Condition2.
Macutagon 2.98 8.83 337 501 712 975 1295 0.97
Abitibi (2001) 2.99 9.11 368 549 781 1070 1423 1.01
Minisinakwa 3.05 7.07 406 611 874 1206 1613 1.15
Round (2000) 3.05 6.71 386 579 830 1144 1530 1.08
Burntbush 3.10 4.74 365 552 796 1103 1482 1.00
Stumpy 3.12 4.43 384 582 840 1167 1572 1.02
Round (1994) 3.12 5.40 468 709 1024 1423 1916 1.28
Crab’s Toe 3.13 4.03 370 562 813 1130 1523 1.05
Jumping Cariboo 3.14 3.61 351 535 774 1077 1453 0.98
Timiskaming 3.14 3.65 355 540 782 1089 1469 0.98
Nagagami 3.14 3.68 358 545 789 1098 1481 0.98
Bearhead 3.15 3.47 358 546 791 1102 1488 0.96
Shenango 3.15 3.76 388 591 857 1194 1612 1.00
Groundhog 3.16 3.49 382 583 845 1179 1594 1.04
Missinaibi 3.18 2.89 356 544 791 1106 1497 0.97
Mattagami 3.18 3.40 418 640 930 1301 1761 1.12
Wabatongushi (2000) 3.18 2.86 352 538 783 1094 1482 0.99
Fushimi 3.18 3.34 411 628 914 1278 1730 1.10
Sisseney (2001) 3.18 2.85 351 536 780 1090 1476 0.99
Sideburned 3.19 3.07 401 613 893 1250 1694 1.10
Nipissing (2000) 3.19 3.16 412 631 919 1286 1744 1.12
Hammer 3.20 2.51 347 532 776 1087 1475 0.97
Marian 3.20 2.66 368 564 823 1152 1563 1.04
Kagiano 3.21 2.60 381 586 855 1199 1628 1.02
Tomiko 3.21 2.58 379 581 848 1189 1615 1.02
Kushog 3.21 2.47 362 556 812 1139 1546 0.99
Rabbit 3.21 2.58 379 581 848 1189 1615 1.02
Temagami 3.22 2.57 400 615 898 1261 1714 1.07
McDiarmid 3.22 2.54 395 607 888 1246 1694 1.08
Wakami (2001) 3.22 2.43 378 581 849 1192 1620 1.05
White 3.22 2.32 361 555 811 1138 1547 1.01
Wakami (1998) 3.22 2.50 389 598 874 1226 1667 1.08
Neville 3.23 2.40 396 609 891 1253 1704 1.10
Biscotasi (2000) 3.24 2.19 383 590 865 1216 1657 1.02
Gowganda 3.24 2.24 392 604 884 1244 1694 1.09
Wenebegon 3.24 2.07 362 558 817 1150 1566 0.97
McArthur 3.25 1.99 369 570 835 1176 1603 1.03
Nipissing (1999) 3.25 2.13 395 610 894 1259 1716 1.10
Nagagamisis 3.25 1.96 363 561 823 1159 1579 0.98
Lac la Cave 3.25 2.18 404 624 915 1289 1756 1.08
West Kabenung 3.25 1.96 363 561 823 1159 1579 0.99
Waboosekon 3.26 1.86 366 565 830 1170 1596 1.00
NE - 16
Table NE-4a. (continued)
Weight-length relation1. Calculated weight at standard total lengthsLake
x10-6 350 400 450 500 550 Condition2.
Ivanhoe 3.26 1.96 385 596 874 1233 1682 1.03
Saganash 3.26 1.94 381 590 866 1220 1665 1.02
Ramsey 3.26 1.87 368 568 834 1176 1605 1.04
Nabakwasi (1998) 3.26 1.71 336 520 763 1076 1468 0.95
Opeepeesway 3.27 1.83 382 590 868 1225 1673 1.04
Shikwamkwa 3.27 1.72 359 555 816 1151 1572 0.97
Highbrush 3.28 1.68 371 576 847 1197 1636 1.04
Anjigami 3.29 1.57 368 571 841 1190 1628 1.03
Howells 3.29 1.41 331 513 756 1069 1462 0.92
Nipissing (1998) 3.29 1.74 408 633 932 1319 1804 1.10
Montrose 3.29 1.62 380 589 868 1228 1680 1.04
Red Cedar 3.29 1.51 354 549 809 1144 1566 0.97
Vermilion 3.30 1.59 395 614 96 1282 1756 1.08
Granitehill 3.30 1.40 348 541 797 1129 1546 0.99
Night Hawk 3.30 1.57 390 606 894 1266 1734 1.06
Mindemoya (1997) 3.30 1.47 365 568 837 1186 1624 1.02
Sinclair 3.31 1.52 401 623 920 1304 1788 1.08
Mindemoya (2001) 3.32 1.26 352 549 811 1151 1579 0.98
Round (1997) 3.32 1.56 436 679 1004 1425 1955 1.16
Jowsey 3.32 1.34 374 583 863 1224 1679 1.03
Kecil 3.33 1.38 409 638 944 1341 1842 1.12
Wabatongushi (1998) 3.33 1.19 353 550 814 1156 1588 1.00
Garnham 3.34 1.10 346 540 800 1138 1564 0.95
Whitefish 3.34 1.24 390 609 902 1282 1763 1.05
Kebskwasheshi 3.34 1.21 380 594 880 1251 1720 1.04
Rennie 3.35 1.22 406 636 943 1342 1847 1.09
Nipissing (2001) 3.35 1.16 386 604 897 1276 1757 1.09
Kabinakagami 3.36 1.05 371 581 863 1229 1694 1.05
Dog 3.36 1.03 364 570 847 1206 1661 1.01
Wicksteed 3.36 1.08 381 598 888 1265 1742 1.03
Kenogamissi 3.36 1.12 396 620 920 1311 1806 1.09
Flanders 3.38 0.92 365 574 854 1220 1684 1.02
Whitewater 3.43 0.67 357 564 844 1212 1681 1.02
Mesomikenda 3.55 0.34 366 587 892 1297 1819 1.02
Popeye 3.58 0.28 359 579 882 1287 1810 1.07
Wowun 3.66 0.19 389 634 976 1435 2035 1.12
1. Two-tailed Student t-tests were employed to test significance of slope and intercept ( o: or = 0). 2. Condition calculated as the average of the individual ratios of observed weight-to-predicted weight from northeast region
median regression model (see below).
NE - 17
Northeast region female walleye weight-at-total length benchmarks.
Weight-length relation Calculated weight at standard total lengths
x10-6 350 400 450 500 550
Mean 3.25 2.43 377 582 854 1204 1642
Minimum 2.98 0.19 331 501 712 975 1295
Maximum 3.66 9.11 468 709 1024 1435 2035
Median 3.25 1.98 376 581 848 1198 1628
5% Percentile 3.05 0.67 346 532 774 1076 1462
25% Quartile 3.19 1.41 361 555 813 1144 1564
75% Quartile 3.31 2.86 390 607 892 1261 1720
95% Percentile 3.43 6.71 412 638 944 1342 1847
Sample Size 78
NE - 18
Table NE-4b. Predicted male walleye weight-at-total length for northeast region waterbodies sampled using the fall walleye index netting standard. A geometric mean predictive regression model was fitted to logarithmically transformed (base 10) individual observations (only
lakes with 30 fish sampled). Weight in grams and total length in millimetres.
Weight-length relation1. Calculated weight at standard total lengthsLake
x10-6 350 400 450 500 550 Condition2.
Round (1994) 2.81 35.37 498 725 1010 1358 1774 1.18
Minisinakwa 2.96 12.75 432 642 910 1243 1648 1.10
Abitibi (2001) 2.98 9.84 375 559 794 1086 1443 0.95
Mattagami 2.98 10.49 400 596 846 1158 1538 1.03
Stumpy 2.99 9.29 376 560 796 1091 1451 0.96
Abitibi (1995) 3.00 9.71 416 621 885 1214 1616 1.04
Round (2001) 3.03 7.10 363 544 777 1069 1427 0.98
Jumping Cariboo 3.07 542 350 528 757 1047 1403 0.94
Nabakwasi (1998) 3.07 5.60 362 545 783 1081 1449 0.91
Round (2000) 3.07 5.97 386 581 834 1153 1545 1.02
West Kabenung 3.10 4.75 366 553 797 1105 1485 0.95
Departure 3.11 4.51 368 558 805 1117 1502 0.94
Papakomeka 3.12 4.81 417 632 912 1267 1706 1.07
Tomiko 3.13 3.99 366 556 805 1119 1508 0.95
McDiarmid 3.13 4.41 405 615 889 1237 1666 1.06
Kecil 3.13 4.19 385 584 845 1175 1583 0.99
Shenango 3.14 3.83 373 567 821 1143 1541 0.94
Oke 3.14 4.09 398 606 877 1220 1646 1.04
Highbrush 3.15 3.77 389 593 859 1197 1616 1.01
Obushkong 3.15 3.60 372 566 820 1143 1543 0.95
Burntbush 3.16 3.36 368 561 814 1135 1534 0.97
Ramsey 3.17 3.15 366 558 811 1133 1532 0.92
Neville 3.18 3.29 405 619 900 1259 1704 1.02
Missinaibi 3.18 2.86 352 538 783 1094 1482 0.92
Opeepeesway 3.19 2.98 389 595 867 1213 1644 0.98
Saganash 3.19 3.02 394 603 879 1229 1666 0.99
Kagiano 3.19 2.93 382 585 852 1193 1617 0.97
Red Cedar 3.20 2.56 354 543 792 1109 1505 0.92
Wabatongushi (2000) 3.20 2.68 371 568 829 1161 1575 0.95
Wabatongushi (1998) 3.20 2.58 357 547 798 1118 1516 0.93
Nipissing (2000) 3.21 2.79 409 628 917 1286 1746 1.07
Wenebegon 3.21 2.47 362 556 812 1139 1546 0.94
Wakami (1998) 3.21 2.64 387 595 868 1217 1653 1.01
Waboosekon 3.21 2.58 379 581 848 1189 1615 0.97
Sisseney (2001) 3.21 2.41 354 543 792 1111 1509 0.91
McArthur 3.21 2.57 377 579 845 1185 1609 0.94
Timiskaming 3.21 2.49 365 561 818 1148 1559 0.94
Shikwamkwa 3.22 2.33 362 557 814 1143 1554 0.91
Whitefish 3.22 2.44 380 583 853 1197 1627 0.99
White 3.22 2.39 372 572 835 1172 1594 0.96
Nipissing (2001) 3.23 2.43 401 617 903 1268 1726 1.04
Anjigami 3.23 2.27 374 576 843 1185 1612 0.96
NE - 19
Table NE-4b. (continued)
Weight-length relation1. Calculated weight at standard total lengthsLake
x10-6 350 400 450 500 550 Condition2.
Echo 3.23 2.41 398 612 895 1258 1712 1.03
Macutagon 3.24 1.98 346 534 782 1100 1498 0.91
Marian 3.24 2.21 387 596 873 1228 1672 0.97
Biscotasi (2000) 3.24 2.21 387 596 873 1228 1672 0.98
Garnham 3.24 1.91 334 515 754 1061 1445 0.86
Kushog 3.24 2.11 369 569 833 1172 1596 0.94
Wizard 3.24 2.32 406 625 916 1289 1755 1.04
Longpoint 3.25 1.78 330 509 747 1052 1434 0.86
Lac la Cave 3.25 2.14 397 613 898 1265 1724 1.01
Kabinakagami 3.25 2.03 376 581 852 1200 1636 0.98
Crab’s Toe 3.25 2.10 389 601 881 1241 1692 1.00
Separation 3.25 2.24 415 641 940 1324 1805 1.07
Hammer 3.25 1.96 363 561 823 1159 1579 0.92
Granitehill 3.25 1.96 363 561 823 1159 1579 0.94
Sinclair 3.25 2.07 384 592 869 1224 1668 0.99
Mistinikon 3.27 1.91 398 616 906 1278 1746 0.99
Gowganda 3.27 1.92 400 619 911 1285 1755 1.02
Sideburned 3.27 1.90 396 613 901 1272 1737 1.03
Howells 3.28 1.59 352 545 802 1132 1548 0.88
Nipissing (1998) 3.28 1.84 407 630 928 1311 1791 1.05
Montrose 3.28 1.71 378 586 862 1218 1665 0.96
Mindemoya (1997) 3.29 1.56 366 567 836 1182 1618 0.96
Fushimi 3.29 1.73 406 629 927 1311 1794 1.04
Ivanhoe 3.29 1.63 382 593 873 1235 1690 0.99
Wicksteed 3.30 1.51 375 583 860 1218 1668 0.97
Wakami (2001) 3.30 1.61 400 622 917 1298 1778 0.99
Bearhead 3.30 1.38 343 533 786 1113 1524 0.87
Dog 3.30 1.44 358 556 820 1161 1591 0.94
Nipissing (1999) 3.30 1.63 405 629 929 1315 1800 1.03
Thistle 3.31 1.38 364 566 836 1184 1624 0.92
Rabbit 3.31 1.43 377 586 866 1227 1682 0.96
Jowsey 3.31 1.47 387 603 890 1262 1729 0.98
Mindemoya (2001) 3.32 1.29 360 562 830 1178 1617 0.94
Rennie 3.34 1.28 402 628 931 1324 1820 1.05
Temagami 3.35 1.21 43 631 936 1331 1832 1.00
Nagagamisis 3.36 1.01 357 559 830 1183 1629 0.92
Kebskwasheshi 3.36 1.11 392 614 912 1300 1790 0.97
Knife 3.36 1.07 378 592 879 1253 1726 0.92
Redwing 3.37 1.02 382 599 891 1271 1752 0.99
Kenogamissi 3.37 1.04 390 611 909 1296 1787 1.03
Nagagami 3.38 0.86 342 536 799 1140 1574 0.91
Whitewater 3.38 0.93 369 580 864 1233 1702 0.96
Mesomikenda 3.39 0.89 375 589 879 1256 1735 0.98
Pivabiska 3.41 0.82 388 612 915 1310 1813 0.99
Manitou 3.42 0.90 452 713 1067 1530 2120 1.22
NE - 20
Table NE-4b. (continued)
Weight-length relation1. Calculated weight at standard total lengthsLake
x10-6 350 400 450 500 550 Condition2.
Wowun 3.43 0.70 373 589 882 1266 1756 0.99
Round (1997) 3.44 0.80 452 715 1072 1540 2138 1.13
1. Two-tailed Student t-tests were employed to test significance of slope and intercept ( o: or = 0). 2. Condition calculated as the average of the individual ratios of observed weight-to-predicted weight from northeast region
median regression model (see below).
Northeast region male walleye weight-at-total length benchmarks.
Weight-length relation Calculated weight at standard total lengths
x10-6 350 400 450 500 550
Mean 3.22 3.22 382 588 860 1208 1643
Minimum 2.81 0.70 330 509 747 1047 1403
Maximum 3.44 35.37 498 725 1072 1540 2138
Median 3.24 2.24 378 585 859 1200 1629
5% Percentile 2.99 0.89 346 534 782 1081 1445
25% Quartile 3.18 1.59 366 560 818 1143 1546
75% Quartile 3.30 3.15 398 613 898 1265 1726
95% Percentile 3.39 9.71 417 641 936 1324 1813
Sample Size 89
NE - 21
Table NE-5a. Average female walleye total length-at-age (millimetres) data for northeast region waterbodies sampled using the fall walleye index netting method. Only
mean values with a minimum sample size 4 are included.
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Montreal River 342
Echo 522 567
Oke 333 350
Abitibi (1995) 387
Abitibi (1996) 425
Sisseney (1999) 413
Crab’s Toe 267 298 318 328 367 392
Skeleton 300
Kabinakagami 321 348 368 422 444
Departure 322 340 383
Night Hawk 332 390 459
Separation 340 428 469
Nugent 347
Waboosekon 347 395 460
Kenogaming 348
Duborne 365 418
Amikougami 373
Whitson 411
Wenebegon 238 297 324 371 459 466 531 513
Macutagon 248 284 357 423
Kenogamissi 250 323 333 388 453 560
Saganash 252 317 367 381 484
Burntbush 252 264 302 349 442
Vermilion 253 305 428
Wabatongushi (1998) 256 299 342 400 446 546
Garnham 259 310 367 412 466 477 603
McDiarmid 260 340 363 412 432 453
Knife 261 322 338
Ramsey 263 356 389
Abitibi (2001) 265 316 353 386 457 469
Kushog 266 316 366
Grassy 267
Wizard 271
Mountain 273 331 373
Opikinimika 273 355
Neville 274
Ritchie 274 320 361
Granitehilll 274 342 395 468 464 488 526 525
Whitewater 278 352 548
Nabakwasi (1998) 280 327 365 444 476
Agnew (1999) 283 345
Pivabiska 284 341 396 433
Red Cedar 285 335 359 480
NE - 22
Table NE-5a. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Crouch 286 426
Sisseney (2001) 288 350 534
Biscotasi (2000) 291 315 416 469
Fushimi 291 328 350 375 418 445
Mistinikon 291 359
McPhail Reservoir 293
Kagiano 293 363 432
Nagagamisis 295 326 380 434 550
McArthur 297
Longpoint 303 392
Mindemoya (2001) 312 373 391 447 422 554
Lower Stetham 315
Junction 318
Birch 323
Round (1994) 324 388 510 585
Anjigami 332 520
Duncam 332 359
Obushkong 338 380
Rabbit 341 419 483 525
Temagami 348 402 497 589
Manitou 490
Nagagami 191 278 333 386 402 478 558
Shikwamkwa 196 250 319 344 436 555
Timiskaming 198 240 273 324 358 395
Nabakwasi 200 260
Round 2000 205 277 337 418 488
Wabatongushi (2000) 207 273 316 349 397 431
Whitefish 210 312 481
Missinaibi 212 250 314 350 406 415
Mindemoya (1997) 218 275 340 376 430
Groundhog 218 287 343 424 490
Minisinakwa 219 287 331 389 442
Ivanhoe 221 287 335 412 461 558
Dog 223 288 354 420 510 551
Marian 224 327 385
Round (2001) 227 279 333 373
Thistle 228 299 337
Montrose 228 317 476
Wicksteed 229 306 376 507
White 231 305 340 410 458
Sinclair 232 303 365 461
Jowsey 233 309 406 444
Burnfield 233 333
Opeepeesway 236 302 372 455
Round (1997) 236 334 411 546 596
NE - 23
Table NE-5a. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Mattagami 237 267 323 372
Somme 237 299
Kebskwasheshi 239 322 367 443 449 456 544
Ministic 244 332
Wakami (2001) 244 346 443 507 564 577
Nipissing (1998) 245 299 348 409 447 483
Howells 248 308 372 434
Flanders 248 317 369 407 467 505 541
Tomiko 251 328 378 536
Lac la Cave 251 334 428 457 583
Highbrush 251 351 416
Post 252 395 447
Papakomeka 252 330 470
Stumpy 253 337 611
Geneva 255
West Kabenung 255 327 363 411 449 465
Nipissing (2000) 257 312 352 404 445 478 497 556
Wakami (1998) 258 337 401 470 503 577
Nipissing (2001) 259 326 367 415 471 487 538
Sideburned 260 352 436 509 561
Shenango 262 437 478 587 578
Mesomikenda 262 349 447 534
Hammer 262 369 539
Nipissing (1999) 263 321 364 419 461 516 552
Jumping Cariboo 270 341 385
Wawagoshe 272 339
Gowganda 276 354 419 477 643 693
Redwing 277
Wowun 277 330 382 483 517
Kenetogami 294 372 442 492
Sugar 312
Rennie 315 363 444
Kecil 315 374 458 479 598
Bearhead 319 370
Popeye 367
NE - 24
Northeast region female walleye total length-at-age benchmarks.
Average total length (mm) at age
1 2 3 4 5 6 7 8 9 10
Mean 247 303 356 400 442 471 495 531 517 553
Minimum 191 238 264 298 318 328 350 444 442 392
Maximum 367 490 458 522 567 589 643 596 560 693
Median 245 299 350 396 447 466 480 547 530 554
5% Percentile 198 250 297 322 349 333 367 444 442 392
25% Quartile 227 274 327 357 397 428 432 526 484 525
75% Quartile 262 330 378 428 476 516 577 558 556 603
95% Percentile 315 369 442 492 539 587 611 596 560 693
Sample Size 59 99 85 71 53 36 22 14 6 5
NE - 25
Table NE-5b. Average male walleye total length-at-age (millimetres) data for northeast region waterbodies sampled using the fall walleye index netting method. Only mean
values with a minimum sample size 4 are included.
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Abitibi (1995) 335 361 396 408
Mountain 366 401
Crouch 454 492
McDiarmid 360 354 421
Oke 289 306 320
Firth 443 481
Flanders 440 434
Saganash 326 352 365 396 428 411
Macutagon 288 359 431
Skeleton 296 345
Wabatongushi (1998) 308 331 345 389 420
Vermilion 310 349
Kabinakagami 319 350 366 400 428
Nabakwasi 320 359 411 424 414
Ritchie 327 345
Night Hawk 333 374
Nugent 345
Grassy 351
Kenogaming 359
Wowun 380 461
Amikougami 383
Round (1994) 386 437 476 497
Post 389 430
Burntbush 216 307 342 358 390
Crab’s Toe 216 261 293 311 329
Wenebegon 251 309 332 353 393 430 441
Garnham 252 310 367 381 413 436 464
Abitibi (1996) 253 358 379 402 417 405
Kushog 255 353 353 426
Onaping (2000) 257
Knife 258 281 314 329
Kenogamissi 260 322 331 361 408 421 473
Wizard 269 335
Whitewater 270 362 440 487
Granitehill 271 337 379 397 428 428 458
Departure 272 312 362 363
Waboosekon 273 344 383
Nagagamisis 276 332 362 396 446
Abitibi (2001) 278 317 349 375 423 438 452 451
McPhail Reservoir 284
Mistinikon 284 322 357 446
Nagagami 284 325 363 400 406 482
Ramsey 286 357 403
NE - 26
Table NE-5b. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Opikinimika 287 348
Sisseney (2001) 291 356 440
Somme 291 365
Longpoint 298 370
Scott (Lower) 301
Jowsey 308 360 384 400 422 436
Mindemoya (2001) 311 361 402 418 416 443
Howells 312 350 385 389 466 452
Junction 316
Rabbit 318 380 435 460
Anjigami 319
Birch 323 418 540
Tomiko 325 361 382
Ministic 332 390 501
Obushkong 332 392 464
Duborne 334 383 410
Round (1997) 334 393 481 524
Kenetogami 364 443
Echo 372 428 478 503
Manitou 463 546
Shikwamkwa 193 234 317 354 393 427 451
Wabatongushi (2000) 197 279 317 329 370 383
Round (2000) 200 294 315 403
Whitefish 205 288 445
Biscotasi (2000) 205 293 326 381 398 452 474 468
Timiskaming 206 224 275 315 340 364 385 408
Groundhog 208 297 346 418 439
Kagiano 209 292 356 404
Dog 210 294 349 402 461 472
Biscotasi (1999) 214 363
Missinaibi 214 263 292 341 372 400 450
Mindemoya (1997) 214 281 329 355 356 388 402 420
Neville 216 271
Marian 218 320 370 440
Ivanhoe 219 277 317 384 417 464
Red Cedar 224 278 333
Wicksteed 225 305 341 398 432 467
White 226 295 335 403 422
Minisinakwa 227 293 329 388 430
Sinclair 229 302 348 409
Mattagami 231 272 327 364 463 443
Thistle 234 273 321 371 368
Opeepeesway 234 306 367 403 495
Round (2001) 235 280 320 359
Kebskwasheshi 236 322 347 379 409
NE - 27
Table NE-5b. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Burnfield 238
Pivabiska 238 292 339 365 381 397
Wakami (2001) 239 338 412 464 508 525
Fushimi 240 290 323 351 365 354 398 391 425
Nipissing (1998) 241 300 343 373 388 409 417 452
Montrose 241 305 369
Nipissing (2001) 242 318 355 382 412 462
Highbrush 242 342 401 470 503
Temagami 244 339 412
Papakomeka 247 329 408
Duncan 251 387
Mesomikenda 252 350 408 451 486 506
Wakami (1998) 253 341 389 429 467 507
Separation 255 373 401
Nipissing (2000) 256 313 348 371 401 425 490
Stumpy 257 330 391 431 499
McArthur 259 295 342
Jumping Cariboo 259 337
West Kabenung 260 319 349 392 409 404 465
Nipissing (1999) 261 313 360 380 406 433 470
Shenango 262 412 448 503 513
Hammer 263 370
Sideburned 265 356 416
Redwing 271 362 425 469
Lac la Cave 277 340 368 390
Gowganda 279 346 409 445
Wawagoshe 280
Rennie 306 363
Bearhead 309 392
Kecil 312 368 422 466
Sugar 315 539
Popeye 367 496
NE - 28
Northeast region male walleye total length-at-age benchmarks.
Average total length (mm) at age
1 2 3 4 5 6 7 8 9 10
Mean 244 303 352 386 398 420 433 452 441 446
Minimum 193 216 261 293 289 306 320 391 385 408
Maximum 367 463 496 546 539 508 513 540 490 492
Median 240 297 348 380 396 423 430 451 444 448
5% Percentile 200 251 292 315 329 354 329 396 385 408
25% Quartile 219 278 323 356 361 396 402 422 420 411
75% Quartile 259 329 380 410 422 433 463 468 469 468
95% Percentile 312 368 416 464 487 506 507 525 490 492
Sample Size 57 89 91 76 54 37 29 24 14 6
NE - 29
Table NE-6a. Average female walleye weight-at-age (grams) data for northeast region waterbodies sampled using the fall walleye index netting method. Only mean
values with a minimum sample size 4 are included.
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Montreal River 313
Echo 1517 1963
Oke 355 386
Abitibi (1995) 536
Abitibi (1996) 680
Sisseney (1999) 697
Crab’s Toe 161 216 288 320 415 556
Skeleton 237
Departure 284 345 499
Kabinakagami 297 380 445 746 847
Night Hawk 323 526 911
Waboosekon 356 538 887
Kenogaming 358
Nugent 367
Separation 457 799 1033
Amikougami 489
Whitson 729
Duborne 865 1094
Wenebegon 106 223 290 470 865 891 1358 1252
Macutagon 120 197 375 609
Burntbush 130 150 243 365 758
Kenogamissi 130 331 352 565 1029 1970
Saganash 133 274 430 508 1100
Vermillion 137 278 936
Knife 140 381 319
Garnham 142 223 404 655 950 1019 2121
Wabatongushi (1998) 143 218 337 561 820 1603
Whitewater 144 419 1734
McDiarmid 155 365 461 672 790 932
Kushog 157 279 425
Ritchie 163 282 397
Granitehill 163 336 545 909 932 1074 1380 1512
Abitibi (2001) 164 281 375 484 815 848
Opikinimika 165 417
Grassy 168
Mountain 173 301 432
Nabakwasi (1998) 175 274 410 775 969
Agnew (1999) 177 387
Ramsey 182 432 534
Wizard 185
Neville 187
Red Cedar 189 306 411 1013
Crouch 193 661
NE - 30
Table NE-6a. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Pivabiska 194 359 620 803
Sisseney (2001) 195 357 1431
Biscotasi (2000) 207 267 727 1025
Mistinikon 213 439
McPhail Reservoir 215
Kagiano 215 423 748
McArthur 222
Fushimi 227 324 409 512 737 887
Longpoint 229 511
Nagagamisis 237 297 464 713 1574
Mindemoya (2001) 250 432 513 774 697 1714
Junction 274
Lower Stetham 288
Duncan 324 442
Obushkong 328 508
Anjigami 333 1368
Birch 336
Rabbit 354 681 1072 1401
Round (1994) 360 685 1555 2290
Temagami 391 666 1300 2158
Manitou 1464
Shikwamkwa 57 125 264 357 832 1685
Timiskaming 58 113 164 283 389 573
Nagagami 58 184 295 476 557 947 1578
Nabakwasi (2000) 69 155
Wabatongushi (2000) 70 167 265 355 572 709
Missinaibi 74 124 244 361 555 668
Mindemoya (1997) 77 177 337 476 756
Round (2000) 77 220 362 734 1113
Whitefish 78 281 1166
Dog 88 197 390 729 1309 1681
Groundhog 89 209 370 705 1063
Ivanhoe 93 199 328 611 910 1853
Thistle 94 222 324
Marian 95 309 509
Montrose 96 310 1070
Wicksteed 100 241 481 1286
White 100 243 339 621 925
Round (2001) 103 179 334 457
Jowsey 103 237 626 811
Sinclair 106 259 472 1011
Opeepeesway 108 237 469 908
Burnfield 108 322
Kebskwasheshi 111 299 479 872 947 920 1614
Howells 113 223 387 657
NE - 31
Table NE-6a. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Round (1997) 113 369 738 1863 2445
Minisinakwa 115 216 348 605 829
Somme 116 240
Ministic 116 299
Flanders 123 264 424 575 963 1199 1455
Wakami (2001) 124 382 863 1235 1790 1991
Mattagami 125 179 317 557
Nipissing (1998) 127 246 403 683 939 1166
Mesomikenda 128 381 848 1570
Tomiko 131 307 476 1485
West Kabenung 134 289 407 626 836 950
Post 136 531 869
Highbrush 136 371 683
Geneva 137
Lac la Cave 138 338 763 1001 2154
Stumpy 140 333 2197
Hammer 144 413 1436
Wakami (1998) 149 352 604 1064 1394 1941
Nipissing (2001) 151 315 470 716 1093 1207 1685
Shenango 155 751 1025 2019 1861
Papakomeka 155 338 1050
Sideburned 157 427 828 1743
Nipissing (2000) 158 290 419 650 936 1167 1244 1880
Nipissing (1999) 158 293 471 734 1014 1444 1950
Jumping Cariboo 158 317 463
Wawagoshe 166 335
Redwing 175
Wowun 182 319 512 1194 2056
Gowganda 183 409 732 1150 2863 3606
Kenetogami 250 625 963 1210
Bearhead 262 412
Kecil 274 507 1036 1207 2495
Rennie 288 443 917
Sugar 297
Popeye 429
NE - 32
Northeast region female walleye weight-at-age benchmarks.
Average weight (g) at age
1 2 3 4 5 6 7 8 9 10
Mean 136 263 439 645 874 1090 1318 1551 1427 1902
Minimum 57 106 150 216 288 313 386 847 758 556
Maximum 429 1464 1036 1517 1963 2158 2863 2445 1970 3606
Median 124 229 390 605 834 950 1046 1576 1428 1714
5% Percentile 58 125 218 281 365 320 415 847 758 556
25% Quartile 96 177 301 397 556 703 790 1358 1100 1512
75% Quartile 155 322 489 799 1052 1438 1941 1853 1880 2121
95% Percentile 288 427 863 1210 1570 2056 2495 2445 1970 3606
Sample Size 59 99 85 71 52 36 22 14 6 5
NE - 33
Table NE-6b. Average male walleye weight-at-age (grams) data for northeast region waterbodies sampled using the fall walleye index netting method. Only mean
values with a minimum sample size 4 are included.
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Firth 864 1114
Oke 220 268 309
Mountain 435 564
Saganash 307 396 455 567 744 661
Flanders 826 793
Crouch 950 1089
McDiarmid 478 427 757
Abitibi (1995) 359 451 626 690
Macutagon 205 396 707
Wabatongushi (1998) 231 302 357 502 684
Skeleton 252 358
Nabakwasi (1998) 263 377 621 695 648
Kabinakagami 280 388 450 611 750
Vermilion 298 424
Ritchie 302 360
Night Hawk 320 490
Grassy 381
Kenogaming 397
Nugent 424
Post 500 736
Amikougami 507
Wowun 510 996
Round (1994) 641 933 1175 1470
Burntbush 83 266 348 412 534
Crab’s Toe 84 152 210 273
Knife 125 174 267 307
Abitibi (1996) 126 389 486 506 604 645
Garnham 127 219 406 428 639 686 845
Wenebegon 128 256 332 403 550 704 786
Kushog 134 391 380 731
Onaping (2000) 139
Kenogamissi 151 311 350 463 688 789 1109
Departure 155 261 411 439
Granitehill 165 323 469 569 659 701 919
Nagagamisis 169 305 400 550 880
Waboosekon 173 351 531
Whitewater 175 446 851 1135
Wizard 178 362
Nagagami 183 274 390 546 611 987
Abitibi (2001) 188 280 372 469 731 725 841 775
Opikinimika 188 387
Longpoint 203 400
Sisseney (2001) 206 384 741
NE - 34
Table NE-6b. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Scott (Lower) 207
McPhail Reservoir 208
Ramsey 208 385 593
Mistinikon 211 307 404 869
Somme 219 456
Howells 239 340 483 484 871 802
Mindemoya (2001) 245 406 597 667 636 798
Jowsey 256 395 513 595 721 858
Junction 283
Rabbit 286 491 752 951
Anjigami 289
Tomiko 299 419 480
Birch 303 750 1642
Ministic 310 539 1157
Obushkong 316 530 850
Round (1997) 377 645 1299 1940
Duborne 478 849 900
Kenetogami 489 881
Echo 508 736 1100 1359
Manitou 1185 2169
Shikwamkwa 56 100 275 401 548 713 785
Wabatongushi (2000) 62 187 274 310 452 506
Kagiano 70 216 397 574
Biscotasi (2000) 70 223 304 474 634 934 1008 976
Biscotasi (1999) 71 392
Timiskaming 72 87 174 267 326 450 509 586
Missinaibi 72 146 213 341 458 565 819
Round (2000) 72 249 298 612
Whitefish 73 210 905
Dog 74 212 365 591 938 986
Groundhog 74 241 419 639 798
Mindemoya (1997) 77 187 308 407 415 533 605 714
Neville 84 178
Ivanhoe 88 166 289 527 689 1014
Red Cedar 90 174 320
Marian 90 291 458 794
Wicksteed 91 243 353 575 774 1046
White 96 222 323 609 696
Pivabiska 97 222 340 452 529 617
Thistle 100 158 282 445 455
Sinclair 103 250 381 660
Opeepeesway 107 259 442 618 1181
Burnfield 110
Kebskwasheshi 110 302 380 520 651
Round (2001) 111 191 291 421
NE - 35
Table NE-6b. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Wakami (2001) 112 346 674 1060 1338 1660
Montrose 113 241 475
Mesomikenda 119 389 665 873 1116 1283
Nipissing (1998) 120 245 432 513 587 679 698 930
Fushimi 121 220 319 416 476 456 581 610 769
Highbrush 121 373 622 1060 1308
Temagami 122 360 720
Mattagami 125 200 342 457 965 819
Minisinakwa 126 245 337 565 756
Nipissing (2001) 126 307 429 531 708 1068
Duncan 134 520
Wakami (1998) 138 370 567 747 1032 1339
Papakomeka 139 346 650
Jumping Cariboo 140 311
Hammer 144 446
Stumpy 147 321 529 734 1080
Separation 149 511 643
Shenango 149 613 811 1276 1262
West Kabenung 149 269 371 533 616 604 913
Nipissing (2000) 156 299 425 517 667 761 1185
Nipissing (1999) 127 278 443 541 665 824 1070
Redwing 159 428 751 1032
Sideburned 160 438 717
Wawagoshe 179
Gowganda 190 382 658 908
Lac la Cave 210 366 466 570
Bearhead 227 495
Rennie 261 453
McArthur 267 213 349
Kecil 298 424 705 1004
Sugar 329 1925
Popeye 434 1247
NE - 36
Northeast region male walleye weight-at-age benchmarks.
Average weight (g) at age
1 2 3 4 5 6 7 8 9 10
Mean 134 264 419 570 635 734 800 934 841 819
Minimum 56 83 152 210 220 268 309 567 509 586
Maximum 434 1185 1247 2169 1925 1338 1470 1940 1185 1089
Median 120 239 384 513 557 688 704 843 816 800
5% Percentile 70 125 213 267 307 412 332 604 509 586
25% Quartile 90 183 304 398 439 565 611 736 684 661
75% Quartile 149 310 500 655 708 774 965 986 1032 976
95% Percentile 298 478 720 1004 1175 1308 1339 1660 1185 1089
Sample Size 57 89 91 76 54 37 29 24 14 6
NE - 37
Table NE-7a. Empirical female walleye von Bertalanffy growth equation for northeast region waterbodies
sampled using the fall walleye index netting standard. L defines the asymptotic or maximum total length, K is the Brody growth coefficient and defines growth rate towards the maximum, and to shifts the growth curve along the axis to allow for apparent non-zero
body length at age zero (to=-1). Omega ( ) is early growth and corresponds to the growth rate near to. h is pre-maturation growth (i.e., growth to 350mm total length for combined sexes).
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Crab’s Toe 55 529 0.140 73.87 No data
White 164 569 0.211 119.78 83.90
Pivabiska 28 591 0.198 116.97 80.37
Macutagon 34 594 0.180 106.45 71.37
McDiarmid 70 606 0.156 94.12 64.62
Nipissing (1998) 361 610 0.206 125.46 84.97
Jowsey 57 616 0.175 108.02 93.98
Nabakwasi (1998) 66 620 0.182 113.07 74.87
Mesomikenda 116 631 0.306 193.07 116.18
Jumping Cariboo 51 634 0.215 136.06 108.05
Mindemoya (1997) 77 639 0.166 105.78 76.19
Biscotasi (2000) 41 640 0.166 106.15 76.52
Fushimi 100 651 0.132 85.79 70.52
Nipissing (2000) 235 660 0.184 121.29 86.36
Nipissing (2001) 171 676 0.182 122.92 93.17
Burntbush 45 678 0.109 74.02 54.31
Whitefish 77 679 0.253 171.31 No data
Wabatongushi (2000) 134 680 0.157 106.55 66.09
Kushog 46 683 0.157 107.04 72.64
Obushkong 29 686 0.163 111.62 105.81
Thistle 27 687 0.166 113.89 78.31
Timiskaming 229 689 0.118 81.17 58.33
Shikwamkwa 89 689 0.197 136.00 69.38
West Kabenung 110 690 0.159 109.40 91.67
Anjigami 81 692 0.203 140.10 No data
Round (1997) 90 696 0.196 136.23 106.11
Lac la Cave 74 701 0.174 122.11 96.38
Redwing 28 703 0.164 115.18 123.12
Rennie 48 706 0.214 151.20 126.85
Kabinakagami 68 710 0.113 79.88 66.85
Montrose 75 711 0.181 128.74 97.71
Papakomeka 27 713 0.207 147.48 No data
Bearhead 36 724 0.247 178.87 140.00
Sisseney (2001) 35 728 0.174 126.69 86.83
Nipissing (1999) 209 731 0.166 120.89 92.94
Whitewater 35 736 0.230 169.34 87.50
Hammer 62 741 0.199 147.72 124.08
Mindemoya (2001) 71 744 0.137 101.86 93.44
Kecil 38 748 0.203 152.09 133.00
Nagagamisis 49 749 0.135 101.08 76.92
NE - 38
Table NE-7a. (continued)
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Groundhog 62 751 0.141 106.09 86.07
Tomiko 41 752 0.193 145.30 97.50
Departure 25 754 0.129 97.18 71.96
Wakami (2001) 82 759 0.187 141.66 112.94
Wicksteed 56 762 0.152 115.37 90.05
Longpoint 27 772 0.179 137.73 96.31
Highbrush 84 773 0.190 146.66 114.17
Round (1994) 42 774 0.172 132.72 98.85
Sinclair 68 774 0.175 135.47 90.43
Mattagami 59 778 0.115 89.70 75.68
Shenango 78 779 0.180 140.41 No data
Wowun 64 779 0.163 126.81 102.82
Abitibi (2001) 123 782 0.113 88.24 70.41
Sugar 29 783 0.210 164.88 No data
Flanders 53 785 0.134 104.99 98.49
Kebskwasheshi 123 788 0.124 97.68 92.65
Temagami 61 792 0.189 149.46 112.62
Sideburned 104 794 0.163 129.54 118.80
Kagiano 100 795 0.159 126.78 90.79
Howells 44 802 0.129 103.79 92.29
Round (2001) 88 811 0.103 83.26 75.53
Stumpy 61 814 0.176 143.17 106.44
Vermilion 35 816 0.141 114.92 76.59
Wakami (1998) 146 817 0.154 125.65 108.29
Saganash 80 823 0.090 74.16 61.01
Nagagami 100 829 0.121 100.27 78.14
Kenogamissi 59 834 0.110 91.73 63.32
Missinaibi 102 838 0.105 87.93 68.66
Garnham 139 839 0.111 92.97 74.20
Granitehill 82 840 0.114 96.06 82.66
Wenebegon 112 840 0.105 87.94 62.10
Duncan 27 844 0.156 131.90 99.65
Night Hawk 33 845 0.110 92.67 77.95
Waboosekon 38 847 0.126 106.76 85.03
Minisinakwa 88 847 0.119 100.91 80.56
Separation 26 852 0.120 101.91 No data
Wabatongushi (1998) 96 858 0.097 83.50 65.43
Dog 117 864 0.125 107.60 88.21
Red Cedar 40 868 0.119 103.35 73.50
Round (2000) 48 896 0.128 114.30 82.06
Ivanhoe 130 897 0.111 99.89 81.04
Birch 26 897 0.115 102.80 106.27
Gowganda 67 957 0.133 127.12 116.67
NE - 39
Northeast region female walleye von Bertalanffy growth equation benchmarks.
L (mm) K (mm year –1) h (mm year–1)1.
Mean 748 0.159 116.99 89.78
Minimum 529 0.090 73.87 54.31
Maximum 957 0.306 193.07 185.90
Median 752 0.159 113.89 87.50
5% Percentile 606 0.105 81.17 61.97
25% Quartile 687 0.124 100.27 75.00
75% Quartile 816 0.184 135.47 99.45
95% Percentile 868 0.215 164.68 126.85
Sample Size 83 99
1. Pre-maturation growth benchmarks are calculated from all northeast lakes.
NE - 40
Table NE-7b. Empirical male walleye von Bertalanffy growth equation for northeast region waterbodies
sampled using the fall walleye index netting standard. L defines the asymptotic or maximum total length, K is the Brody growth coefficient and defines growth rate towards the maximum, and to shifts the growth curve along the axis to allow for apparent non-zero
body length at age zero (to=-1). Omega ( ) is early growth and corresponds to the growth rate near to. h is pre-maturation growth (i.e., growth to 350mm total length for combined sexes).
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Wabatongushi (2000) 142 492 0.219 107.43 66.09
Crab’s Toe 54 501 0.147 73.30 No data
Nipissing (1998) 465 507 0.235 119.06 84.97
Fushimi 246 508 0.187 94.74 70.52
McPhail Reservoir 28 508 0.216 109.84 No data
Pivabiska 92 527 0.211 111.23 80.37
Burntbush 46 536 0.147 78.75 54.31
Mindemoya (2001) 103 539 0.185 99.44 93.44
Jowsey 69 541 0.184 99.33 93.98
Mindemoya (1997) 210 544 0.168 91.14 76.19
Abitibi (1995) 102 550 0.176 96.77 64.81
West Kabenung 142 553 0.200 110.79 91.67
Nagagamisis 63 559 0.188 105.17 76.92
Vermilion 25 559 0.268 149.66 76.59
Kabinakagami 106 560 0.162 90.46 66.85
Sideburned 113 564 0.302 170.61 118.80
White 130 565 0.237 133.93 83.90
Tomiko 50 565 0.217 122.37 97.50
Nipissing (2000) 278 566 0.199 112.81 86.36
Sincalir 70 566 0.266 150.60 90.43
Departure 51 567 0.206 116.81 71.96
Waboosekon 59 568 0.203 115.43 85.03
McDiarmid 51 572 0.150 86.08 64.62
Abitibi (2001) 135 572 0.175 100.07 70.41
Nipissing (2001) 222 573 0.211 121.08 93.17
Nipissing (1999) 277 573 0.198 113.63 92.94
Wicksteed 80 581 0.178 103.28 90.05
Groundhog 74 582 0.202 117.44 86.07
Garnham 107 585 0.173 101.09 74.20
Separation 36 589 0.184 108.53 No data
Ivanhoe 112 592 0.181 106.95 81.04
Hammer 53 592 0.223 131.88 124.08
Wenebegon 139 593 0.164 97.17 62.10
Abitibi (1996) 96 594 0.137 80.96 61.97
Thistle 43 598 0.162 96.99 78.31
Rabbit 64 599 0.242 145.01 102.26
Kebskwasheshi 97 599 0.182 108.94 92.65
Kenogamissi 127 599 0.155 92.70 63.32
Timiskaming 220 600 0.116 69.45 58.33
Biscotasi (2000) 60 601 0.170 102.10 76.52
NE - 41
Table NE-7b. (continued)
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Montrose 64 606 0.219 132.36 97.71
Sisseney (2001) 31 608 0.200 121.56 86.83
Biscotasi (1999) 27 611 0.154 94.20 70.95
Round (2001) 115 611 0.191 116.74 75.53
Marian 81 612 0.192 117.39 99.45
Kagiano 96 613 0.201 123.51 90.79
Whitefish 65 614 0.244 149.61 No data
Shikwamkwa 82 616 0.179 110.29 69.38
Obushkong 44 618 0.178 109.92 105.81
Round (2000) 31 618 0.211 130.69 82.06
Kushog 44 619 0.167 102.91 72.64
Dog 105 620 0.171 106.05 88.21
Nabakwasi (1998) 47 621 0.154 95.74 74.87
Redwing 32 623 0.159 99.35 123.12
Temagami 63 626 0.258 161.27 112.62
Shenango 105 627 0.212 132.50 No data
Bearhead 46 627 0.237 148.68 140.00
Macutagon 38 630 0.152 95.39 71.37
Wabatongushi (1998) 50 635 0.114 72.41 65.43
Round (1997) 86 635 0.197 124.95 106.11
Echo 53 636 0.251 159.43 124.55
Howells 60 639 0.140 89.49 92.29
Mattagami 58 642 0.148 94.88 75.68
Mistinikon 46 642 0.148 94.78 75.00
Highbrush 105 644 0.196 126.43 114.17
Saganash 92 648 0.113 73.14 61.01
Red Cedar 38 650 0.156 101.35 73.50
Missinaibi 106 652 0.130 84.60 68.66
Crouch 28 653 0.157 102.64 No data
Duborne 24 654 0.152 99.32 86.15
Mesomikenda 111 655 0.201 131.73 116.18
Flanders 25 660 0.159 104.84 98.49
Wakami (1998) 184 660 0.185 121.67 108.29
Round (1994) 62 662 0.191 126.12 98.85
Stumpy 122 672 0.155 103.97 106.44
Ministic 27 686 0.178 122.10 106.06
Wakami (2001) 77 687 0.177 121.63 112.94
Opikinimika 29 689 0.142 97.99 86.02
Rennie 38 702 0.162 113.64 126.85
Nagagami 88 712 0.129 91.68 78.14
Minisinakwa 87 743 0.144 107.36 80.56
Granitehill 97 744 0.114 84.90 82.66
Longpoint 38 763 0.141 107.19 96.31
Popeye 28 772 0.199 153.55 185.90
Birch 28 809 0.129 104.08 106.27
NE - 42
Northeast region male walleye von Bertalanffy growth equation benchmarks.
L (mm) K (mm year –1) h (mm year–1)1.
Mean 612 0.182 110.67 89.78
Minimum 492 0.113 69.45 54.31
Maximum 809 0.302 170.61 185.90
Median 611 0.179 107.36 87.50
5% Percentile 508 0.129 78.75 61.97
25% Quartile 568 0.155 96.99 75.00
75% Quartile 642 0.202 121.67 99.45
95% Percentile 743 0.251 150.60 126.85
Sample Size 85 99
1. Pre-maturation growth benchmarks are calculated from all northeast lakes.
NE - 43
Table NE-8a. Female walleye age and size-at-maturity (total length in millimetres) schedules for northeast region waterbodies sampled using the fall walleye index netting standard. A logistic regression model was fitted to logarithmically transformed (base 10) maturity ogive (as percents).
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Sideburned 2.18 1.41 2.95 395.9 373.9 417.8
Popeye 2.41 1.47 3.35 502.3 469.4 535.2
Papakomeka 2.46 2.05 2.86 413.4 394.7 432.2
Bearhead 2.53 2.27 2.79 459.9 439.7 480.1
Kenetogami 2.60 1.85 3.35 391.9 375.7 408.2
Shenango 2.62 1.63 3.60 412.6 360.2 465.0
Wakami (2001) 2.87 2.17 3.56 407.6 386.4 428.7
Mindemoya (2001) 2.94 1.99 3.88 373.3 350.6 395.9
Obushkong 2.98 1.84 4.11 419.4 399.9 438.9
Redwing 3.01 2.46 3.55 438.1 417.9 458.3
Highbrush 3.04 2.34 3.73 420.1 379.3 460.9
Amikougami 3.11 2.36 3.85 416.3 385.0 447.7
Rabbit 3.14 2.31 3.98 441.7 416.2 467.1
Duborne 3.27 1.86 4.68 362.2 345.6 378.8
Sugar 3.37 2.92 3.81 439.7 415.9 463.5
Fawcett 3.41 2.98 3.84 440.1 425.7 454.4
Separation 3.44 2.02 4.86 418.6 390.0 447.1
Anjigami 3.50 3.05 3.95 440.4 423.6 457.3
Opeepesway 3.51 2.95 4.08 429.7 380.3 479.0
Wicksteed 3.54 2.86 4.21 425.9 377.9 474.0
Round (1997) 3.54 3.10 3.99 479.5 461.5 497.5
Montrose 3.57 2.74 4.39 419.0 400.4 437.6
Temagami 3.57 2.83 4.31 448.0 425.5 470.5
Wakami (1998) 3.58 2.74 4.43 448.9 422.7 475.2
Nugent 3.58 2.84 4.31 398.5 378.1 419.0
Mindemoya (1997) 3.65 2.86 4.44 369.0 321.5 416.6
Jowsey 3.65 3.12 4.17 379.9 361.4 398.5
Minisinakwa 3.67 2.49 4.85 359.9 325.3 394.5
Kecil 3.71 2.96 4.47 467.8 448.9 486.8
Mesomikenda 3.74 2.98 4.50 494.1 451.7 536.5
Birch 3.85 3.19 4.51 479.5 459.6 499.4
Rennie 3.87 3.46 4.28 472.1 455.7 488.5
Whitefish 3.87 3.57 4.18 476.4 454.6 498.2
Sinclair 3.93 3.26 4.60 455.8 434.2 477.4
Kebskwasheshi 3.98 3.14 4.83 417.8 396.7 438.9
Jumping Cariboo 4.00 3.60 4.40 420.1 397.7 442.4
Lac la Cave 4.03 3.40 4.66 425.8 349.3 502.4
Groundhog 4.07 3.53 4.62 426.3 407.7 444.8
Nipissing (2000) 4.11 3.03 5.19 406.1 366.0 446.2
Kagiano 4.13 3.27 4.98 465.0 441.3 488.8
Nabakwasi (1998) 4.13 3.25 5.00 378.9 343.7 414.1
NE - 44
Table NE-8a. (continued)
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Nipissing (1998) 4.13 3.31 4.95 413.5 376.7 450.3
Opikinimika 4.14 2.79 5.50 439.1 415.2 463.0
Nipissing (2001) 4.14 3.70 4.58 428.3 412.0 444.6
Night Hawk 4.16 2.67 5.66 405.9 382.2 429.5
Round (2000) 4.18 3.48 4.88 440.3 415.4 465.2
Gowganda 4.25 3.34 5.16 519.0 494.0 543.9
Nipissing (1999) 4.36 3.25 5.47 434.4 395.0 473.7
Sisseney (2001) 4.36 3.50 5.21 401.9 386.7 417.0
Crouch 4.51 4.10 4.92 434.6 415.6 453.5
Tomiko 4.52 4.23 4.81 478.5 458.3 498.7
Stumpy 4.53 3.88 5.17 499.7 478.5 520.8
West Kabenung 4.66 3.78 5.55 433.1 407.3 459.0
Hammer 4.68 4.21 5.15 504.2 475.6 532.7
Whitewater 4.77 4.52 5.03 495.0 468.3 521.6
Wowun 4.87 3.90 5.84 491.0 471.6 510.4
Granitehill 4.99 3.78 6.21 427.5 393.1 461.8
Waboosekon 5.00 4.17 5.82 457.1 406.2 508.1
Biscotasi (2000) 5.00 4.55 5.45 418.1 360.0 476.1
Dog 5.03 4.12 5.94 482.2 450.4 514.0
Flanders 5.03 4.29 5.76 475.1 450.9 499.2
Red Cedar 5.07 4.34 5.79 449.9 426.1 473.7
Shikwamkwa 5.14 4.70 5.57 489.4 465.3 513.6
Wenebegon 5.24 4.45 6.02 391.4 366.7 416.1
Abitibi (2001) 5.26 2.70 7.81 420.1 283.9 556.1
Timiskaming 5.29 4.26 6.31 389.4 324.0 454.8
Vermilion 5.37 4.14 6.60 460.7 434.9 486.5
Garnham 5.46 4.72 6.20 439.0 408.9 469.1
Kenogamissi 5.47 4.54 6.40 425.9 385.7 466.1
McDiarmid 5.50 4.78 6.22 394.5 372.4 416.7
Mattagami 5.52 4.36 6.67 418.5 375.2 461.8
Macutagon 5.55 4.62 6.48 419.7 402.4 436.9
Fushimi 5.61 4.93 6.29 414.3 381.3 447.2
Kabinakagami 5.74 4.64 6.84 385.9 353.3 418.5
Marian 5.92 4.37 7.47 585.9 494.6 677.2
Saganash 5.97 4.71 7.23 392.9 338.6 447.1
Oke 6.17 5.06 7.29 337.9 331.3 344.5
Wabatongushi (2000) 6.35 4.82 7.88 480.4 382.9 577.9
Howells 6.49 3.79 9.20 538.8 509.8 567.9
Missinaibi 6.59 5.81 7.38 486.8 454.1 519.6
Crab’s Toe 6.63 5.91 7.35 363.4 327.7 399.1
Ivanhoe 6.76 5.89 7.62 505.1 475.2 535.0
Wabatongushi (1998) 7.12 5.14 9.10 477.6 397.0 558.2
Nagagamisis 7.35 4.85 9.85 484.1 416.0 552.3
Nagagami 7.44 6.20 8.68 530.1 499.3 560.8
Round (2001) 8.76 7.49 10.03 539.8 505.9 573.7
NE - 45
Table NE-8a. (continued)
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Burntbush 9.06 8.32 9.80 406.4 375.0 437.9
Northeast region female walleye maturity benchmarks.
Maturity by age (years) Maturity by size (mm)
50% 10% 90% 50% 10% 90%
Mean 4.51 3.62 5.39 439.6 407.0 472.1
Minimum 2.18 1.41 2.79 337.9 283.9 344.5
Maximum 9.06 8.32 10.03 585.9 509.8 677.2
Median 4.14 3.46 4.98 433.1 402.4 465.0
5% Percentile 2.60 1.85 3.35 369.0 327.0 398.5
25% Quartile 3.57 2.83 4.28 412.6 375.7 438.9
75% Quartile 5.29 4.37 6.22 476.4 448.9 499.4
95% Percentile 7.12 5.89 9.10 519.0 494.0 560.8
Sample Size 87
NE - 46
Table NE-8b. Male walleye age and size-at-maturity (total length in millimetres) schedules for northeast region waterbodies sampled using the fall walleye index netting standard. A logistic regression model was fitted to logarithmically transformed (base 10) maturity ogive (as percents).
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Bearhead 1.26 0.56 1.95 329.1 276.8 381.3
Popeye 1.34 0.42 2.27 403.2 333.6 472.9
Rennie 1.41 0.34 2.48 321.5 303.0 340.0
Sugar 1.60 1.00 2.19 379.9 364.6 395.2
Hammer 1.67 1.25 2.09 340.0 327.4 352.6
Highbrush 1.79 0.94 2.64 331.6 310.0 353.2
Sideburned 1.80 1.28 2.32 338.4 322.6 354.2
Wakami (1998) 1.82 0.79 2.86 325.1 286.9 363.3
Echo 1.87 1.20 2.53 374.7 358.3 391.2
Jumping Cariboo 1.95 1.65 2.26 335.0 319.9 350.2
Ministic 1.96 0.89 3.03 338.2 316.0 360.4
Marian 2.00 1.11 2.90 311.7 232.3 391.2
Shenango 2.04 1.37 2.70 361.9 345.1 378.7
Longpoint 2.06 0.10 4.02 352.7 316.0 389.5
Nipissing (2001) 2.09 1.15 3.04 311.4 268.6 354.1
Tomiko 2.09 1.41 2.77 318.3 271.7 364.8
Mindemoya (2001) 2.12 0.80 3.44 320.4 283.9 356.9
Nipissing (1998) 2.12 0.97 3.26 300.8 246.9 354.6
Papakomeka 2.13 1.89 2.36 354.9 339.7 370.1
Wakami (2001) 2.16 1.24 3.08 366.3 336.9 395.7
West Kabenung 2.16 1.27 3.06 306.6 260.4 352.9
Anjigami 2.16 1.80 2.51 340.7 319.8 361.6
Nipissing (1999) 2.20 1.03 3.38 316.1 275.2 357.0
Nipissing (2000) 2.23 1.08 3.38 316.5 242.3 390.7
Whitefish 2.29 1.96 2.62 345.3 306.4 384.2
Montrose 2.30 1.38 3.21 341.5 326.8 356.2
Wicksteed 2.37 1.42 3.33 316.6 287.5 345.7
Smith (Walford) 2.38 1.97 2.79 399.6 380.7 418.4
Mattagami 2.40 0.95 3.84 298.7 247.4 350.1
Opeepeesway 2.40 1.73 3.07 337.0 316.4 357.6
Burnfield 2.41 2.05 2.77 359.8 345.5 374.1
Junction 2.44 1.96 2.92 361.3 351.1 371.6
Sinclair 2.46 1.75 3.16 332.5 310.1 354.8
Wowun 2.47 2.07 2.88 245.8 174.7 317.0
Somme 2.51 2.25 2.78 335.7 320.9 350.5
Kebskwasheshi 2.52 1.07 3.96 339.2 321.2 357.2
Temagami 2.52 1.61 3.43 372.8 330.6 415.0
Saganash 2.54 0.19 4.90 263.5 180.7 346.2
Post 2.55 1.88 3.22 368.5 353.1 383.9
Round (1997) 2.56 1.63 3.50 366.1 331.5 400.7
Mindemoya (1997) 2.57 1.71 3.43 307.1 266.7 347.5
Agnew (1998) 2.60 1.26 3.95 275.2 256.0 294.4
NE - 47
Table NE-8b. (continued)
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Wizard 2.75 2.22 3.27 324.9 294.6 355.1
Rabbit 2.80 1.94 3.66 366.2 313.0 419.5
Duncan 2.81 2.57 3.04 368.6 354.5 382.6
Groundhog 2.82 1.77 3.87 340.8 304.8 376.7
Fushimi 2.85 1.80 3.90 314.3 276.0 352.7
Pivabiska 2.89 1.73 4.04 329.5 273.8 385.1
Waboosekon 2.92 1.68 4.16 336.9 278.0 395.8
Sisseney (2001) 2.94 2.60 3.28 344.7 330.1 359.2
Opikinimika 2.98 1.94 4.03 364.5 333.8 395.2
McArthur 2.98 2.21 3.76 345.9 329.0 362.9
Wabatongushi (2000) 3.00 1.74 4.26 311.1 262.4 359.8
Vermilion 3.02 2.69 3.35 300.5 272.4 328.5
Minisinakwa 3.04 1.82 4.27 340.7 297.4 384.0
Dog 3.06 2.13 3.98 349.9 310.2 389.7
Kagiano 3.09 2.02 4.15 360.3 334.3 386.3
Round (2000) 3.14 2.39 3.89 337.1 320.2 354.1
Ramsey 3.20 2.69 3.70 381.7 322.4 441.1
Scott (Lower) 3.27 2.61 3.93 392.5 370.5 414.4
Ivanhoe 3.32 2.68 3.96 347.5 319.3 375.6
Wenebegon 3.32 2.25 4.39 313.2 288.3 338.1
Biscotasi (2000) 3.32 2.76 3.87 340.2 320.2 360.2
Duborne 3.37 0.30 6.43 366.7 350.2 383.1
Mistinikon 3.40 2.08 4.72 332.8 316.3 349.3
Timiskaming 3.41 1.77 5.04 285.3 245.9 324.6
Kabinakagami 3.41 2.30 4.52 306.0 242.9 369.2
Thistle 3.53 2.40 4.66 339.9 324.4 355.4
Knife 3.55 2.75 4.35 298.6 278.0 319.3
Kenogamissi 3.59 1.96 5.22 320.1 271.3 368.9
Red Cedar 3.62 1.88 5.36 345.7 329.1 362.2
Departure 3.62 2.65 4.59 335.7 291.1 380.4
Biscotasi (1999) 3.65 3.12 4.17 345.2 315.4 374.9
Kushog 3.66 2.45 4.88 336.4 285.3 387.5
Granitehill 3.70 2.55 4.85 364.6 319.9 409.2
White 3.70 2.96 4.44 387.3 363.9 410.7
Burntbush 3.78 2.74 4.82 295.3 281.1 309.6
Oke 3.81 3.14 4.48 276.9 241.0 312.8
Wabatongushi (1998) 3.82 3.04 4.59 313.7 283.6 343.8
Nagagamisis 3.95 2.40 5.51 362.6 322.0 403.3
Shikwamkwa 3.97 2.38 5.57 356.8 299.9 413.6
Nabakwasi (1998) 4.04 3.22 4.85 363.3 316.3 410.3
McPhail Reservoir 4.07 3.38 4.75 333.0 314.7 351.2
Round (2001) 4.08 2.30 5.87 365.3 295.1 435.5
Crab’s Toe 4.19 3.11 5.27 293.4 280.4 306.4
Larder 4.29 3.44 5.15 475.9 456.7 495.2
Nagagami 4.42 3.49 5.35 377.8 358.5 397.1
NE - 48
Table NE-8b. (continued)
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Abitibi (1996) 4.51 1.58 7.44 319.3 74.6 564.1
Macutagon 4.75 3.94 5.57 384.3 368.3 400.3
Missinaibi 4.84 3.48 6.20 368.6 342.8 394.4
Garnham 4.90 3.06 6.74 385.6 317.3 453.8
Northeast region male walleye maturity benchmarks.
Maturity by age (years) Maturity by size (mm)
50% 10% 90% 50% 10% 90%
Mean 2.90 1.92 3.87 339.7 303.9 375.4
Minimum 1.26 0.10 1.95 245.8 74.6 294.4
Maximum 4.90 3.94 7.44 475.9 456.7 564.1
Median 2.82 1.89 3.87 339.2 315.4 369.2
5% Percentile 1.67 0.42 2.27 285.3 241.0 317.0
25% Quartile 2.20 1.27 3.04 316.6 278.0 353.2
75% Quartile 3.55 2.55 4.59 363.3 330.1 394.4
95% Percentile 4.42 3.38 5.87 387.3 364.6 441.1
Sample Size 91
NE
- 4
9
Ta
ble
NE
-9.
Wa
lleye
re
lative
fe
cu
ndity (
eg
gs p
er
gra
m o
f to
tal
we
igh
t) d
ata
fo
r n
ort
he
ast
reg
ion w
ate
rbo
die
s s
am
ple
d u
sin
g t
he
fall
wa
lleye
in
de
x n
ett
ing
sta
nd
ard
. O
nly
me
an
va
lue
s w
ith
a m
inim
um
sa
mp
le s
ize
3
are
in
clu
de
d.
La
ke
M
ea
n
Sta
nd
ard
Err
or
Min
imu
m
Maxim
um
Media
n
5%
Pe
rce
ntile
25
%Q
ua
rtile
75
%Q
ua
rtile
95
%P
erc
en
tile
Sa
mp
leS
ize
Gra
nite
hill
2
6.3
6
1.0
0
17
.69
3
8.2
1
25
.49
1
8.8
0
23
.62
2
9.2
2
35
.62
2
7
Wa
ka
mi
(200
1)
30
.18
1
.25
1
4.8
3
46
.56
3
0.8
9
18
.06
2
4.3
5
34
.53
4
3.6
3
42
Bu
rntb
ush
3
0.4
4
1.6
0
21
.61
4
0.6
7
29
.15
2
2.3
2
28
.51
3
3.2
3
38
.57
1
2
Re
d C
ed
ar
31
.91
1
.47
2
2.3
8
42
.11
3
1.1
4
23
.47
2
8.5
9
35
.57
4
0.2
2
15
Sh
en
an
go
3
2.6
5
1.1
2
23
.08
8
0.7
1
31
.00
2
4.6
2
27
.90
3
5.3
6
40
.73
5
5
Sid
eb
urn
ed
3
2.7
3
0.7
6
13
.71
4
3.4
9
32
.28
2
2.2
7
28
.62
3
7.2
5
42
.01
6
7
Mis
tin
iko
n
32
.83
2
.64
2
2.9
9
38
.67
3
3.5
9
25
.07
3
3.4
0
35
.50
3
8.0
3
5
Ob
ush
ko
ng
3
3.0
9
2.1
6
23
.61
4
6.5
8
31
.21
2
5.6
5
30
.00
3
4.3
7
44
.81
1
0
Nip
issin
g(2
001)
34
.19
1
.56
1
5.8
1
61
.61
3
4.4
1
21
.62
2
7.5
9
38
.98
4
9.8
3
39
Ro
un
d(2
00
0)
35
.38
2
.34
1
7.7
2
48
.48
3
7.5
7
23
.08
3
1.5
6
39
.33
4
6.1
4
12
Ra
bb
it
35
.47
5
.54
2
.81
7
1.4
3
34
.33
4
.40
1
8.4
9
55
.56
6
4.3
1
16
Cra
b’s
To
e
36
.03
3
.45
2
4.8
3
60
.68
3
4.1
4
26
.69
3
0.0
1
35
.84
5
3.0
4
9
Ga
rnh
am
3
6.6
1
1.4
4
15
.24
5
5.1
7
36
.34
2
0.3
2
31
.51
4
3.1
4
48
.40
3
6
Nu
ge
nt
36
.75
2
.54
3
2.5
0
41
.27
3
6.4
7
32
.89
3
4.4
8
38
.87
4
0.7
9
3
Bis
co
tasi
(2000)
37
.17
6
.31
7
.44
7
6.6
7
39
.71
8
.46
1
4.3
2
56
.63
7
1.6
5
15
Te
ma
ga
mi
37
.77
2
.28
2
5.3
5
49
.79
3
6.5
1
26
.07
3
2.4
2
44
.01
4
8.9
1
13
To
mik
o
37
.92
2
.68
2
6.2
4
48
.80
3
6.5
4
28
.73
3
4.8
1
40
.69
4
8.7
0
8
Re
dw
ing
3
8.5
5
1.4
8
34
.58
4
4.4
7
37
.66
3
4.9
2
36
.20
4
0.3
2
43
.60
6
Du
nca
n
39
.95
1
.82
3
4.8
2
45
.22
3
9.5
3
35
.38
3
7.6
3
42
.53
4
4.6
8
5
Sh
ikw
am
kw
a
40
.07
1
.29
3
4.6
0
47
.83
3
7.4
1
34
.79
3
6.6
4
44
.64
4
6.7
4
13
Ha
mm
er
40
.82
3
.82
9
.80
5
1.3
8
42
.17
2
0.7
3
41
.47
4
8.3
8
51
.04
1
0
We
ne
be
go
n
40
.99
1
.70
1
9.0
4
70
.99
4
0.6
8
24
.74
3
6.7
6
44
.92
6
3.4
8
39
McD
iarm
id
41
.43
0
.92
2
6.3
5
49
.63
4
2.1
3
32
.36
3
8.1
1
46
.02
4
9.2
9
38
Wh
ite
wa
ter
41
.58
2
.14
2
7.7
0
46
.72
4
3.0
7
31
.98
4
0.9
5
45
.25
4
6.3
8
8
Nig
ht
Ha
wk
41
.69
1
.61
2
9.0
4
58
.08
4
1.2
8
31
.93
3
6.9
9
46
.97
4
9.5
0
19
Ro
un
d(1
99
7)
42
.94
1
.76
3
0.5
0
57
.26
4
1.7
6
35
.13
3
9.3
2
46
.17
5
4.0
9
14
Ka
gia
no
4
3.4
8
4.2
5
29
.92
6
8.7
2
42
.78
3
0.5
1
37
.49
4
4.8
8
61
.86
8
Sin
cla
ir
43
.88
1
.97
2
6.3
4
52
.53
4
5.9
8
32
.86
3
8.5
9
48
.67
5
2.4
4
14
Wo
wu
n
44
.89
3
.49
2
2.4
0
63
.60
4
5.5
6
23
.65
4
2.3
5
52
.08
6
0.0
6
12
Hig
hb
rush
4
4.9
4
3.0
8
27
.58
1
30
.09
4
1.1
2
28
.76
3
4.9
2
50
.59
6
1.6
0
34
Na
ga
ga
mi
45
.30
2
.07
3
5.6
0
68
.56
4
4.2
6
36
.47
4
1.6
7
46
.48
5
6.2
9
14
NE
- 5
0
Ta
ble
NE
-9.
(co
ntin
ue
d)
La
ke
M
ea
n
Sta
nd
ard
Err
or
Min
imu
m
Maxim
um
Media
n
5%
Pe
rce
ntile
25
%Q
ua
rtile
75
%Q
ua
rtile
95
%P
erc
en
tile
Sa
mp
leS
ize
Fla
nd
ers
4
5.5
1
1.3
3
37
.69
5
7.5
3
45
.74
3
8.1
6
40
.11
4
9.5
0
55
.72
2
0
Bu
rnfie
ld
45
.66
1
.04
4
3.5
8
46
.81
4
6.6
0
43
.89
4
5.0
9
46
.70
4
6.7
9
3
Ku
sh
og
4
5.7
4
3.2
7
33
.75
5
1.2
2
49
.07
3
5.7
8
43
.91
5
0.7
3
51
.12
5
Ma
cu
tag
on
4
7.0
4
16
.89
1
9.7
6
11
3.3
8
31
.42
2
1.9
6
30
.73
3
9.9
2
98
.69
5
Ju
nctio
n
47
.34
3
.39
4
2.0
8
53
.68
4
6.2
7
42
.50
4
4.1
7
49
.97
5
2.9
3
3
We
st
Ka
be
nu
ng
4
8.6
2
1.5
0
30
.84
6
1.0
7
47
.46
4
1.8
1
44
.68
5
2.4
6
59
.78
2
2
Piv
ab
iska
4
8.6
9
2.1
7
45
.10
5
6.8
1
46
.10
4
5.2
6
45
.89
4
9.5
7
55
.37
5
Wic
kste
ed
4
8.7
8
2.5
5
34
.96
6
0.9
8
49
.43
3
7.1
4
47
.23
5
2.3
5
58
.57
9
Na
ba
kw
asi
(200
0)
48
.92
4
.82
3
9.3
2
54
.48
5
2.9
5
40
.69
4
6.1
4
53
.71
5
4.3
3
3
Iva
nh
oe
4
8.9
2
1.8
8
30
.64
6
0.7
0
51
.14
3
4.9
9
41
.79
5
5.3
4
59
.64
2
1
Ke
no
ga
min
g
49
.20
4
.16
4
1.8
2
60
.54
4
7.2
2
42
.19
4
3.6
8
52
.74
5
8.9
8
4
Fir
th
49
.74
3
.68
4
4.0
6
63
.93
4
7.5
3
44
.08
4
4.1
6
49
.00
6
0.9
5
5
Sis
se
ne
y(1
999)
50
.26
1
.64
4
7.9
1
53
.41
4
9.4
5
48
.07
4
8.6
8
51
.43
5
3.0
2
3
Go
wg
an
da
5
0.5
7
1.3
7
32
.14
6
0.2
5
50
.81
4
4.0
4
46
.03
5
5.3
8
59
.01
2
2
Wa
bo
ose
kon
5
1.0
7
3.0
3
41
.25
7
6.2
3
48
.91
4
1.4
1
45
.12
5
1.4
8
68
.79
1
1
Mo
ntr
ose
5
2.6
2
2.7
9
42
.70
7
0.0
3
49
.11
4
3.1
1
45
.87
5
7.1
0
68
.50
1
1
Ke
bskw
ashe
sh
i 5
2.8
2
1.3
8
34
.01
6
8.5
7
53
.53
4
0.1
3
48
.45
5
9.3
7
66
.34
3
5
Re
nn
ie
53
.73
2
.00
4
8.5
6
62
.27
5
3.1
1
48
.99
5
0.5
6
54
.99
6
0.4
6
6
Th
istle
5
4.4
0
2.0
3
50
.43
6
0.4
8
52
.26
5
0.5
1
50
.85
5
7.9
8
59
.98
5
Wa
ka
mi
(199
8)
54
.61
1
.42
3
3.4
2
68
.61
5
5.5
3
37
.99
4
8.1
4
60
.97
6
6.5
2
42
Na
ga
ga
mis
is
54
.96
2
.79
4
4.1
1
81
.50
5
3.9
0
45
.38
4
9.3
3
55
.94
7
0.7
5
12
Ve
rmili
on
5
5.0
8
2.8
5
45
.85
6
4.8
5
54
.29
4
6.9
2
51
.07
5
9.5
0
63
.91
6
Ka
bin
aka
gam
i 5
5.2
9
1.1
5
45
.53
6
5.5
0
56
.69
4
5.8
7
49
.76
5
9.4
8
63
.78
2
9
Po
st
55
.72
4
.55
5
0.1
6
64
.73
5
2.2
6
50
.37
5
1.2
1
58
.50
6
3.4
8
3
Cro
uch
5
5.7
2
2.5
7
43
.69
6
7.8
9
52
.64
4
6.3
2
50
.70
6
2.7
0
67
.41
1
0
Fu
sh
imi
55
.79
1
.93
4
5.7
4
81
.79
5
5.5
5
45
.93
5
2.0
3
58
.54
6
3.6
7
17
De
sch
en
es
56
.54
0
.76
5
5.2
2
58
.62
5
6.1
6
55
.28
5
5.5
1
57
.20
5
8.3
3
4
Wa
ba
ton
gu
sh
i(2
000)
56
.65
2
.88
3
7.4
5
84
.90
5
6.0
5
41
.63
4
7.8
3
59
.83
7
7.7
5
18
Po
pe
ye
5
7.3
4
4.2
3
40
.69
7
5.5
9
56
.07
4
2.0
8
49
.78
6
6.9
5
72
.69
8
Be
arh
ea
d
57
.43
1
.07
5
3.5
0
61
.18
5
7.1
2
54
.23
5
6.4
6
58
.84
6
0.6
9
6
Nip
issin
g(1
998)
58
.30
1
.21
1
6.6
2
90
.28
5
9.3
3
42
.81
5
3.8
8
64
34
2
71
.79
7
5
NE
- 5
1
Ta
ble
NE
-9.
(co
ntin
ue
d)
La
ke
M
ea
n
Sta
nd
ard
Err
or
Min
imu
m
Maxim
um
Media
n
5%
Pe
rce
ntile
25
%Q
ua
rtile
75
%Q
ua
rtile
95
%P
erc
en
tile
Sa
mp
leS
ize
Jo
wse
y
58
.44
1
.58
4
0.0
8
87
.91
5
7.9
2
45
.11
5
2.7
6
63
.10
7
3.7
2
39
Wa
ba
ton
gu
sh
i(1
998)
58
.92
1
.54
4
1.6
3
74
.71
5
9.5
7
46
.16
5
2.0
6
64
.76
7
1.9
6
30
Min
de
mo
ya
(200
1)
59
.37
2
.25
3
8.1
2
81
.36
5
8.8
6
38
.35
4
9.0
3
66
.94
8
0.7
0
35
Ju
mp
ing
Ca
rib
oo
6
0.1
6
4.0
9
46
.88
7
4.8
4
59
.39
4
8.8
0
54
.58
6
5.4
7
72
.61
6
An
jiga
mi
60
.54
4
.27
3
6.0
4
79
.76
6
3.2
3
37
.31
5
9.9
5
65
.88
7
6.1
6
10
Op
ee
pe
esw
ay
60
.80
2
.93
4
8.3
5
72
.27
6
1.7
3
49
.68
5
1.7
6
69
.05
7
2.2
2
10
La
rde
r 6
1.3
5
3.5
3
53
.64
7
0.6
9
60
.53
5
4.5
3
58
.07
6
3.8
1
69
.31
4
La
c la
Ca
ve
6
1.8
1
2.4
4
38
.06
9
0.9
3
61
.99
4
1.6
6
51
.62
6
9.6
4
85
.41
3
0
Mis
sin
aib
i 6
2.5
0
1.3
4
54
.23
7
3.2
3
61
.48
5
5.4
2
59
.31
6
5.8
1
72
.27
1
6
Do
g
63
.87
1
.92
3
8.6
3
87
.12
6
3.8
3
46
.76
5
9.3
2
70
.23
8
0.4
2
29
Wh
ite
fish
6
3.9
3
6.3
6
43
.29
1
15
.47
6
0.0
5
43
.41
4
8.9
0
69
.96
9
9.3
3
11
Min
de
mo
ya
(199
7)
64
.54
2
.46
4
5.0
6
91
.27
6
2.8
8
47
.60
5
3.1
4
72
.88
8
2.3
3
25
Ke
cil
67
.42
3
.76
5
2.0
8
80
.10
6
9.6
5
52
.23
6
0.6
8
75
.64
7
8.9
0
8
Tim
iska
min
g
68
.95
2
.41
5
1.7
3
86
.39
6
9.2
7
52
.62
6
0.4
3
74
.78
8
4.7
5
18
Wh
ite
6
9.4
4
17
.88
4
8.9
3
12
2.9
8
52
.93
4
9.3
4
50
.98
7
1.4
0
11
2.6
6
4
Pa
pa
ko
me
ka
7
2.4
4
21
.83
3
6.2
1
24
.71
5
2.7
9
37
.16
4
1.9
0
63
.97
1
73
.65
9
Ho
we
lls
72
.65
1
.74
6
7.2
2
83
.27
7
2.0
3
67
.43
7
0.5
0
73
.53
8
0.0
7
8
Sa
ga
na
sh
7
5.5
2
1.5
7
52
.19
1
05
.06
7
6.4
7
56
.10
7
1.4
8
79
.84
9
0.6
3
45
Bir
ch
8
7.9
8
6.7
9
39
.91
1
65
60
7
9.9
7
59
.21
7
5.0
5
92
.03
1
44
.48
1
9
No
rth
ea
st
reg
ion
wa
lleye
re
lati
ve
fe
cu
nd
ity b
en
ch
ma
rks
.
Ave
rag
es
M
ea
n
Min
imu
mM
axim
um
Me
dia
n
5%
P
erc
en
tile
25
%Q
ua
rtile
75
%Q
ua
rtile
95
%P
erc
en
tile
Re
gio
n A
ve
rag
e1
.4
9.7
4
35
.07
7
0.0
2
48
.68
3
7.9
6
44
.06
5
3.9
1
63
.90
Re
gio
n A
ve
rag
e2
.4
8.9
4
31
.31
7
0.7
6
48
.50
3
5.3
9
42
.85
5
4.3
7
63
.62
1.
All
lakes c
om
bin
ed (
n=
81).
2.
Only
lakes w
ith
sam
ple
siz
e
10 (
n=
50).
NE
- 5
2
Ta
ble
NE
-10
. W
alle
ye
fe
cu
nd
ity-t
ota
l le
ng
th r
ela
tion
sh
ips f
or
no
rth
ea
st
reg
ion
wa
terb
od
ies s
am
ple
d u
sin
g t
he
fall
wa
lleye
in
de
x n
ett
ing
sta
nd
ard
. A
ge
om
etr
ic m
ea
n p
red
ictive
re
gre
ssio
n m
od
el w
as f
itte
d t
o
log
ari
thm
ically
tr
an
sfo
rme
d
(Na
pie
ria
n
log
ari
thm
s)
ind
ivid
ua
l o
bse
rva
tio
ns.
To
tal
len
gth
in
m
illim
etr
es.
Fe
cu
nd
ity-l
en
gth
re
latio
n1
.C
alc
ula
ted
fe
cu
nd
ity a
t sta
nd
ard
to
tal le
ng
th (
mm
)
La
ke
x 1
0-6
R2
n
40
0
45
0
50
0
55
0
60
0
65
0
Ro
un
d(2
00
0)
2.1
8
58
19
4.2
4
0.8
3
12
2
78
95
3
60
74
4
54
03
5
59
05
6
76
01
8
05
09
Jo
wse
y
2.6
3
52
67
.29
0
.64
3
9
35
98
1
49
02
6
64
65
7
83
04
9
10
43
73
1
28
79
4
Piv
ab
iska
2
.66
3
92
5.6
6
0.7
6
5
33
08
4
45
26
6
59
91
8
77
22
0
97
34
4
12
04
58
Mo
ntr
ose
2
.68
3
91
0.8
1
0.8
3
11
3
70
25
5
07
73
6
73
46
8
69
53
1
09
79
9
13
60
81
Sin
cla
ir
2.7
3
24
43
.38
0
.87
1
4
31
72
9
43
78
2
58
39
6
75
77
8
96
12
8
11
96
41
We
st
Ka
be
nu
ng
2
.76
2
02
5.7
2
0.8
7
22
3
03
41
4
19
85
5
61
40
7
30
16
9
28
17
1
15
74
1
Bu
rntb
ush
2
.78
1
02
1.4
5
0.7
9
12
1
74
10
2
41
53
3
23
69
4
21
86
5
37
27
6
71
12
Du
nca
n
2.8
7
96
9.6
8
0.7
6
5
28
89
3
40
52
5
54
84
7
72
11
9
92
59
7
11
65
33
Go
wg
an
da
2
.90
1
02
8.1
1
0.8
6
22
3
70
41
5
21
47
7
08
14
9
33
95
1
20
24
5
15
17
12
Fu
sh
imi
2.9
7
66
2.9
6
0.9
2
17
3
53
63
5
01
71
6
86
01
9
10
44
1
17
88
8
14
95
19
Sid
eb
urn
ed
3
.00
3
28
.14
0
.79
6
7
20
54
5
29
23
9
40
09
4
53
34
6
69
23
5
88
00
1
Ku
sh
og
3
.00
4
25
.84
0
.93
5
2
70
47
3
85
05
5
28
11
7
02
84
9
12
37
1
15
98
8
Wic
kste
ed
3
.08
3
05
.66
0
.83
9
3
10
49
4
46
12
6
16
96
8
27
23
1
08
12
0
13
83
16
Mis
sin
aib
i 3
.09
3
48
.54
0
.96
1
6
37
38
6
53
77
4
74
43
7
99
89
4
13
06
65
1
67
28
0
Tim
iska
min
g
3.1
1
29
5.4
9
0.9
2
18
3
64
62
5
25
90
7
29
77
9
81
52
1
28
64
9
16
50
07
Po
pe
ye
3
.12
3
18
.36
0
.92
8
4
26
14
6
15
61
8
55
49
1
15
21
0
15
11
85
1
94
12
3
Do
g
3.1
3
28
8.2
2
0.9
1
29
4
00
41
5
78
87
8
04
96
1
08
46
9
14
24
17
1
82
95
5
Sh
ikw
am
kw
a
3.1
3
17
4.2
6
0.7
7
13
2
42
94
3
51
24
4
88
45
6
58
22
8
64
27
1
11
03
3
Ju
mp
ing
Ca
rib
oo
3
.18
1
84
.76
0
.92
6
3
39
80
4
93
97
6
90
29
9
34
33
1
23
17
6
15
88
31
Nip
issin
g(1
998)
3.1
9
18
9.5
5
0.6
4
75
3
67
91
5
35
39
7
48
89
1
01
45
2
13
38
52
1
72
72
3
Ve
rmili
on
3
.20
1
68
.60
0
.94
6
3
52
78
5
14
14
7
20
11
9
76
70
1
29
00
2
16
66
31
Wa
ba
ton
gu
sh
i(1
998)
3.2
2
14
5.3
5
0.9
6
30
3
48
16
5
08
75
7
14
25
9
70
84
1
28
48
1
16
62
58
La
c la
Ca
ve
3
.24
1
42
.01
0
.77
3
0
37
15
6
54
38
8
76
47
7
10
40
97
1
37
93
8
17
87
07
Wo
wu
n
3.2
8
99
.66
6
0.9
0
12
3
32
59
4
89
17
6
90
79
9
43
92
1
25
52
1
16
31
49
Sa
ga
na
sh
3
.32
1
00
.89
0
.95
4
5
43
07
6
63
66
4
90
29
3
12
38
64
1
65
30
3
21
55
65
Ke
bskw
ashe
sh
i 3
.36
5
5.8
18
0
.91
3
5
30
83
3
45
80
1
65
25
5
89
88
3
12
04
03
1
57
55
4
Na
ga
ga
mis
is
3.3
8
46
.76
2
0.8
6
12
2
92
37
4
35
36
6
21
62
8
57
92
1
15
13
0
15
09
03
Gra
nite
hill
3
.40
2
0.4
20
0
.89
2
7
14
04
6
20
95
5
29
97
1
41
42
7
55
67
1
73
06
2
NE
- 5
3
Ta
ble
NE
-10
. (c
on
tin
ued
)
Fe
cu
nd
ity-l
en
gth
re
latio
n1
.C
alc
ula
ted
fe
cu
nd
ity a
t sta
nd
ard
to
tal le
ng
th (
mm
)
La
ke
x 1
0-6
R2
n
40
0
45
0
50
0
55
0
60
0
65
0
Nig
ht
Ha
wk
3.4
2
30
.21
3
0.8
9
19
2
37
77
3
55
66
5
09
88
7
06
29
9
50
99
1
25
03
1
Ga
rnh
am
3
.43
2
2.7
39
0
.85
3
6
19
36
9
29
01
8
41
66
0
57
78
0
77
88
7
10
25
11
Wa
ba
ton
gu
sh
i(2
000)
3.4
3
35
.37
8
0.8
9
18
3
04
29
4
55
96
6
54
69
9
08
17
1
22
43
9
16
11
69
Ro
un
d(1
99
7)
3.4
8
23
.78
8
0.7
0
14
2
64
38
3
98
17
5
74
30
7
99
90
1
08
24
4
14
29
73
Sh
en
an
go
3
.48
1
5.4
12
0
.86
5
5
17
20
2
25
90
9
37
37
2
52
05
6
70
44
8
93
05
5
Fir
th
3.5
0
20
.97
9
0.7
0
5
26
56
9
40
11
5
57
99
4
80
94
3
10
97
42
1
45
20
4
McD
iarm
id
3.5
1
18
.42
3
0.6
9
38
2
44
15
3
68
97
5
33
84
7
45
63
1
01
15
9
13
39
28
Ho
we
lls
3.6
2
14
.01
2
0.9
8
8
35
95
8
55
05
1
80
58
0
11
37
39
1
55
79
6
20
80
94
Ka
bin
aka
gam
i 3
.62
1
2.1
89
0
.97
2
9
31
41
9
48
10
7
70
42
1
99
40
6
13
61
72
1
81
89
3
Wa
bo
ose
kon
3
.62
1
0.0
52
0
.92
1
1
25
68
2
39
31
5
57
54
2
81
21
5
11
12
39
1
48
57
1
Wa
ka
mi
(199
8)
3.6
5
9.3
06
0
.94
4
2
29
88
3
45
95
3
67
52
8
95
65
7
13
14
55
1
76
10
9
Iva
nh
oe
3
.73
4
.71
2
0.9
5
21
2
41
18
3
74
29
5
54
56
7
91
40
1
09
49
5
14
76
06
Re
nn
ie
3.7
5
5.1
52
0
.99
6
2
95
25
4
59
23
6
81
75
9
74
67
1
35
07
4
18
23
63
Hig
hb
rush
3
.78
3
.25
4
0.8
2
34
2
28
97
3
57
58
5
32
77
7
64
17
1
06
21
8
14
37
99
Ka
gia
no
3
.84
1
.91
5
0.8
3
8
18
24
8
28
66
7
42
93
9
61
88
7
86
40
1
11
74
44
We
ne
be
go
n
3.8
4
1.8
21
0
.88
3
9
18
23
6
28
67
6
42
99
2
62
01
1
86
63
8
11
78
44
To
mik
o
3.8
4
1.7
79
0
.89
8
1
77
0
27
83
1
41
72
0
60
17
1
84
05
8
11
43
26
Th
istle
3
.88
2
.19
0
0.9
8
5
27
47
7
43
40
0
65
32
4
94
56
2
13
25
48
1
80
83
5
Re
d C
ed
ar
3.8
8
1.0
93
0
.91
1
5
13
86
3
21
90
2
32
97
2
47
73
8
66
92
5
91
32
0
Ob
ush
ko
ng
3
.89
1
.25
2
0.8
6
10
1
63
53
2
58
51
3
89
38
5
64
02
7
91
05
1
07
98
2
Na
ga
ga
mi
3.9
3
1.1
24
0
.95
1
4
19
04
3
30
25
7
45
78
3
66
59
2
93
75
0
12
84
18
Re
dw
ing
3
.93
1
.07
2
0.9
2
6
18
58
5
29
54
3
44
72
0
65
06
9
91
63
6
12
55
61
Min
de
mo
ya
(199
7)
3.9
4
2.0
26
0
.91
2
5
36
53
6
58
12
1
88
04
3
12
81
87
1
80
63
0
24
76
32
Te
ma
ga
mi
4.0
0
0.6
69
0
.93
1
3
17
01
7
27
25
5
41
53
7
60
80
9
86
11
7
11
86
05
Min
de
mo
ya
(200
1)
4.0
2
1.0
29
0
.92
3
5
29
26
6
46
97
5
71
72
9
10
51
95
1
49
21
4
20
58
11
Fla
nd
ers
4
.14
0
.35
2
0.9
2
20
2
05
73
3
34
93
5
17
97
7
68
39
1
10
14
2
15
33
89
Nip
issin
g(2
001)
4.1
7
0.2
48
0
.75
3
9
17
47
6
28
55
7
44
30
9
65
92
8
94
75
9
13
22
98
Wa
ka
mi
(200
1)
4.3
7
0.0
50
8
0.7
7
42
1
21
75
2
03
78
3
23
04
4
90
09
7
17
02
1
01
75
4
Op
ee
pe
esw
ay
4.4
1
0.1
03
0
.95
1
0
30
23
8
50
81
7
88
52
1
23
06
4
18
05
86
2
56
98
0
Ke
cil
4.6
2
0.0
26
2
0.7
6
8
27
51
4
47
41
1
77
14
2
11
98
21
1
79
11
1
25
92
57
Cra
b’s
To
e
4.7
7
0.0
08
97
0
.83
9
2
30
07
4
03
47
6
66
84
1
05
05
7
15
90
87
2
33
03
1
NE
- 5
4
Ta
ble
NE
-10
. (c
on
tin
ued
)
Fe
cu
nd
ity-l
en
gth
re
latio
n1
.C
alc
ula
ted
fe
cu
nd
ity a
t sta
nd
ard
to
tal le
ng
th (
mm
)
La
ke
x 1
0-6
R2
n
40
0
45
0
50
0
55
0
60
0
65
0
Wh
ite
fish
4
.85
0
.00
56
6
0.7
4
11
2
41
86
4
28
41
7
14
45
1
13
47
3
17
31
09
2
55
30
4
1. T
wo-t
aile
d S
tudent
t-te
sts
were
em
plo
yed t
o t
est
sig
nific
ance o
f slo
pe a
nd inte
rcept
(o:
or
= 0
).
No
rth
ea
st
reg
ion
wa
lleye
fe
cu
nd
ity-
tota
l le
ng
th-f
ec
un
dit
y b
en
ch
ma
rks
.
F
ecu
nd
ity-l
en
gth
re
latio
n
Ca
lcu
late
d fe
cu
nd
ity a
t sta
nd
ard
to
tal le
ng
th (
mm
)
x 1
0-6
R2
n
40
0
45
0
50
0
55
0
60
0
65
0
Me
an
3
.48
1
38
8.8
6
0.8
6
21
2
76
17
4
13
52
5
95
42
8
30
42
1
12
78
4
14
97
85
Min
um
um
2
.18
0
.00
56
6
0.6
4
5
12
17
5
20
37
8
29
97
1
41
42
8
53
72
8
67
11
3
Ma
xim
um
4
.85
5
81
94
.24
0
.99
7
5
43
07
6
63
66
4
90
29
4
12
81
88
1
80
63
0
25
92
57
Me
dia
n
3.4
3
23
.26
3
0.8
9
14
2
81
37
4
25
21
6
00
52
8
19
69
1
09
77
1
14
54
39
5%
Pe
rce
ntile
2
.67
0
.03
85
0
.70
5
1
52
00
2
30
28
3
26
71
4
83
74
6
72
64
8
42
55
25
% Q
ua
rtile
3
.12
1
.86
8
0.8
1
9
20
55
9
31
87
6
45
59
4
65
87
6
91
43
7
11
76
45
75
% Q
ua
rtile
3
.86
3
00
.58
0
.92
3
0
34
39
9
49
78
4
71
12
0
97
91
1
13
20
02
1
74
41
6
95
% P
erc
en
tile
4
.62
3
92
.56
6
0.9
8
55
4
00
42
5
81
22
8
55
49
1
23
06
4
17
91
12
2
55
30
4
Sa
mp
le S
ize
6
0
NE
-5
5
Ta
ble
NE
-11
a.
Ma
ture
fe
ma
le w
alle
ye
fa
ll g
on
ad
oso
ma
tic i
nd
ex (
GS
I is
ova
ry w
eig
ht
exp
resse
d a
s a
pe
rce
nt
of
the
to
tal
we
igh
t m
inu
s
th
e o
va
ry w
eig
ht)
, vis
cera
l fa
t in
de
x (
VF
I is
fa
t w
eig
ht
exp
resse
d a
s a
pe
rce
nt
of
the t
ota
l w
eig
ht)
, a
nd
re
pro
du
ctive
in
de
x
(R
is 3
h/L
) d
ata
fo
r n
ort
he
ast
reg
ion
wa
terb
od
ies s
am
ple
d u
sin
g t
he
fa
ll w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
. O
nly
me
an
va
lue
s w
ith
a m
inim
um
sa
mp
le s
ize
3
are
in
clu
de
d.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Lo
ng
po
int
1.5
3%
0
.48
%
2.3
7%
5
2
.82
%
0.9
0%
4
.52
%
5
0.3
7
Gra
nite
hill
1
.81
%
0.4
4%
3
.42
%
41
2
.70
%
0%
5
.89
%
40
0
.30
Oke
1
.91
%
1.3
1%
2
.71
%
9
0.2
8%
0
%
0.5
0%
9
N
o d
ata
Ga
rnh
am
1
.98
%
0.3
0%
7
.19
%
38
2
.91
%
0.3
9%
9
.70
%
38
0
.27
Wa
ka
mi
(200
1)
2.1
0%
0
.72
%
3.5
1%
4
2
3.9
7%
1
.69
%
6.9
9%
4
2
0.4
5
Ma
cu
tag
on
2
.11
%
1.6
3%
2
.78
%
9
1.4
8%
0
.80
%
2.4
2%
9
0
.36
Tim
iska
min
g
2.2
4%
1
.16
%
3.8
0%
1
8
1.1
6%
0
.32
%
2.9
8%
1
8
0.2
5
Ro
un
d(2
00
1)
2.2
9%
0
.24
%
5.3
0%
8
0
.92
%
0.1
2%
2
.54
%
8
0.2
8
Ob
ush
ko
ng
2
.57
%
0.1
6%
5
.01
%
12
1
.38
%
0%
2
.74
%
12
0
.46
Min
de
mo
ya
(200
1)
2.6
2%
0
.60
%
4.7
1%
3
6
2.3
7%
0
.11
%
5.7
9%
3
6
0.3
8
Po
st
2.7
7%
2
.14
%
3.1
0%
3
1
.18
%
0.1
5%
2
.10
%
3
No
da
ta
McD
iarm
id
2.9
1%
1
.66
%
4.2
8%
3
8
0.7
4%
0
%
2.0
1%
3
8
0.3
2
Nu
ge
nt
3.0
5%
2
.07
%
4.1
5%
3
2
.21
%
1.0
9%
3
.60
%
3
No
da
ta
Ku
sh
og
3
.06
%
2.0
8%
4
.29
%
5
2.4
0%
1
.07
%
3.4
9%
5
0
.32
Sa
ga
na
sh
3
.10
%
0.0
4%
5
.28
%
46
1
.87
%
0.2
5%
6
.97
%
46
0
.22
Ka
bin
aka
gam
i 3
.12
%
0.6
7%
5
.21
%
29
1
.53
%
0.7
1%
2
.82
%
8
0.2
8
Wh
ite
wa
ter
3.1
4%
2
.55
%
4.1
9%
8
1
.92
%
0.8
8%
2
.29
%
8
0.3
6
Piv
ab
iska
3
.19
%
2.5
9%
4
.29
%
5
2.5
0%
1
.70
%
3.0
1%
5
0
.41
Ha
mm
er
3.2
6%
1
.05
%
4.5
9%
1
0
1.7
9%
0
.88
%
2.4
2%
1
0
0.5
0
Min
de
mo
ya
(199
7)
3.4
1%
1
.98
%
5.3
3%
2
5
1.9
9%
0
.27
%
4.6
1%
2
5
0.3
6
Stu
mp
y
3.4
7%
2
.25
%
5.6
6%
1
0
2.0
7%
0
.47
%
5.3
9%
9
0
.39
Sis
se
ne
y(2
001)
3.4
8%
2
.07
%
6.1
9%
1
4
3.6
9%
1
.86
%
5.5
3%
1
4
0.3
6
Fu
sh
imi
3.5
1%
2
.11
%
6.5
2%
1
7
2.9
6%
0
.65
%
4.9
9%
1
7
0.3
2
NE
-5
6
Ta
ble
NE
-11
a.
(co
ntin
ue
d)
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Ro
un
d(1
99
7)
3.5
1%
2
.70
%
4.6
3%
1
4
6.7
7%
4
.06
%
11
.60
%
14
0
.46
Na
ga
ga
mis
is
3.5
6%
2
.46
%
4.9
6%
1
2
2.5
2%
0
.79
%
4.0
3%
1
2
0.3
1
Wo
wu
n
3.5
6%
1
.79
%
5.0
8%
1
2
3.3
2%
1
.45
%
5.1
0%
1
2
0.4
0
Po
pe
ye
3
.58
%
0.2
1%
5
.08
%
9
2.8
1%
0
.46
%
5.9
3%
9
0
.65
Am
iko
ug
am
i 3
.65
%
1.7
1%
1
0.1
5%
1
0
2.9
7%
1
.05
%
8.1
3%
1
0
No
da
ta
Nip
issin
g(1
998)
3.6
9%
0
.90
%
5.9
7%
7
5
2.7
8%
0
.20
%
4.7
0%
7
5
0.4
2
Op
ee
pe
esw
ay
3.8
5%
2
.62
%
5.1
8%
1
0
2.5
9%
1
.73
%
3.1
6%
1
0
0.4
5
La
rde
r 3
.89
%
3.2
1%
4
.59
%
4
4.5
5%
1
.30
%
6.4
7%
4
N
o d
ata
Ju
mp
ing
Ca
rib
oo
3
.92
%
3.0
5%
4
.61
%
6
0.5
7%
0
%
0.8
4%
6
0
.51
Wa
ka
mi
(199
8)
3.9
3%
1
.38
%
6.4
7%
4
2
3.6
8%
1
.74
%
6.0
1%
4
2
0.4
0
We
ne
be
go
n
4.1
0%
0
.30
%
7.8
0%
4
6
1.6
2%
0
.28
%
3.4
7%
4
4
0.2
2
Fla
nd
ers
4
.22
%
2.9
8%
5
.96
%
20
1
.66
%
0.2
8%
2
.96
%
20
0
.38
Ho
we
llls
4.4
1%
3
.30
%
6.9
0%
8
0
.42
%
0%
1
.23
%
8
0.3
5
La
c la
Ca
ve
4
.49
%
0.3
6%
1
0.1
7%
3
3
3.6
9%
1
.36
%
7.3
4%
3
3
0.4
1
We
st
Ka
be
nu
ng
4
.56
%
2.8
3%
8
.19
%
22
1
.51
%
0.4
8%
3
.81
%
21
0
.40
Bu
rnfie
ld
4.6
1%
3
.54
%
5.7
2%
3
0
.62
%
0.2
0%
1
.17
%
3
No
da
ta
Re
d C
ed
ar
4.7
2%
2
.39
%
7.2
0%
1
5
1.2
9%
0
%
3.6
3%
1
5
0.2
5
Nig
ht
Ha
wk
4.7
9%
3
.19
%
7.3
9%
1
9
1.2
4%
0
.16
%
2.5
8%
1
9
0.2
8
Re
nn
ie
4.8
6%
2
.66
%
7.0
0%
6
4
.40
%
3.5
5%
5
.06
%
6
0.5
4
Th
istle
4
.86
%
3.5
9%
6
.05
%
5
1.8
2%
0
.67
%
3.4
3%
5
0
.34
Be
arh
ea
d
5.0
7%
3
.58
%
6.0
9%
6
2
.10
%
1.7
7%
2
.64
%
6
0.5
8
Bir
ch
5
.13
%
2.3
1%
7
.96
%
19
1
.22
%
0.1
3%
3
.78
%
19
0
.36
Du
nca
n
5.3
1%
4
.67
%
5.8
9%
5
2
.25
%
1.5
1%
2
.94
%
5
0.3
5
Mis
tin
iko
n
5.7
0%
4
.04
%
7.0
0%
5
2
.16
%
0.7
4%
3
.15
%
5
No
da
ta
Ve
rmili
on
5
.85
%
3.9
6%
6
.87
%
6
1.9
3%
1
.23
%
2.7
1%
6
0
.28
Do
g
6.0
2%
3
.83
%
8.8
8%
2
9
2.4
1%
1
.03
%
4.7
0%
2
9
0.3
1
NE
-5
7
Ta
ble
NE
-11
a.
(co
ntin
ue
d)
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Wh
ite
6
.02
%
3.2
6%
1
1.9
0%
4
2
.45
%
0.4
3%
4
.45
%
4
0.4
4
Fir
th
6.0
3%
5
.04
%
7.9
6%
5
2
.50
%
0.9
1%
3
.75
%
5
No
da
ta
Go
wg
an
da
6
.11
%
4.5
2%
7
.97
%
22
2
.49
%
0.9
5%
3
.75
%
22
0
.37
To
mik
o
6.1
3%
3
.53
%
8.0
4%
8
1
.66
%
0%
3
.06
%
6
0.3
9
Ke
no
ga
min
g
7.4
8%
6
.90
%
8.6
0%
4
2
.24
%
1.8
9%
3
.03
%
4
No
da
ta
Sis
se
ne
y
8.6
5%
8
.19
%
8.9
9%
3
1
.77
%
1.0
5%
2
.21
%
3
No
da
ta
No
rth
ea
st
ma
ture
fe
ma
le g
on
ad
os
om
ati
c,
vis
ce
ral
fat,
an
d r
ep
rod
uc
tiv
e i
nd
ice
s b
en
ch
ma
rks
.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at In
de
x
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Re
gio
n A
ve
rag
e1
.4
.09
%
2.6
7%
5
.83
%
14
2
.06
%
0.8
5%
3
.62
%
11
4
0.3
8
Re
gio
n A
ve
rag
e2
.3
.65
%
1.7
2%
6
.15
%
26
2
.43
%
0.7
8%
4
.91
%
25
0
.35
1.
All
lakes c
om
bin
ed (
n=
55).
2.
O
nly
la
ke
s w
ith
sam
ple
siz
e
10 (
n=
30).
NE
- 5
8
Ta
ble
NE
-11
b.
Ma
ture
ma
le w
alle
ye
fa
ll g
on
ad
oso
ma
tic in
de
x (
GS
I is
te
ste
s w
eig
ht
exp
resse
d a
s a
pe
rce
nt
of
the
to
tal w
eig
ht
min
us t
he
teste
s w
eig
ht)
, vis
ce
ral fa
t in
de
x (
VF
I is
fa
t w
eig
ht
exp
resse
d a
s a
pe
rce
nt
of
the
tota
l w
eig
ht)
, a
nd
re
pro
ductive
in
de
x (
R
is
3h
/L)
da
ta f
or
no
rth
ea
st
reg
ion
wa
terb
od
ies s
am
ple
d u
sin
g t
he
fa
ll w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
. O
nly
me
an
va
lue
s
w
ith
a
min
imu
m s
am
ple
siz
e
3 a
re in
clu
de
d.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Ro
un
d(2
00
1)
1.3
2%
0
.01
%
3.4
1%
2
9
1.3
2%
0
%
3.6
5%
3
2
0.3
7
Ke
no
ga
min
g
1.6
4%
1
.21
%
2.4
7%
1
4
1.9
9%
0
.81
%
4.5
7%
1
4
No
da
ta
Nig
ht
Ha
wk
1.7
9%
1
.24
%
2.4
2%
1
9
1.2
6%
0
%
2.8
2%
1
9
No
da
ta
Ritch
ie
1.8
2%
1
.39
%
3.2
9%
1
6
0.5
4%
0
%
1.2
4%
1
6
No
da
ta
Tim
iska
min
g
1.9
4%
0
.12
%
3.6
7%
6
5
1.1
0%
0
.18
%
6.0
5%
5
1
0.2
9
Gra
nite
hill
1
.95
%
0.0
1%
4
.08
%
62
1
.70
%
0%
4
.52
%
62
0
.33
Ob
ush
ko
ng
1
.97
%
0.0
3%
3
.19
%
27
1
.04
%
0%
2
.92
%
25
0
.51
Gra
ssy
2.0
7%
0
.26
%
3.6
4%
8
1
.57
%
0.7
5%
3
.39
%
8
No
da
ta
Oke
2
.07
%
0.9
8%
2
.97
%
23
0
.13
%
0%
0
.43
%
23
N
o d
ata
Wa
ka
mi
(200
1)
2.0
8%
0
.02
%
4.0
9%
3
0
3.3
3%
0
.75
%
5.9
2%
3
0
0.4
9
Mis
tin
iko
n
2.1
0%
1
.54
%
3.1
8%
2
7
1.7
0%
0
.46
%
2.8
0%
2
7
0.3
5
Kn
ife
2
.13
%
0.2
6%
3
.91
%
22
0
.25
%
0%
1
.02
%
24
N
o d
ata
To
mik
o
2.1
3%
0
.34
%
3.0
8%
1
8
0.6
7%
0
%
2.4
2%
1
8
0.5
2
Ju
mp
ing
Ca
rib
oo
2
.15
%
1.0
8%
3
.46
%
10
0
.43
%
0%
1
.07
%
10
0
.66
Be
arh
ea
d
2.1
8%
0
.34
%
3.6
4%
2
9
0.7
9%
0
%
2.2
1%
2
9
0.6
7
La
c la
Ca
ve
2
.24
%
0.0
1%
3
.59
%
55
2
.42
%
0%
6
.60
%
55
0
.51
Wa
wa
go
sh
e
2.2
6%
1
.69
%
3.2
2%
3
1
.02
%
0.3
3%
1
.89
%
3
No
da
ta
Th
istle
2
.27
%
1.5
1%
3
.43
%
19
0
.54
%
0%
1
.73
%
20
0
.39
Po
pe
ye
2
.27
%
0.1
2%
3
.79
%
18
2
.57
%
0.6
7%
5
.97
%
18
0
.72
Nu
ge
nt
2.2
9%
0
.75
%
3.2
7%
1
0
1.0
2%
0
.65
%
1.6
3%
1
0
No
da
ta
Ha
mm
er
2.3
3%
0
.64
%
4.0
3%
1
8
1.1
4%
0
.58
%
2.0
8%
1
8
0.6
3
McD
iarm
id
2.3
3%
0
.01
%
3.5
0%
4
6
0.5
5%
0
%
2.2
9%
4
6
0.3
4
Du
nca
n
2.3
9%
1
.86
%
3.1
3%
8
2
.36
%
1.0
9%
3
.73
%
8
No
da
ta
NE
- 5
9
Ta
ble
NE
-11
b.
(co
ntin
ue
d)
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Mo
un
tain
2
.39
%
1.4
5%
3
.49
%
16
0
.96
%
0.0
4%
2
.06
%
16
N
o d
ata
Stu
mp
y
2.4
7%
0
.01
%
21
.89
%
84
0
.84
%
0%
5
.55
%
82
0
.48
Ve
rmili
on
2
.48
%
0.8
1%
4
.68
%
15
1
.44
%
0%
4
.29
%
15
0
.41
We
ne
be
go
n
2.5
1%
1
.16
%
5.0
4%
9
7
1.3
9%
0
%
4.1
5%
8
7
0.3
1
Go
wg
an
da
2
.54
%
1.5
5%
4
.06
%
33
2
.42
%
0.7
7%
4
.14
%
33
0
.44
Ka
bin
aka
gam
i 2
.56
%
1.3
3%
3
.70
%
9
1.2
8%
0
.09
%
3.3
6%
9
0
.36
Wo
wu
n
2.5
7%
1
.26
%
3.6
7%
5
9
1.9
0%
0
%
5.4
5%
5
9
0.5
4
Ho
we
lls
2.5
7%
0
.01
%
4.4
3%
5
2
0.3
0%
0
%
1.0
9%
5
2
0.4
3
Ra
mse
y
2.5
9%
1
.30
%
3.6
2%
1
2
1.2
5%
0
.57
%
2.2
9%
1
2
0.4
7
Min
de
mo
ya
(199
7)
2.6
0%
0
.73
%
4.5
1%
1
24
1
.48
%
0%
4
.41
%
12
3
0.4
2
Ku
sh
og
2
.63
%
0.0
9%
4
.16
%
28
1
.22
%
0.0
8%
2
.58
%
28
0
.35
Am
iko
ug
am
i 2
.64
%
1.1
9%
3
.47
%
13
1
.49
%
0%
3
.18
%
16
N
o d
ata
Piv
ab
iska
2
.68
%
0.2
4%
4
.43
%
56
1
.02
%
0.0
3%
2
.81
%
56
0
.46
Lo
ng
po
int
2.7
0%
0
.22
%
4.3
3%
2
5
1.1
4%
0
%
3.3
2%
2
5
0.3
8
Fir
th
2.7
3%
2
.12
%
3.5
5%
1
0
2.2
4%
0
.94
%
2.9
5%
1
0
No
da
ta
Fu
sh
imi
2.7
4%
0
.23
%
5.1
6%
1
70
1
.69
%
0.2
6%
6
.19
%
17
0
0.4
2
Ge
ne
va
2
.75
%
1.6
3%
4
.07
%
4
0.5
0%
0
%
1.6
4%
4
N
o d
ata
Wh
ite
2
.76
%
0.2
5%
5
.98
%
37
1
.08
%
0%
3
.40
%
37
0
.45
Ga
rnh
am
2
.77
%
0.0
1%
1
9.4
9%
4
6
1.7
2%
0
%
4.7
0%
4
6
0.3
8
Fla
nd
ers
2
.78
%
1.8
8%
3
.94
%
22
1
.32
%
0%
2
.14
%
22
0
.45
We
st
Ka
be
nu
ng
2
.79
%
0.0
7%
6
.31
%
11
2
0.9
8%
0
%
6.7
0%
1
06
0
.50
Wh
ite
wa
ter
2.7
9%
0
.87
%
4.1
9%
1
2
1.0
5%
0
.03
%
2.9
0%
2
8
0.4
0
Nip
issin
g(1
998)
2.7
9%
0
.04
%
9.6
0%
2
93
2
.02
%
0.0
2%
8
.38
%
29
5
0.5
0
Po
st
2.8
4%
1
.29
%
3.6
6%
1
9
0.3
3%
0
%
1.0
7%
1
9
0.4
6
Sis
se
ne
y(2
001)
2.8
7%
1
.71
%
6.7
4%
1
6
1.6
8%
0
%
6.0
5%
1
8
0.4
3
Min
de
mo
ya
(200
1)
2.8
9%
0
.39
%
5.1
0%
7
9
1.6
5%
0
.03
%
4.6
4%
7
8
0.5
2
NE
- 6
0
Ta
ble
NE
-11
b.
(co
ntin
ue
d)
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Re
d C
ed
ar
2.8
9%
1
.37
%
4.8
4%
1
4
0.5
2%
0
%
1.7
4%
1
4
0.3
4
Ro
cky I
sla
nd
2
.93
%
2.0
1%
3
.45
%
4
0.7
7%
0
.13
%
1.6
9%
4
N
o d
ata
Op
ee
pe
esw
ay
2.9
4%
0
.94
%
5.8
5%
3
9
1.4
4%
0
%
2.5
7%
3
8
0.4
4
Ma
cu
tag
on
2
.95
%
1.2
9%
1
0.1
5%
1
6
1.4
5%
0
.41
%
4.3
3%
1
6
0.3
4
Do
g
3.0
1%
1
.23
%
7.4
6%
4
4
1.2
6%
0
%
2.6
1%
4
4
0.4
3
Na
ga
ga
mis
is
3.0
5%
0
.78
%
5.5
9%
2
4
1.3
2%
0
%
3.1
2%
2
4
0.4
1
Ro
un
d(1
99
7)
3.0
5%
1
.05
%
4.6
8%
4
2
5.2
7%
1
.72
%
10
.12
%
42
0
.50
Wh
itso
n
3.1
6%
2
.74
%
3.4
2%
3
0
.54
%
0.0
9%
0
.94
%
3
No
da
ta
Re
nn
ie
3.2
0%
1
.78
%
5.5
5%
2
2
3.1
3%
1
.49
%
5.3
7%
2
2
0.5
4
Sa
ga
na
sh
3
.23
%
0.7
3%
5
.44
%
83
1
.23
%
0%
3
.49
%
83
0
.28
Wa
ka
mi
(199
8)
3.3
4%
0
.25
%
9.1
2%
1
50
2
.69
%
0.5
2%
7
.77
%
15
0
0.4
9
Bir
ch
3
.36
%
0.6
5%
8
.83
%
21
1
.85
%
0.6
6%
3
.13
%
21
0
.39
Ag
ne
w(1
999
) 3
.50
%
1.9
8%
7
.00
%
4
0.1
4%
0
%
0.5
4%
4
N
o d
ata
Bu
rnfie
ld
4.2
6%
3
.18
%
63
51
%
7
1.3
4%
0
.51
%
1.7
8%
7
N
o d
ata
La
rde
r 4
.86
%
4.2
3%
5
.60
%
6
4.2
5%
2
.88
%
5.8
2%
6
N
o d
ata
NE
- 6
1
No
rth
ea
st
ma
ture
ma
le g
on
ad
os
om
ati
c,
vis
ce
ral
fat,
an
d r
ep
rod
uc
tiv
e i
nd
ice
s b
en
ch
ma
rks
.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at In
de
x
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Re
gio
n A
ve
rag
e1
.2
.61
%
1.0
3%
4
.95
%
37
1
.41
%
0.3
1%
3
.39
%
37
0
.45
Re
gio
n A
ve
rag
e2
.2
.52
%
0.7
2%
5
.21
%
45
1
.43
%
0.2
2%
3
.72
%
45
0
.45
1.
All
lakes c
om
bin
ed (
n=
64).
2.
O
nly
la
ke
s w
ith
sam
ple
siz
e
10 (
n=
54).
NE
- 6
2
Ta
ble
NE
-12
. W
alle
ye
life
his
tory
pa
ram
ete
rs f
or
On
tari
o w
ate
rbo
die
s s
am
ple
d u
sin
g th
e w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
199
3 t
o 2
00
1.
Pe
rce
ntile
sL
ife
His
tory
Pa
ram
ete
r (S
am
ple
Siz
e)
Me
an
(±9
5%
C.I
.)1
.S
.D.
(C.V
.)2
.M
inim
um
to
Maxim
um
M
ed
ian
5%
2
5%
7
5%
9
5%
Fe
ma
le a
du
lt m
ort
alit
y (
A%
) (n
=2
96
)2
8(2
7-3
0)
13
.6(4
8%
)7
to
89
2
5
12
1
9
34
5
7
Ma
le a
du
lt m
ort
alit
y (
A%
) (n
=2
95
)3
0(2
9-3
2)
12
.8(4
2%
)6
to
82
2
8
14
2
1
38
5
1
Fe
ma
le r
ela
tive
co
nd
itio
n
(n=
24
1)
1.0
3(1
.02
-1.0
4)
0.0
79
(8%
)0
.90
to
1.4
9
1.0
2
0.9
4
0.9
8
1.0
7
1.1
6
Ma
le r
ela
tive
co
nd
itio
n
(n=
26
5)
1.0
3(1
.02
-1.0
4)
0.0
90
(9%
)0
.87
to
1.4
8
1.0
2
0.9
2
0.9
7
1.0
7
1.1
8
Fe
ma
le a
sym
pto
tic le
ngth
(L
, m
m)
(n=
23
8)
74
4(7
33
-75
5)
84
.2(1
1%
)5
29
to
95
7
74
8
60
6
68
4
80
4
88
0
Ma
le a
sym
pto
tic le
ng
th (
L,
mm
) (n
=2
52
)6
18
(61
0-6
26
)6
4.7
(10
%)
48
5 t
o 8
14
6
16
5
08
5
72
6
54
7
38
Fe
ma
le B
rod
y c
oe
ffic
ien
t (K
, yr-1
)(n
=2
38
)0
.15
8(0
.15
4-0
.16
3)
0.0
37
7(2
4%
)0
.09
0 t
o 0
.32
7
0.1
52
0.1
10
0.1
32
0.1
80
0.2
30
Ma
le B
rod
y c
oe
ffic
ien
t (K
, yr-1
)(n
=2
52
)0
.18
0(0
.17
6-0
.18
5)
0.0
35
0(1
9%
)0
.10
5 t
o 0
.32
00
.17
60
.12
90
.15
5
0.2
00
0.2
44
Fe
ma
le O
me
ga
(,
mm
•yr-1
)(n
=2
38
)1
16
.40
(11
3.4
1-1
19
.38
)2
3.3
22
(20
%)
73
.49
to
20
0.2
7
11
4.4
5
80
.84
9
9.3
1
13
2.8
2
15
5.9
2
Ma
le O
me
ga
(,
mm
•yr-1
)(n
=2
52
)1
10
.57
(10
8.1
2-1
13
.02
)1
9.7
48
(18
%)
69
.45
to
18
1.1
9
10
9.4
1
80
.67
9
7.2
0
12
2.0
7
14
8.0
6
Pre
-ma
tura
tio
n g
row
th (
h,
mm
•yr-1
)(n
=2
78
)9
3.0
0(9
0.4
8-9
5.5
1)
21
.30
6(2
3%
)5
4.3
1 t
o 1
85.9
0
92
.07
6
1.1
3
76
.59
1
05
.95
1
29
.45
Fe
ma
le a
ge
-at-
50
% m
atu
rity
(ye
ars
) (n
=2
61
)4
.5(4
.3-4
.7)
1.5
1(3
4%
)2
.0 t
o 1
0.2
4
.2
2.5
3
.4
5.4
7
.1
Ma
le a
ge
-at-
50
% m
atu
rity
(ye
ars
) (n
=2
66
)3
.0(2
.9-3
.2)
1.1
0(3
7%
)0
.8 t
o 7
.7
2.9
1
.5
2.2
3
.7
4.9
NE
- 6
3
Ta
ble
NE
-12
. (c
on
tin
ued
)
Pe
rce
ntile
sL
ife
His
tory
Pa
ram
ete
r (S
am
ple
Siz
e)
Me
an
(±9
5%
C.I
.)1
.S
.D.
(C.V
.)2
.M
inim
um
to
Maxim
um
M
ed
ian
5%
2
5%
7
5%
9
5%
Fe
ma
le s
ize
-at-
50
% m
atu
rity
(m
m)
(n=
26
1)
44
1(4
36
-44
7)
42
.1(1
0%
)3
38
to
58
6
44
0
36
9
41
6
47
3
50
4
Ma
le s
ize
-at-
50
% m
atu
rity
(m
m)
(n=
26
6)
34
9(3
45
-35
3)
32
.9(9
%)
24
6 t
o 4
76
3
46
2
99
3
31
3
70
4
01
Fe
ma
le g
on
ad
oso
ma
tic in
de
x (
%)
(n=
15
1)
4.4
(4.2
-4.7
)1
.64
(37
%)
1.5
to
11
.1
4.2
2
.1
3.3
5
.4
7.5
Ma
le g
on
ad
oso
ma
tic ind
ex (
%)
(n=
15
5)
2.9
(2.8
-3.0
)0
.76
(26
%)
1.3
to
5.4
2
.8
1.8
2
.5
3.2
4
.5
Re
lative
fe
cu
nd
ity (
eg
gs•g
-1)
(n=
13
6)
49
.79
(47
.41
-52
.17
)1
4.0
49
(28
%)
18
.04
to
87
.98
5
0.4
7
26
.22
40
.01
5
8.6
9
72
.65
Fe
ma
le v
isce
ral fa
t in
de
x (
%)
(n=
15
1)
1.9
(1.7
-2.1
)1
.29
(68
%)
0.1
to
6.8
1
.6
0.3
0
.8
2.7
4
.4
Ma
le v
isce
ral fa
t in
de
x (
%)
(n=
15
5)
1.4
(1.2
-1.5
)0
.98
(72
%)
0.1
to
5.3
1
.1
0.2
0
.6
1.8
3
.2
Re
lative
ab
un
da
nce
(n
um
be
r•n
et-1
)(n
=4
41
)7
.5(6
.9-8
.2)
7.1
3(9
5%
)0
to
36
.1
5.3
0
.4
2.0
1
0.9
2
2.5
1.
95%
Confidence inte
rval of
the m
ean.
2.
Sta
ndard
devia
tion o
f th
e m
ean (
S.D
.) a
nd c
oeffic
ient
of
variation o
f th
e m
ean e
xpre
ssed a
s a
perc
ent
(C.V
.).
Fig
ure
4. M
ap o
f south
-centr
al O
nta
rio s
how
ing the location o
f th
e 9
9
wate
rbodie
s for
whic
h fall
walle
ye index n
ettin
g d
ata
were
availa
ble
.
SC
- 1
Ta
ble
SC
-1.
Th
e s
ou
th-c
en
tra
l re
gio
n w
ate
rbo
die
s s
am
ple
d u
sin
g t
he
fa
ll w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
19
93
to
20
01
.
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Allu
me
tte
1
99
9
45
53
00
-77
13
00
1
35
98
.3
No
da
ta
No
da
ta
3.2
1
82
1
Ba
lsa
m
19
98
& 2
00
14
43
50
1-7
85
03
8
46
65
.0
5.0
1
4.9
2
.5
18
67
Ba
rk
20
01
4
52
65
2-7
75
01
0
38
01
.0
24
.3
87
.5
4.6
1
64
5
Be
ave
r 2
00
1
44
30
30
-77
01
33
5
28
.8
2.4
1
2.0
2
.3
19
79
Be
lmo
nt
19
99
4
42
95
4-7
74
92
3
76
8.3
6
.2
16
.1
3.5
1
91
9
Be
nn
ett
2
00
1
44
54
56
-76
27
54
4
81
.4
4.4
1
2.2
4
.3
19
78
Big
Cle
ar
20
00
4
44
33
3-7
65
42
6
32
3.2
7
.7
19
.5
4.5
1
91
2
Big
Gu
ll 1
99
8
44
49
33
-76
57
24
2
36
3.4
3
.9
25
.9
5.0
1
84
2
Bla
ck
20
01
4
44
90
4-7
93
90
8
97
.6
5.2
1
0.7
2
.3
19
43
Bo
bs
19
97
4
44
10
4-7
63
52
0
24
49
.2
7.5
2
5.6
7
.0
20
00
Bra
nd
y
20
01
4
50
62
9-7
93
13
2
10
4.8
3
.5
7.5
1
.1
17
48
Bu
ck
20
01
4
44
21
8-7
65
71
3
70
.8
5.3
1
8.3
5
.3
19
13
Bu
ckh
orn
2
00
0
44
29
20
-78
23
30
3
19
1.0
2
.1
9.4
2
.3
18
85
Bu
ll 2
00
1
44
41
14
-76
58
34
1
13
.3
4.7
1
6.5
5
.3
19
13
Bu
rro
ws
20
01
4
45
02
1-7
93
94
0
86
.2
3.6
1
1.6
3
.6
19
44
Ca
me
ron
2
00
1
44
33
08
-78
45
41
1
30
3.2
6
.3
18
.3
4.2
1
85
9
Ca
no
nto
2
00
1
45
03
22
-76
47
40
2
21
.0
3.8
2
1.4
6
.3
18
36
Ce
ce
be
2
00
1
45
38
30
-79
34
26
7
70
.0
4.9
1
9.5
2
.6
16
39
Ce
nte
nn
ial
19
98
4
50
90
2-7
70
30
1
34
39
.8
13
.9
44
.2
3.8
1
79
7
Ch
em
on
g
20
00
4
42
31
9-7
82
33
5
22
80
.0
2.4
6
.7
2.6
1
92
0
Ch
esle
y
19
98
4
43
31
2-8
11
32
9
19
7.6
4
.3
16
.7
4.4
1
81
1
Ch
ristie
2
00
1
44
48
16
-76
26
20
6
45
.9
8.5
1
8.3
3
.5
20
07
Co
nse
co
n
20
01
4
40
00
0-7
72
70
0
60
0.0
6
.9
16
.7
0.2
2
11
7
Co
nsta
nt
20
01
4
52
42
0-7
65
85
8
68
.0
5.1
2
1.0
6
.6
18
22
Co
uch
ain
2
00
1
45
41
13
-77
44
04
4
5.3
7
.5
21
.4
4.1
1
75
3
Cra
ne
2
00
1
45
12
36
-79
56
34
5
12
.9
9.2
3
2.0
5
.5
18
26
Cro
tch
2
00
1
44
54
45
-76
48
35
1
67
7.4
8
.4
32
.2
5.5
1
87
6
Cro
we
1
99
9
44
28
49
-77
44
04
8
76
.4
5.6
1
5.9
2
.4
19
27
SC
- 2
Ta
ble
SC
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Da
lho
usie
2
00
1
44
57
56
-76
34
11
5
90
.0
4.5
1
1.0
3
.4
19
36
Da
lrym
ple
1
99
9
44
38
50
-79
06
32
1
33
2.5
2
.4
10
.4
2.6
1
88
8
De
er
20
01
4
53
44
5-7
91
62
9
11
9.0
3
.9
8.5
3
.5
16
37
De
nb
igh
2
00
1
45
07
48
-77
16
18
8
0.3
3
.5
7.0
3
.8
17
23
Do
llars
2
00
1
45
56
54
-80
12
39
9
69
.7
5.2
2
8.1
2
.9
17
33
Ele
ph
an
t 2
00
1
45
07
50
-78
07
56
1
09
5.8
1
.7
7.6
3
.1
16
56
Fe
rgu
so
ns
20
01
4
51
80
7-7
65
04
0
10
4.0
1
0.6
3
2.0
3
.5
18
60
Fo
ur
Mile
2
00
1
44
40
54
-78
44
15
7
86
.2
8.5
1
9.2
4
.3
18
30
Fra
se
r 2
00
1
45
11
23
-77
38
30
2
11
.4
6.6
1
5.5
4
.1
16
70
Fre
nch
Riv
er
19
93
,19
94
,1
99
5,
19
97
, 2
00
0 &
20
01
4
55
60
0-8
05
40
0
80
57
.5
No
da
ta
No
da
ta
No
da
ta1
73
5
Ge
org
es
20
01
4
53
62
1-7
45
82
5
46
.8
3.0
1
0.5
0
.2
19
29
Gib
so
n
20
01
4
45
75
4-7
94
45
9
26
3.5
7
.6
14
.3
2.4
1
91
5
Gir
ou
x
20
01
4
54
25
0-8
03
02
4
11
5.5
2
.9
6.1
1
.8
17
63
Go
Ho
me
2
00
1
45
00
26
-79
50
42
6
66
.0
8.6
3
2.6
5
.3
18
97
Go
lde
n
19
97
4
53
35
6-7
71
92
9
35
52
.2
8.5
2
4.4
4
.1
17
94
Go
od
erh
am
2
00
1
44
54
39
-78
22
59
8
7.4
6
.8
18
.9
5.0
1
73
2
Go
uld
1
99
8
44
35
38
-81
12
09
1
08
.1
1.8
8
.6
2.0
1
81
5
Go
va
n
20
01
4
50
75
9-7
64
74
9
12
5.5
4
.3
16
.0
7.0
1
82
2
Ho
lde
n
19
98
4
61
50
0-7
81
50
0
75
91
.9
No
da
ta
No
da
ta
No
da
ta1
66
2
Ho
rse
sh
oe
2
00
1
44
40
29
-76
57
35
6
8.8
3
.7
18
.3
5.3
1
91
5
Hu
ron
3.
19
96
, 1
99
7
& 2
00
1
44
30
00
-82
15
00
4
16
59
00
.0
61
.0
21
7.6
N
o d
ata
No
da
ta
Isa
be
lla
20
01
4
52
42
0-7
94
84
2
19
0.5
5
.4
15
.4
3.1
1
73
9
Ja
ck
20
01
4
44
13
8-7
80
12
9
12
21
.3
8.4
4
2.7
4
.9
18
20
Ja
cks
(BR
AC
EB
RID
GE
) 2
00
1
45
39
26
-79
20
13
8
8.3
1
.1
4.0
0
.7
16
33
Ja
cks
(NO
RT
H B
AY
) 1
99
9
45
56
00
-79
49
00
2
15
.4
3.6
1
2.7
3
.4
16
95
SC
- 3
Ta
ble
SC
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Ka
hsh
e
20
01
4
45
14
9-7
91
55
9
75
0.6
4
.9
18
.3
2.9
1
82
9
Ka
ka
kiw
ag
an
da
1
99
8
46
12
16
-80
47
33
1
87
.9
9.6
2
2.9
3
.3
17
33
Ka
sh
wa
ka
ma
k
20
00
4
45
13
2-7
70
22
2
12
19
.3
5.2
2
4.0
7
.2
18
22
Ka
ssh
ab
og
2
00
1
44
37
53
-77
57
54
8
90
.7
4.3
2
5.3
4
.1
18
49
Ke
nn
eb
ec
20
01
4
44
42
4-7
65
75
6
54
6.9
7
.9
27
.0
2.3
1
89
0
La
Clo
ch
e
19
96
& 2
00
14
60
84
0-8
20
43
4
11
44
.3
9.3
3
9.6
4
.5
16
61
La
ng
2
00
0
46
09
54
-81
40
02
5
95
.9
6.3
2
1.0
6
.3
16
63
Le
on
ard
2
00
1
45
04
28
-79
26
48
1
89
.0
6.8
1
8.3
7
.3
17
78
Lo
ng
(KIN
GS
TO
N)
20
00
4
44
02
3-7
64
61
7
30
1.1
6
.2
13
.4
1.8
1
94
5
Lo
ng
(SU
DB
UR
Y)
19
96
4
62
20
0-8
10
50
0
13
31
.0
10
.4
36
.5
4.5
1
68
6
Lo
ve
rin
g
19
98
4
61
35
5-8
04
51
2
90
.4
8.6
2
0.0
2
.5
17
33
Lo
we
r P
au
da
sh
2
00
1
44
58
03
-78
00
05
4
61
.3
6.6
2
1.3
6
.1
17
07
Ma
da
wa
ska
2
00
1
45
24
24
-76
24
21
1
00
1.1
9
.2
30
.5
1.5
1
97
6
McF
arl
an
e
19
96
4
62
50
1-8
05
72
3
14
0.6
7
.3
20
.1
2.4
1
69
8
McK
ella
r 2
00
1
45
30
49
-79
56
01
8
5.5
4
.5
9.8
3
.5
17
27
Min
k
20
01
4
53
35
3-7
70
25
5
54
2.0
5
.9
13
.0
2.5
1
83
9
Mis
sis
sa
ga
go
n
20
01
4
45
21
8-7
70
44
5
52
4.8
8
.8
23
.5
6.7
1
81
1
Mis
sis
sip
pi
20
01
4
50
40
1-7
60
95
7
23
51
.4
2.7
9
.2
3.8
2
01
7
Mo
ira
2
00
1
44
29
08
-77
27
07
8
22
.7
3.9
1
1.0
3
.0
19
32
Mo
un
tain
1
99
8 &
20
01
44
42
31
-80
41
52
7
5.7
1
.8
3.2
2
.9
18
48
Ne
pe
wa
ssi
20
01
4
62
20
0-8
03
80
0
11
91
.4
No
da
ta
16
.7
2.8
1
71
5
No
rca
n
20
01
4
51
00
0-7
65
20
0
72
8.5
2
.6
4.6
4
.6
18
46
No
sb
on
sin
g
20
01
4
61
20
9-7
91
23
2
17
04
.5
5.5
1
4.0
2
.9
17
15
Oa
k
20
01
4
43
54
8-7
75
45
2
28
5.2
7
.1
24
.7
5.2
1
85
7
Pa
ke
sh
ka
g
20
01
4
55
65
7-8
03
14
1
53
.7
3.4
2
2.9
1
.2
17
46
Pic
ke
rel R
ive
r 1
99
5,
19
96
&
19
97
4
55
50
0-8
04
60
0
50
5.0
5
.5
18
.3
2.7
1
74
5
SC
- 4
Ta
ble
SC
-1.
(co
ntin
ue
d)
La
ke
Ye
ar
Sa
mp
led
La
titu
de
-
Lo
ng
itu
de
1.
Su
rfa
ce
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
2.
Pig
eo
n
20
00
4
42
81
5-7
82
94
2
53
49
.0
3.0
1
7.4
2
.2
18
85
Re
dh
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er
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7
1.
Latitu
de a
nd longitude in d
egre
es,
min
ute
s,
and s
econds.
2.
GD
D>
5 is g
row
ing-d
egre
e-d
ays a
bove 5
o C
els
ius.
3.
Data
not
inclu
ded in t
he lake c
hara
cte
ristics s
um
mary
sta
tistics b
ecause o
nly
a s
mall
port
ion o
f th
e e
ntire
wate
rbody w
as s
am
ple
d.
SC
- 5
Ch
ara
cte
ris
tic
s o
f s
ou
th-c
en
tra
l re
gio
n l
ak
es
sa
mp
led
us
ing
fa
ll w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
.
S
urf
ace
Are
a
(he
cta
res)
Me
an
De
pth
(m
)M
ax.
Depth
(m
)S
ecch
i(m
)G
DD
>5
Me
an
1
20
9.2
6
.0
19
.6
3.6
1
82
2
Min
imu
m
45
.3
0.9
2
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0.2
1
62
4
Ma
xim
um
1
35
98
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24
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7.3
2
11
7
Me
dia
n
51
2.9
5
.5
18
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3.5
1
82
2
5%
Pe
rce
ntile
6
8.0
1
.8
6.1
1
.1
16
39
25
% Q
ua
rtile
1
37
.6
3.9
1
1.8
2
.4
17
32
75
% Q
ua
rtile
1
21
9.3
7
.7
25
.0
4.6
1
91
3
95
% P
erc
en
tile
5
34
9.0
1
0.7
3
9.6
6
.7
20
07
Sa
mp
le S
ize
9
9
94
9
6
96
9
9
SC - 6
Table SC-2. Walleye relative abundance and average size for south-central region waterbodies sampled using the fall walleye index netting standard 1993 to 2001.
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
Allumette 5.9 0.26 24 22 335 473
Balsam (19983.
) 1.2 0.11 36 27 326 430
Balsam (2001) 1.7 0.15 28 22 351 531
Bark 1.9 0.19 24 16 494 1409
Beaver 3.2 0.19 14 13 358 455
Belmont 0.4 0.11 18 7 298 323
Bennett 1.8 0.17 12 11 395 619
Big Clear 2.5 0.39 9 7 345 490
Big Gull 3.6 0.18 28 25 389 635
Black 0 - 12 0 - -
Bobs 1.6 0.16 25 19 418 865
Brandy 4.1 0.17 12 12 322 419
Buck 2.2 0.70 6 3 392 626
Buckhorn 0.8 0.12 28 16 408 657
Bull 1.8 0.32 12 7 424 830
Burrows 0 - 6 0 - -
Cameron 0.7 0.11 24 13 296 253
Canonto 1.4 0.22 12 9 394 648
Cecebe 3.8 0.29 12 11 364 568
Centennial 0.7 0.10 34 19 363 478
Chemong 0.9 0.12 28 17 396 582
Chesley 10.0 0.24 6 6 378 554
Christie 1.8 0.22 12 10 421 797
Consecon 2.1 0.37 12 8 380 566
Constant 1.6 0.20 12 10 387 592
Couchain 0 - 12 0 - -
Crane 0.4 0.14 12 5 625 2789
Crotch 2.0 0.14 29 24 373 546
Crowe 1.3 0.15 16 12 359 526
Dalhousie 1.1 0.21 12 8 392 567
Dalrymple 1.8 0.14 24 20 328 450
Deer 6.7 0.20 14 14 334 453
Denbigh 0.4 0.16 12 4 491 1193
Dollars 4.8 0.36 12 10 335 431
Elephant 0.1 0.05 24 3 587 2194
Fergusons 0.3 0.12 12 4 521 1269
Four Mile 3.0 0.27 14 12 414 712
Fraser 2.2 0.27 12 9 342 440
French River (19933.
) 2.3 0.20 24 18 393 762
French River (19943.
) 4.3 0.13 58 52 377 788
French River (19953.
) 2.5 0.12 68 50 381 712
French River (19973.
) 3.5 0.11 63 57 354 549
French River (20003.
) 5.7 0.10 69 67 402 802
SC - 7
Table SC-2. (continued)
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
French River (2001) 2.0 0.15 35 26 411 830
Georges 0.9 0.20 12 7 441 1060
Gibson 0.7 0.19 12 6 392 819
Giroux 1.9 0.19 11 10 283 391
Go Home 3.7 0.38 11 9 431 924
Golden 1.0 0.17 23 13 567 2651
Gooderham 1.0 0.19 12 8 441 1013
Gould 9.2 0.40 6 6 303 398
Govan 5.0 0.25 12 12 379 558
Holden 4.4 0.17 36 33 345 482
Horseshoe 1.6 0.39 6 4 369 569
Huron (19963.
) 2.0 0.19 26 20 337 707
Huron (19973.
) 1.6 0.17 35 23 306 425
Huron (2001) 1.2 0.19 26 15 417 1458
Isabella 0.5 0.16 12 5 574 2072
Jack 0.7 0.13 24 12 454 1054
Jacks (BRACEBRIDGE) 2.2 0.22 12 10 341 429
Jacks (NORTH BAY) 2.8 0.28 12 11 395 738
Kahshe 0.3 0.14 12 3 526 2231
Kakakiwaganda 6.7 0.54 6 5 544 2196
Kashwakamak 7.1 0.25 21 21 371 531
Kasshabog 0 - 13 0 - -
Kennebec 1.2 0.22 14 9 454 1067
La Cloche (19963.
) 0.5 0.13 24 9 557 2454
La Cloche (2001) 1.4 0.18 24 17 531 2202
Lang 6.5 0.26 16 15 455 1218
Leonard 0.2 0.09 12 3 607 2367
Long (KINGSTON) 1.5 0.19 10 8 405 613
Long (SUDBURY) 6.3 0.19 12 12 434 1075
Lovering 0.4 0.27 6 2 433 988
Lower Paudash 0.1 0.06 12 1 634 2500
Madawaska 4.3 0.14 12 12 265 192
McFarlane 2.2 0.20 16 14 414 1047
McKellar 1.6 0.22 12 9 339 533
Mink 7.8 0.31 12 11 395 724
Mississagagon 0.8 0.15 14 8 463 991
Mississippi 1.1 0.14 23 16 337 406
Moira 5.9 0.18 12 12 359 589
Mountain (19983.
) 18.0 0.18 6 6 343 406
Mountain (2001) 6.5 0.14 4 4 375 615
Nepewassi 1.7 0.20 18 13 338 515
Norcan 0.5 0.20 12 4 445 980
Nosbonsing 10.6 0.24 12 12 387 719
Oak 5.3 0.15 12 12 428 778
SC - 8
Table SC-2. (continued)
Average Size2.
LakeAverageCatch1.
StandardError of
the Mean
Sample Size (number of net-nights)
Number of Nets that Caught
WalleyeTotal
LengthWeight
Pakeshkag 0.1 0.06 12 1 443 1069
Pickerel River (19953.
) 3.8 0.19 16 16 398 925
Pickerel River (19963.
) 7.7 0.34 11 10 387 854
Pickerel River (1997) 3.2 0.38 11 9 365 654
Pigeon 0.6 0.090 42 22 402 741
Redhorse 0.6 0.15 12 6 415 758
Restoule 8.4 0.29 12 12 318 435
Rice 27.7 0.12 20 20 409 736
Richard 0.9 0.19 9 6 350 418
Rock Island 1.0 0.22 12 7 451 1167
Salmon Trout 3.1 0.32 12 9 367 450
Sandy 6.8 0.17 12 12 292 277
Second Depot 0.5 0.16 12 5 401 686
Sharbot (20003.) 0.5 0.12 21 9 563 2646
Sharbot (2001) 0.3 0.10 28 7 535 2252
Skootamatta 2.2 0.16 30 23 513 1716
Stewartville (20003.
) 0.7 0.13 12 8 352 405
Stewartville (2001) 1.0 0.18 12 8 352 598
Stoco 1.8 0.21 12 10 318 437
Stormy 4.0 0.51 12 8 337 516
Sturgeon 0.5 0.12 28 11 286 288
Sturgeon (Chalk) 1.7 0.24 12 9 366 465
Three Mile 1.0 0.15 12 9 473 1152
Trout 2.0 0.25 16 12 532 1868
Wilber 1.0 0.23 12 7 465 1249
Wolfe 8.1 0.27 14 13 341 412
Woodcock 5.9 0.27 12 11 320 360
1. Average catch is the geometric mean number of walleye caught net-1
.2. Average size is the arithmetic mean of total length in mm and wet weight in grams. 3. Data not used to calculate relative abundance or average size benchmarks.
SC - 9
South-central region walleye relative abundance benchmarks.
Average Size AverageCatch Total
LengthWeight
Mean 2.8 406 877
Minimum 0 265 192
Maximum 27.7 634 2789
Median 1.7 393 641
5% Percentile 0.1 296 323
25% Quartile 0.7 348 480
75% Quartile 3.8 442 1057
95% Percentile 8.3 574 2252
Sample Size 100 96
SC - 10
Table SC-3a. Annual female walleye mortality (%) for south-central region waterbodies sampled using the fall walleye index netting standard. “Observed mortality” is a direct estimate based on age structure of female fish and was calculated from age 5 onwards using Robson and Chapman’s estimator. “Estimated mortality” is an indirect estimate based on age structure of the population regardless of sex using the empirical formula: Female mortality = 0.952(Combined sex mortality) (n=246, R2=0.77, standard error of the estimate=6.493).
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Huron (1996) No data 10.5% Restoule 21% 25.7%
Stewartville (2001) No data 12.4% Three Mile No data 25.7%
Sharbot 2001 No data 12.4% Mountain (1998) 17% 25.7%
Gibson No data 13.3% Canonto No data 25.7%
Huron 2001 14% 13.3% Stoco 20% 25.7%
Skootamatta 16% 14.3% French River (2000) 29% 26.7%
Pickerel River (1995) 17% 15.2% Mountain (2001) No data 26.7%
Mississagagon No data 16.2% Sandy 14% 26.7%
Jacks (BRACEBRIDGE) No data 17.1% Big Gull 32% 26.7%
Sturgeon 11% 17.1% Nosbonsing 29% 27.6%
Christie No data 17.1% Kennebec No data 27.6%
Georges No data 17.1% Pigeon 29% 27.6%
La Cloche (2001) 14% 18.1% Long (SUDBURY) 25% 28.6%
Nepewassi 21% 18.1% Govan 14% 28.6%
Lang 19% 19.0% Dalrymple 50% 29.5%
Sharbot (2000) No data 19.0% French River (1993) 29% 29.5%
Kakakiwaganda 25.0% 20.0% Deer 15% 29.5%
Stormy 21% 20.9% Dalhousie No data 29.5%
French River (1994) 22% 20.9% French River (1995) 35% 30.5%
Trout 21% 20.9% Bull No data 30.5%
Rock Island No data 20.9% French River (2001) 34% 31.4%
Gould No data 20.9% Holden 27% 31.4%
Wolfe No data 20.9% Buck No data 31.4%
Crotch 24% 20.9% Bobs No data 31.4%
Huron (2001) No data 20.9% Bennett No data 31.4%
Jack 14% 21.9% Mississippi No data 31.4%
Big Clear 23% 21.9% French River (1997) 35% 30.5%
Moira 25% 21.9% Pickerel River (1997) 29% 32.4%
Golden 21% 22.8% Allumette 35% 32.4%
Balsam (2001) 32% 22.8% Crowe No data 33.3%
McFarlane 23% 23.8% Chemong 32% 33.3%
Pickerel River (1996) 24% 23.8% Beaver 67% 35.2%
Brandy No data 23.8% Kashwakamak 40% 36.2%
Cecebe 32% 23.8% Horseshoe No data 36.2%
Giroux No data 23.8% Salmon Trout 33% 37.1%
McKellar No data 23.8% Jacks (NORTH BAY) 43% 37.1%
Wilber No data 23.8% Gooderham No data 38.1%
Consecon 29% 23.8% La Cloche (1996) No data 39.0%
Four Mile 27% 24.8% Rice 40% 40.0%
Dollars 18% 24.8% Constant 50% 40.0%
Go Home 18% 24.8% Mink 42% 40.9%
Centennial No data 24.8% Long (KINGSTON) No data 40.9%
SC - 11
Table SC-3a. (continued)
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Balsam (1998) 36% 24.8% Buckhorn 53% 42.8%
Oak 42% 45.7% Woodcock 44% 57.1%
Madawaska No data 47.6% Fraser No data 59.0%
Sturgeon (Chalk) No data 47.6% Bark 63% 75.2%
South-central region female walleye mortality benchmarks.
Observed Mortality
EstimatedMortality
Mean 29% 27.8%
Minimum 11% 10.5%
Maximum 67% 75.2%
Median 27% 25.7%
5% Percentile 14% 13.3%
25% Quartile 20% 20.9%
75% Quartile 35% 31.9%
95% Percentile 53% 47.6%
Sample Size 56 92
SC - 12
Table SC-3b. Annual male walleye mortality (%) for south-central region waterbodies sampled using the fall walleye index netting standard. “Observed mortality” is a direct estimate based on age structure of male fish and was calculated from age 5 onwards using Robson and Chapman’s estimator. “Estimated mortality” is an indirect estimate based on age structure of the population regardless of sex using the empirical formula: Male mortality = 0.945(Combined sex mortality) + 2.844 (n=242, R2=0.65, standard error of the estimate=7.477).
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Huron (1996) No data 13.2% Canonto No data 28.4%
Stewartville (2001) No data 15.1% Mountain (1998) 29% 28.4%
Sharbot (2001) No data 15.1% Restoule 40% 28.4%
Gibson No data 16.1% Stoco No data 28.4%
Huron (1997) No data 16.1% Three Mile No data 28.4%
Skootamatta 14% 17.0% Big Gull 26% 29.3%
Pickerel River (1995) 12% 18.0% French River (2000) 25% 29.3%
Mississagagon No data 18.9% Mountain (2001) No data 29.3%
Christie No data 19.9% Sandy 30% 29.3%
Georges No data 19.9% Kennebec 33% 30.3%
Jacks (BRACEBRIDGE) 33% 19.9% Nosbonsing 29% 30.3%
Sturgeon No data 19.9% Pigeon 26% 30.3%
La Cloche (2001) 20% 20.8% Govan 32% 31.2%
Nepewassi 14% 20.8% Long (SUDBURY) 33% 31.2%
Lang 21% 21.8% Dalhousie No data 32.2%
Sharbot (2000) No data 21.8% Dalrymple No data 32.2%
Kakakiwaganda 17% 22.7% Deer 48% 32.2%
Crotch 21% 23.7% French River (1993) 29% 32.2%
French River (1994) 21% 23.7% Bull 44% 33.1%
Gould 13% 23.7% French River (1995) 25% 33.1%
Huron (2001) No data 23.7% Bennett No data 34.1%
Rock Island No data 23.7% Bobs 40% 34.1%
Stormy 25% 23.7% Buck 25% 34.1%
Trout 21% 23.7% French River (2001) 31% 34.1%
Wolfe 22% 23.7% Holden 41% 34.1%
Big Clear No data 24.6% Mississippi No data 34.1%
Jack 40% 24.6% Allumette 32% 35.0%
Moira 20% 24.6% French River (1997) 32% 35.0%
Balsam (2001) 20% 25.5% Pickerel River (1997) 44% 35.0%
Golden 26% 25.5% Chemong 37% 35.9%
Brandy No data 26.5% Crowe No data 35.9%
Cecebe No data 26.5% Rice 33% 35.9%
Consecon 23% 26.5% Beaver 32% 37.8%
Giroux No data 26.5% Horseshoe No data 38.8%
McFarlane No data 26.5% Kashwakamak 37% 38.8%
McKellar No data 26.5% Jacks (NORTH BAY) 30% 39.7%
Pickerel River (1996) 25% 26.5% Salmon Trout 38% 39.7%
Wilber No data 26.5% Gooderham No data 40.7%
Balsam (1998) 21% 27.4% La Cloche (1996) 39% 41.6%
Centennial 22% 27.4% Constant No data 42.6%
Dollars 29% 27.4% Long (KINGSTON) 43% 43.5%
Four Mile 25% 27.4% Mink 42% 43.5%
SC - 13
Table SC-3b. (continued)
Lake Observed Mortality
EstimatedMortality
Lake Observed Mortality
EstimatedMortality
Go Home 38% 27.4% Buckhorn 38% 45.4%
Oak 48% 48.2% Woodcock 74% 59.6%
Madawaska 40% 50.1% Fraser No data 61.5%
Sturgeon (Chalk) 40% 50.1% Bark No data 77.6%
South-central region male walleye mortality benchmarks.
Observed Mortality
EstimatedMortality
Mean 31% 30.5%
Minimum 12% 13.2%
Maximum 74% 77.6%
Median 30% 28.4%
5% Percentile 14% 16.1%
25% Quartile 22% 23.6%
75% Quartile 38% 34.5%
95% Percentile 48% 50.1%
Sample Size 60 92
SC - 14
Table SC-4a. Predicted female walleye weight-at-total length for south-central region waterbodies sampled using the fall walleye index netting standard. A geometric mean predictive regression model was fitted to logarithmically transformed (base 10) individual
observations (only lakes with 30 fish sampled). Weight in grams and total length in millimetres.
Weight-length relation1.
Calculated weight at standard total lengths
Lake
x10-6 350 400 450 500 550 Condition2.
Crotch 3.01 8.10 368 550 785 1077 1435 0.96
Wolfe 3.08 5.38 369 556 799 1106 1483 0.98
French River (1993) 3.09 5.59 406 613 883 1222 1641 1.04
Deer 3.12 4.10 355 539 778 1080 1455 0.95
Mountain (1998) 3.12 4.19 363 550 795 1104 1486 0.95
Chesley 3.13 4.21 387 587 849 1180 1591 0.99
Balsam (2001) 3.14 3.71 361 549 795 1107 1493 0.94
Sandy 3.14 4.07 396 603 872 1214 1638 1.03
Pickerel River (1997) 3.17 3.78 439 670 973 1359 1838 1.13
Govan 3.18 2.97 366 559 813 1136 1539 0.94
Kashwakamak 3.18 2.93 361 551 802 1121 1518 0.96
Big Gull 3.19 3.08 402 615 896 1254 1699 1.05
Moira 3.20 2.77 383 588 857 1200 1628 1.02
Skootamatta 3.21 2.97 436 669 976 1369 1859 1.13
Mink 3.23 2.35 388 597 873 1227 1669 1.01
Pickerel River (1996) 3.23 2.56 422 650 951 1336 1818 1.07
Rice 3.24 2.20 385 593 869 1222 1664 0.99
Allumette 3.26 2.02 397 614 901 1271 1734 1.06
Go Home 3.27 1.72 359 555 816 1151 1572 0.96
Huron (2001) 3.27 2.01 419 649 953 1345 1837 1.12
Pickerel River (1995) 3.27 1.85 386 597 877 1238 1691 1.03
French River (1994) 3.28 1.83 405 627 923 1303 1782 1.07
Holden 3.28 1.72 387 600 882 1246 1704 0.99
Huron (1997) 3.28 1.87 413 641 943 1332 1821 1.06
Lang 3.30 1.67 415 645 951 1347 1845 1.09
Stormy 3.30 1.58 393 610 900 1274 1745 0.99
French River (1997) 3.31 1.44 380 590 872 1236 1694 0.98
French River (2000) 3.33 1.23 364 569 842 1195 1642 1.00
French River (2001) 3.33 1.31 388 606 896 1273 1749 1.00
French River (1995) 3.35 1.16 386 604 897 1276 1757 1.02
Restoule 3.37 1.07 401 629 935 13333 1838 1.06
Long (SUDBURY) 3.38 0.95 377 592 882 1260 1738 1.01
Nosbonsing 3.39 0.87 366 576 859 1228 1696 0.96
1. Two-tailed Student t-tests were employed to test significance of slope and intercept ( o: or = 0). 2. Condition calculated as the average of the individual ratios of observed weight to predicted weight from south-central
region median regression model (see below).
SC - 15
South-central region female walleye weight-at-total length benchmarks.
Weight-length relation Calculated weight at standard total lengths
x10-6 350 400 450 500 550
Mean 3.23 2.71 389 598 876 1231 1676
Minimum 3.01 0.87 355 539 778 1077 1435
Maximum 3.39 8.10 439 670 976 1369 1859
Median 3.24 2.20 387 597 877 1236 1694
5% Percentile 3.08 0.95 359 549 785 1080 1455
25% Quartile 3.17 1.67 368 569 842 1180 1591
75% Quartile 3.30 3.71 402 615 901 1276 1757
95% Percentile 3.38 5.59 436 669 973 1359 1845
Sample Size 33
SC - 16
Table SC-4b. Predicted male walleye weight-at-total length for south-central region waterbodies sampled using the fall walleye index netting standard. A geometric mean predictive regression model was fitted to logarithmically transformed (base 10) individual
observations (only lakes with 30 fish sampled). Weight in grams and total length in millimetres.
Weight-length relation1.
Calculated weight at standard total lengths
Lake
x10-6 350 400 450 500 550 Condition2.
Huron (1996) 2.95 13.19 422 626 886 1208 1601 1.08
Crotch 2.98 9.74 371 553 785 1075 1428 0.95
Kakakiwaganda 3.03 10.36 530 794 1134 1560 2083 1.29
Madawaska 3.03 6.99 357 535 765 1053 1405 1.00
Jacks (NORTH BAY) 3.04 8.13 441 661 946 1303 1741 1.10
Mountain (1998) 3.08 5.19 356 536 771 1067 1430 0.94
Dalrymple 3.09 5.13 373 563 810 1122 1506 0.98
Sandy 3.09 4.99 362 548 788 1091 1465 0.98
Big Gull 3.10 5.16 397 601 866 1201 1614 1.05
Consecon 3.11 4.59 375 568 819 1137 1529 0.94
Four Mile 3.11 4.33 354 536 773 1072 1442 0.92
Govan 3.11 4.38 358 542 782 1085 1459 0.94
Stormy 3.13 4.13 379 576 833 1158 1561 0.98
Balsam (2001) 3.15 3.43 354 539 781 1089 1470 0.93
Mink 3.15 3.84 396 604 875 1219 1646 1.02
Bobs 3.16 3.49 382 583 845 1179 1594 0.99
Deer 3.16 3.36 368 561 814 1135 1534 0.93
Salmon Trout 3.16 3.15 345 526 763 1064 1438 0.90
Go Home 3.19 2.85 372 569 829 1160 1573 0.96
Rice 3.19 3.08 402 615 896 1254 1699 1.03
Kashwakamak 3.20 2.70 374 573 835 1170 1587 0.98
Oak 3.20 2.57 356 545 795 1113 1510 0.92
Dollars 3.21 2.81 412 633 924 1295 1759 1.04
Moira 3.21 2.81 412 633 924 1295 1759 1.05
Wolfe 3.21 2.49 365 561 818 1148 1559 0.95
Cecebe 3.22 2.38 370 569 832 1168 1587 0.95
French River (1997) 3.22 2.46 383 588 860 1207 1640 1.00
Restoule 3.22 2.59 403 619 905 1271 1727 1.05
Skootamatta 3.23 2.56 422 650 951 1336 1818 1.12
Beaver 3.25 1.92 356 550 806 1135 1547 0.93
Trout 3.26 1.93 380 587 861 1214 1656 1.02
French River (1994) 3.27 1.93 402 623 915 1292 1764 1.04
Holden 3.28 1.68 371 576 847 1197 1636 0.98
Pickerel River (1996) 3.28 1.81 400 620 912 1289 1762 1.05
Huron 1997 3.29 1.80 422 655 965 1364 1867 1.08
Pickerel River (1995) 3.29 1.66 389 604 890 1258 1721 1.04
Bark 3.30 1.68 418 649 957 1355 1856 1.09
Allumette 3.32 1.54 430 670 991 1406 1930 1.07
French River (2000) 3.32 1.40 391 609 901 1279 1754 1.01
French River (1995) 3.33 1.28 379 592 876 1244 1709 1.01
SC - 17
Table SC-4b. (continued)
Weight-length relation1.
Calculated weight at standard total lengths
Lake
x10-6 350 400 450 500 550 Condition2.
French River (2001) 3.35 1.14 380 594 881 1254 1726 1.00
Nosbonsing 3.35 1.08 360 563 835 1188 1635 0.96
Long (SUDBURY) 3.37 0.99 371 582 865 1234 1701 0.99
Lang 3.38 1.04 413 649 966 1379 1903 1.12
La Cloche (2001) 3.39 1.17 493 775 1155 1651 2280 1.39
Woodcock 3.57 0.30 363 584 889 1296 1821 0.92
1. Two-tailed Student t-tests were employed to test significance of slope and intercept ( o: or = 0). 2. Condition calculated as the average of the individual ratios of observed weight to predicted weight from south-central
region median regression model (see below).
South-central region male walleye weight-at-total length benchmarks.
Weight-length relation Calculated weight at standard total lengths
x10-6 350 400 450 500 550
Mean 3.21 3.41 389 598 872 1223 1662
Minimum 2.95 0.30 345 526 763 1053 1405
Maximum 3.57 13.19 530 794 1155 1651 2280
Median 3.21 2.64 380 586 863 1208 1638
5% Percentile 3.03 1.04 354 536 771 1067 1430
25% Quartile 3.11 1.68 365 561 814 1135 1534
75% Quartile 3.29 4.33 403 623 912 1292 1759
95% Percentile 3.38 9.74 441 670 991 1406 1930
Sample Size 46
SC - 18
Table SC-5a. Average female walleye total length-at-age (millimetres) data for south-central region waterbodies sampled using the fall walleye index netting method. Only
mean values with a minimum sample size 4 are included.
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
La Cloche (1996) 644
Trout 661 692
Kakakiwaganda 566 651
Sharbot (2000) 680
Buckhorn 469
Balsam (1998) 420
McFarlane 340
Cameron 340
Woodcock 349 417
Salmon Trout 372
Jack 373
Cecebe 386 477 529
Sturgeon (Chalk) 387
Canonto 403
Mississagagon 413
Bennett 433
La Cloche (2001) 528 590
Nepewassi 299
Dollars 300
Long (SUDBURY) 301 363 412 569 682
Balsam (2001) 305 399
Madawaska 309 353
Big Clear 310 368 401
Crowe 320
Four Mile 320 388
Consecon 321 368
Mississippi 329
French River (1993) 330 413 503
McKellar 331
Beaver 331
Wolfe 334 404 421
Horseshoe 335
Fraser 339
Chesley 344 406 462
Constant 348
Stoco 354
Georges 368
Go Home 373 445 486 590
Bark 374 489 591
Oak 391 451 494
Long (KINGSTON) 392
Gould 429 464
Pigeon 217 316 378 448
SC - 19
Table SC-5a. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
French River (1997) 234 286 346 416 459 477 513
Moira 241 336 399 464 443 567
Nosbonsing 244 333 424 478 528 543 638
French River (1995) 244 354 342 448 523 552 590 606
Holden 246 307 379 436 467
Restoule 253 308 349
Huron (1997) 256 332 404
Allumette 258 310 387 421 450 483 538
Crotch 260 312 365
Deer 260 337 413
Big Gull 261 339 376 427
Sandy 269 389
French River (1994) 284 313 404 484 547 587 623 679
Stormy 284 347 439 542
Pickerel River (1997) 286 435 532
Mountain 1998 286 330 393
Kashwakamak 286 357 403 458
French River (2001) 286 372 414 471 495 559 631
Govan 286 378 465
Pickerel River (1995) 289 350 493
French River (2000) 296 358 403 462 490 534 574 620
Mink 303 408 453 607
Huron 2001 306 701
Rice 309 362 435 483 509 551 572 592
Pickerel River (1996) 313 377 471
Jacks (NORTH BAY) 315 409
Dalrymple 327 453
Sharbot (2001) 346
Lang 365 482 542 661
Skootamatta 373 458 493 587 618
South-central region female walleye total length-at-age benchmarks.
Average total length (mm) at age
1 2 3 4 5 6 7 8 9 10
Mean 283 348 405 459 492 550 599 604 658 646
Minimum 217 286 316 378 417 477 513 538 620 567
Maximum 373 482 542 590 591 644 680 682 692 701
Median 286 337 403 460 490 542 590 612 661 658
5% Percentile 234 300 340 401 417 477 513 538 620 567
25% Quartile 256 320 370 421 450 529 572 568 620 602
75% Quartile 306 372 434 483 523 587 651 627 692 690
95% Percentile 365 429 493 566 591 644 680 682 692 701
Sample Size 31 51 48 26 15 13 9 8 3 4
SC - 20
Table SC-5b. Average male walleye total length-at-age (millimetres) data for south-central region waterbodies sampled using the fall walleye index netting method. Only
mean values with a minimum sample size 4 are included.
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Denbigh 504
Jack 437
Sharbot (2000) 619
Pigeon 431 483
Madawaska 321 337
Buckhorn 355
Chemong 360
Crotch 364 398 497
Salmon Trout 378 397 442
Centennial 380
Rock Island 390
Richard 401
La Cloche (1996) 404 590
Bennett 412
Gould 438
Trout 444
Golden 463
Nepewassi 292 337
Dollars 294 343 383 429
Jacks (BRACEBRIDGE) 304 363
French River (1993) 306 394
Four Mile 316 370 439 461 520
Balsam (2001) 316 372 407
Consecon 318 366 383 451
Wolfe 327 389 405
Fraser 330 449
Sturgeon (Chalk) 330 369
Beaver 332 387 399 414
Deer 335 377 424 425 462
Mississippi 336
Moira 339
McFarlane 344 376 424
Brandy 344 416
Stoco 347
Chesley 351 399
Bobs 356 465
Buck 360 416
Huron (1996) 365 435 511
Long (SUDBURY) 368 372 438 522 570
Go Home 375 389 456 474
Govan 375 406
Bark 385 455 529
Long (KINGSTON) 388
SC - 21
Table SC-5b. (continued)
Average total length (mm) at age Lake
1 2 3 4 5 6 7 8 9 10
Oak 389 441 465 494 505
Christie 396
Kennebec 416 458
La Cloche (2001) 433 505 526
Kakakiwaganda 441 480 511
Nosbonsing 226 327 378 400 451 475 512
Balsam (1998) 231 356
Cecebe 231 368 431
French River (1997) 232 271 333 396 423 439
Woodcock 234 318 358 363
Holden 237 295 356 399 412 481
French River (1995) 242 358 378 404 485
Restoule 245 304 340 379
Allumette 249 301 336 379 444
Huron (1997) 250 356 425 471
Big Clear 256 303 338
Sandy 259 376
Kashwakamak 264 358 393 425 462 474 496 494
French River (1994) 267 324 371 434 511
Mountain (1998) 269 349 374 401 439
Stormy 273 330 405
Big Gull 274 325 375 404 439 467 492
Pickerel River (1995) 282 370 474
French River (2000) 287 353 383 435 419 473 484 565
Dalrymple 291 437
Pickerel River (1997) 296 406
Huron (2001) 300 499
Rice 301 355 396 424 455 467 485 500
French River (2001) 302 364 402 431 454 494
Mink 305 390 428 500
Pickerel River (1996) 311 386 450 513
Bull 314 360 417 448
Jacks (NORTH BAY) 321 382 412
Skootamatta 347 448 489 554
Lang 364 448 499 585
SC - 22
South-central region male walleye total length-at-age benchmarks.
Average total length (mm) at age
1 2 3 4 5 6 7 8 9 10
Mean 275 352 394 429 444 475 479 528 517
Minimum 226 271 318 337 363 425 429 486 483
Maximum 364 448 505 526 529 590 619 570 585
Median 271 351 388 425 442 464 475 520 497
5% Percentile 231 294 336 358 363 425 429 486 483
25% Quartile 245 325 368 399 416 444 451 512 492
75% Quartile 301 375 416 456 474 500 494 554 565
95% Percentile 347 441 480 513 529 590 619 570 585
Sample Size 30 55 62 39 16 13 15 5 7 0
SC - 23
Table SC-6a. Average female walleye weight-at-age (grams) data for south-central region waterbodies sampled using the fall walleye index netting method. Only mean
values with a minimum sample size 4 are included.
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
La Cloche (1996) 3535
Trout 3310 3830
Kakakiwaganda 2397 3480
Sharbot (2000) 3912
Buckhorn 1001
Balsam (1998) 617
Cameron 314
Woodcock 357 723
McFarlane 380
Jack 414
Salmon Trout 431
Sturgeon (Chalk) 522
Cecebe 540 984 1671
Canonto 544
Mississagagon 589
Bennett 717
La Cloche (2001) 2004 2919
Nepewassi 229
Balsam (2001) 230 568
Dollars 245
Long (SUDBURY) 247 452 580 1921 3788
Crowe 249
Madawaska 250 365
Big Clear 267 438 607
Four Mile 277 507
Consecon 290 438
Mississippi 291
McKellar 293
Beaver 316
Fraser 318
Horseshoe 322
Wolfe 323 571 647
French River (1993) 327 672 1338
Constant 353
Chesley 371 616 927
Stoco 402
Georges 443
Go Home 465 863 1068 2117
Oak 488 785 1070
Long (KINGSTON) 541
Bark 544 1230 2230
Gould 713 921
Pigeon 82 268 479 881
SC - 24
Table SC-6a. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
French River (1997) 110 201 373 678 984 1199 1613
Nosbonsing 111 303 680 1009 1481 1717 2974
Moira 120 343 560 966 856 1929
Holden 125 260 489 823 1074
French River (1995) 125 414 356 962 1643 1793 2265 2363
Restoule 143 259 413
Huron (1997) 147 369 674
Deer 148 302 581
Crotch 149 258 430
Big Gull 156 358 503 732
Allumette 160 289 566 796 922 1232 1743
Sandy 177 538
Mountain (1998) 194 314 524
Govan 194 459 892
Stormy 200 359 809 1654
Kashwakamak 200 415 570 941
French River (2001) 208 459 666 1081 1216 1857 2769
French River (1994) 213 283 651 1236 1801 2083 2752 3476
Pickerel River (1995) 222 416 1272
French River (2000) 224 417 611 1012 1180 1635 2020 2618
Pickerel River (1997) 225 858 1789
Mink 242 644 922 2298
Rice 263 421 773 1109 1306 1752 1959 2152
Huron (2001) 273 4300
Pickerel River (1996) 312 503 1128
Dalrymple 319 931
Jacks (NORTH BAY) 335 680
Sharbot (2001) 443
Lang 477 1229 1881 3293
Skootamatta 538 1063 1398 2402 2802
South-central region female walleye weight-at-age benchmarks.
Average weight (g) at age
1 2 3 4 5 6 7 8 9 10
Mean 220 399 667 1025 1272 1886 2504 2511 3247 3170
Minimum 82 201 268 479 723 1199 1613 1717 2618 1929
Maximum 538 1229 2004 2919 2230 3535 3912 3788 3830 4300
Median 200 343 569 952 1180 1752 2117 2558 3293 3225
5% Percentile 110 230 356 580 723 1199 1613 1717 2618 1929
25% Quartile 147 283 438 732 922 1635 1959 1948 2618 2452
75% Quartile 242 459 745 1070 1643 2083 3310 2786 3830 3888
95% Percentile 477 713 1398 2397 2230 3535 3912 3788 3830 4300
Sample Size 31 51 48 26 15 13 9 8 3 4
SC - 25
Table SC-6b. Average male walleye weight-at-age (grams) data for south-central region waterbodies sampled using the fall walleye index netting method. Only mean
values with a minimum sample size 4 are included.
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Denbigh 1245
Jack 722
Sharbot (2000) 2887
Pigeon 757 1089
Madawaska 284 326
Chemong 383
Buckhorn 411
Crotch 427 583 1043
Centennial 455
Salmon Trout 458 532 738
Rock Island 523
Richard 574
Bennett 610
La Cloche (1996) 746 2795
Gould 792
Trout 853
Golden 1750
Nepewassi 216 338
Dollars 238 388 535 805
Jacks (BRACEBRIDGE) 249 430
French River (1993) 250 579
Four Mile 261 443 721 884 1237
Balsam (2001) 279 453 573
Wolfe 286 522 600
Beaver 291 493 534 643
Fraser 292 748
Consecon 295 425 474 837
Mississippi 304
Deer 318 454 678 671 897
Sturgeon (Chalk) 345 456
Moira 353
Chesley 353 593
Brandy 357 636
McFarlane 376 579 847
Stoco 410
Bobs 417 917
Buck 417 686
Long (SUDBURY) 460 464 769 1440 2038
Go Home 461 529 891 1009
Govan 464 566
Huron (1996) 472 786 1483
Oak 500 723 903 1073 1154
Long (KINGSTON) 521
SC - 26
Table SC-6b. (continued)
Average weight (g) at age Lake
1 2 3 4 5 6 7 8 9 10
Christie 548
Bark 565 1007 1601
Kennebec 724 1009
La Cloche (2001) 1051 1743 2016
Kakakiwaganda 1063 1510 1688
Woodcock 88 263 445 420
Nosbonsing 92 294 472 571 974 1070 1314
Balsam (1998) 99 362
Cecebe 105 445 707
French River (1997) 105 185 333 597 761 881
Holden 109 219 415 598 679 1095
French River (1995) 123 435 518 668 1191
Restoule 133 259 380 561
Huron (1997) 141 435 1005 1034
Big Clear 146 260 343
Allumette 147 263 364 534 931
Sandy 150 465
Kashwakamak 157 417 544 708 922 996 1152 1151
Mountain (1998) 164 378 450 576 684
Stormy 173 316 580
French River (1994) 175 327 502 831 1481
Big Gull 183 358 492 605 851 1000 1250
Pickerel River (1995) 202 486 1102
French River (2000) 205 410 533 826 717 1098 1185 1945
Dalrymple 222 769
Huron (2001) 239 1263
Pickerel River (1997) 245 683
Bull 252 425 686 729
Mink 252 560 731 1247
Rice 255 416 581 745 947 1035 1173 1261
French River (2001) 263 437 622 798 935 1256
Pickerel River (1996) 281 562 985 1340
Jacks (NORTH BAY) 342 545 711
Skootamatta 422 965 1271 1926
Lang 482 1010 1456 2265
SC - 27
South-central region male walleye weight-at-age benchmarks.
Average weight (g) at age
1 2 3 4 5 6 7 8 9 10
Mean 198 428 633 798 880 1143 1130 1536 1436
Minimum 88 185 263 326 420 671 684 1166 1043
Maximum 482 1063 1750 2016 1601 2795 2887 2038 2265
Median 174 410 526 721 806 996 1070 1314 1250
5% Percentile 92 219 338 445 420 671 684 1166 1043
25% Quartile 133 292 443 576 678 884 837 1237 1089
75% Quartile 252 472 711 891 1004 1154 1185 1926 1945
95% Percentile 422 1010 1456 1688 1601 2795 2887 2038 2265
Sample Size 30 55 62 39 16 13 15 5 7 0
SC - 28
Table SC-7a. Empirical female walleye von Bertalanffy growth equation for south-central region
waterbodies sampled using the fall walleye index netting standard. L defines the asymptotic or maximum total length, K is the Brody growth coefficient and defines growth rate towards the maximum, and to shifts the growth curve along the axis to allow for apparent non-zero
body length at age zero (to=-1). Omega ( ) is early growth and corresponds to the growth rate near to. h is pre-maturation growth (i.e., growth to 350 mm total length for combined sexes).
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Dalrymple 25 620 0.237 146.70 No data
Big Gull 52 630 0.199 125.04 103.70
Govan 39 652 0.232 151.24 129.10
Allumette 154 656 0.193 126.56 91.67
Kashwakamak 51 660 0.217 143.21 120.23
Crotch 31 662 0.176 116.52 92.31
Balsam (2001) 35 697 0.196 136.39 98.88
Rice 198 729 0.196 142.54 126.96
Big Clear 27 741 0.194 143.54 89.84
Pickerel River (1997) 39 743 0.170 129.09 No data
Moira 36 748 0.135 100.88 106.30
Mink 48 755 0.215 162.06 139.70
Holden 119 771 0.152 116.97 92.95
French River (2001) 121 775 0.172 133.35 125.44
French River (1995) 165 777 0.174 135.00 118.77
Restoule 48 791 0.134 106.01 No data
Nosbonsing 68 795 0.149 118.70 104.82
Stormy 66 801 0.143 114.60 No data
French River (1997) 141 807 0.139 111.72 83.75
French River (2000) 240 812 0.151 122.56 118.36
French River (1993) 47 817 0.160 130.94 104.87
French River (1994) 185 820 0.167 136.64 101.08
Bark 29 833 0.215 178.81 No data
Deer 43 842 0.153 129.07 106.13
Skootamatta 40 861 0.139 119.64 180.72
Lang 63 869 0.155 134.34 No data
Go Home 32 870 0.135 117.82 No data
Pickerel River (1996) 47 875 0.157 137.22 136.41
Long (SUDBURY) 41 880 0.168 147.76 97.62
Pickerel River (1995) 47 886 0.159 141.21 120.57
Huron (2001) 37 939 0.144 135.28 No data
SC - 29
South-central region female walleye von Bertalanffy growth equation benchmarks.
L (mm) K (mm year –1) h (mm year–1)1.
Mean 778 0.172 131.84 109.54
Minimum 620 0.134 100.88 70.95
Maximum 939 0.237 178.81 180.72
Median 791 0.167 133.35 106.21
5% Percentile 630 0.135 106.01 78.26
25% Quartile 729 0.149 118.70 96.91
75% Quartile 842 0.194 141.33 120.61
95% Percentile 886 0.232 162.06 140.74
Sample Size 31 48
1. Pre-maturation growth benchmarks are calculated from all south-central lakes.
SC - 30
Table SC-7b. Empirical male walleye von Bertalanffy growth equation for south-central region
waterbodies sampled using the fall walleye index netting standard. L defines the asymptotic or maximum total length, K is the Brody growth coefficient and defines growth rate towards the maximum, and to shifts the growth curve along the axis to allow for
apparent non-zero body length at age zero (to=-1). Omega ( ) is early growth and corresponds to the growth rate near to. h is pre-maturation growth (i.e., growth to 350 mm total length for combined sexes).
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Restoule 81 508 0.221 112.04 No data
Woodcock 42 536 0.194 103.71 77.04
Sandy 47 542 0.242 130.65 127.47
Mountain (1998) 64 549 0.187 102.93 106.52
Allumette 137 554 0.206 114.91 91.67
Stormy 52 568 0.200 113.68 No data
Holden 132 573 0.223 127.72 92.95
Deer 47 583 0.196 114.54 106.13
Kashwakamak 168 583 0.225 131.18 120.23
Salmon Trout 32 588 0.192 112.46 102.63
Big Gull 95 591 0.193 114.19 103.70
Rice 245 592 0.217 128.55 126.96
Wolfe 85 595 0.229 136.20 105.69
Dollars 58 600 0.197 117.98 82.89
Moira 37 600 0.155 92.70 106.30
Centennial 26 604 0.190 114.86 No data
Beaver 36 604 0.165 99.38 105.18
Dalrymple 31 620 0.173 107.12 No data
Nosbonsing 76 623 0.180 112.12 104.82
Bull 26 626 0.212 132.58 124.83
Consecon 35 627 0.172 107.36 96.20
Go Home 36 640 0.171 109.24 No data
Govan 41 640 0.197 125.85 129.10
French River (1997) 165 643 0.194 124.82 83.75
Mink 81 645 0.202 129.83 139.70
Balsam (1998) 28 650 0.154 100.14 No data
Oak 45 651 0.209 136.20 140.74
Jacks (NORTH BAY) 35 654 0.240 157.14 144.00
Crotch 40 654 0.152 99.37 92.31
Balsam (2001) 38 667 0.172 114.51 98.88
French River (1995) 131 679 0.177 120.15 118.77
French River (1993) 26 685 0.184 125.66 104.87
Four Mile 42 687 0.164 112.45 99.06
Bobs 32 689 0.184 127.10 115.19
French River (1994) 172 703 0.193 135.43 101.08
Pickerel River (1997) 26 703 0.208 146.01 No data
French River (2000) 246 709 0.150 106.70 118.36
French River (2001) 117 711 0.156 110.63 125.44
Lang 95 714 0.184 130.90 No data
Pickerel River (1996) 65 719 0.205 147.58 136.41
SC - 31
Table SC-7b. (continued)
Lake Sample Size L (mm) K (mm year–1) h (mm year–1)
Long (SUDBURY) 48 737 0.178 131.08 97.62
Skootamatta 57 738 0.153 112.78 180.72
Trout 34 771 0.161 123.75 No data
Huron (1996) 30 771 0.169 130.10 120.64
La Cloche (2001) 33 776 0.167 129.82 No data
Kakakiwaganda 32 814 0.156 127.20 No data
South-central region male walleye von Bertalanffy growth equation benchmarks.
L (mm) K (mm year –1) h (mm year–1)1.
Mean 646 0.188 120.25 109.54
Minimum 508 0.150 92.70 70.95
Maximum 814 0.242 157.14 180.72
Median 642 0.188 118.78 106.21
5% Percentile 542 0.153 99.38 78.26
25% Quartile 592 0.169 112.04 96.91
75% Quartile 703 0.202 130.10 120.61
95% Percentile 771 0.229 146.01 140.74
Sample Size 46 48
1. Pre-maturation growth benchmarks are calculated from all south-central lakes.
SC - 32
Table SC-8a. Female walleye age and size-at-maturity (total length in millimetres) schedules for south-central region waterbodies sampled using the fall walleye index netting standard. A logistic regression model was fitted to logarithmically transformed (base 10) maturity ogive (as percents).
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Lang 1.95 1.10 2.79 486.5 450.0 523.0
Skootamatta 1.95 0.83 3.06 475.3 440.1 510.5
Gould 2.14 1.86 2.43 418.7 357.1 480.3
Go Home 2.33 1.03 3.63 423.4 397.2 449.5
Huron (2001) 2.50 1.63 3.38 419.2 393.0 445.5
Govan 2.78 2.39 3.17 439.9 421.2 458.7
Big Clear 2.90 2.22 3.57 373.7 291.7 455.7
Dalrymple 2.90 2.59 3.22 409.7 388.3 431.1
Pickerel River (1996) 2.95 2.40 3.50 459.4 437.4 481.5
Mink 2.97 2.34 3.60 481.0 428.1 534.0
Bark 2.99 2.52 3.47 490.7 453.4 527.9
Deer 3.03 2.31 3.75 425.0 399.9 450.0
Wolfe 3.05 2.74 3.35 402.6 365.7 439.6
Oak 3.12 2.07 4.18 486.2 471.6 500.7
Kashwakamak 3.22 2.94 3.51 422.6 405.8 439.4
Pickerel River (1995) 3.22 1.90 4.54 479.6 457.9 501.3
Cecebe 3.25 2.53 3.97 401.3 375.3 427.2
French River (1993) 3.27 2.58 3.96 463.6 433.0 494.1
Restoule 3.32 2.66 3.97 367.5 332.8 402.1
Woodcock 3.34 2.24 4.45 368.2 354.6 381.8
McFarlane 3.38 2.83 3.94 440.8 421.2 460.3
Stormy 3.71 3.12 4.31 411.4 391.7 431.1
French River (1994) 3.72 3.04 4.40 468.2 424.8 511.5
French River (2001) 3.78 3.25 4.32 463.7 431.4 496.0
Moira 3.88 2.74 5.02 416.5 395.3 437.5
Rice 3.92 3.44 4.39 472.2 425.0 519.5
Chemong 3.93 3.23 4.64 405.6 388.3 423.0
French River (1995) 3.94 3.21 4.67 450.6 402.4 498.7
Nosbonsing 4.02 3.30 4.73 474.8 437.9 511.8
Crotch 4.05 2.67 5.43 416.0 345.3 486.6
Big Gull 4.06 3.62 4.50 433.1 418.1 448.1
Allumette 4.24 2.95 5.52 419.2 397.4 441.0
Chesley 4.25 2.97 5.53 478.1 416.0 540.3
French River (2000) 4.28 3.01 5.54 468.9 421.1 516.6
Holden 4.30 3.31 5.30 445.3 419.9 470.6
Nepewassi 4.41 3.94 4.89 440.3 423.8 456.8
Balsam 2001 4.50 3.13 5.87 450.0 423.2 476.9
Pickerel River (1997) 4.52 3.56 5.48 468.1 445.9 490.3
Huron (1997) 4.75 4.03 5.46 538.5 506.0 570.9
Pigeon 4.75 4.27 5.23 439.3 413.7 464.9
Long (SUDBURY) 4.80 4.38 5.23 556.6 531.9 581.3
SC - 33
Table SC-8a. (continued)
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Buckhorn 4.87 1.92 7.81 404.3 389.9 418.7
French River (1997) 5.43 3.89 6.98 479.7 434.4 525.0
South-central region female walleye maturity benchmarks.
Maturity by age (years) Maturity by size (mm)
50% 10% 90% 50% 10% 90%
Mean 3.60 2.76 4.43 445.0 413.0 477.0
Minimum 1.95 0.83 2.43 367.5 291.7 381.8
Maximum 5.43 4.38 7.81 556.6 531.9 581.3
Median 3.71 2.74 4.39 440.8 419.9 476.9
5% Percentile 2.14 1.10 3.06 373.7 345.3 418.7
25% Quartile 2.99 2.31 3.57 416.5 391.7 441.0
75% Quartile 4.25 3.25 5.23 474.8 434.4 511.5
95% Percentile 4.80 4.03 5.87 490.7 471.6 540.3
Sample Size 43
SC - 34
Table SC-8b. Male walleye age and size-at-maturity (total length in millimetres) schedules for south-central region waterbodies sampled using the fall walleye index netting standard. A logistic regression model was fitted to logarithmically transformed (base 10) maturity ogive (as percents).
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Gould 0.80 0.11 1.49 339.6 320.4 358.7
Big Clear 1.00 0.70 1.31 258.8 249.5 268.1
Dalrymple 1.30 0.69 1.91 316.4 299.3 333.4
Rice 1.36 0.50 2.22 321.1 295.7 346.6
Huron (2001) 1.36 0.76 1.95 336.3 312.8 359.8
Stoco 1.43 1.18 1.68 300.2 290.7 309.7
Moira 1.44 0.34 2.53 330.3 314.8 345.7
Govan 1.45 0.68 2.22 340.3 311.3 369.3
Pickerel River (1996) 1.45 0.56 2.34 331.2 247.3 414.9
Restoule 1.48 0.67 2.28 275.1 241.3 309.0
Mink 1.51 0.87 2.15 367.3 349.3 385.2
Pickerel River (1997) 1.56 1.16 1.96 366.9 341.1 392.8
Sandy 1.60 1.08 2.11 335.9 322.8 348.9
Wolfe 1.60 0.48 2.72 338.0 316.1 359.8
Stormy 1.62 1.04 2.19 318.9 289.7 348.1
Oak 1.63 0.77 2.48 366.7 348.1 385.4
Cecebe 1.78 0.43 3.14 325.8 310.7 341.0
French River (1995) 1.83 0.47 3.20 341.9 303.2 380.5
Kashwakamak 1.84 1.08 2.60 330.4 268.5 392.3
Jacks (BRACEBRIDGE) 1.85 0.93 2.77 292.9 249.6 336.2
French River (2000) 1.90 0.20 3.60 348.1 290.5 405.7
French River (2001) 1.96 0.49 3.42 352.3 300.5 404.0
Beaver 2.06 0.95 3.16 331.5 250.4 412.7
Mississippi 2.14 0.66 3.62 350.3 274.9 425.6
Balsam (2001) 2.19 1.46 2.92 343.3 324.1 362.5
Allumette 2.21 1.46 2.95 314.9 269.5 360.3
Go Home 2.23 1.75 2.70 348.8 299.6 398.1
Salmon Trout 2.25 1.50 3.01 336.4 300.5 372.3
Sturgeon (Chalk) 2.27 1.78 2.76 331.1 307.5 354.6
Big Gull 2.28 1.27 3.28 342.5 315.6 369.5
Nosbonsing 2.28 1.55 3.00 340.9 301.8 380.1
Bobs 2.29 1.22 3.35 369.8 338.7 400.9
Holden 2.31 1.67 2.94 306.4 267.6 345.2
Consecon 2.38 1.60 3.15 339.8 323.6 356.0
Bark 2.39 1.04 3.73 392.5 266.7 518.3
Crotch 2.41 1.41 3.42 333.2 293.4 373.0
French River (1994) 2.44 1.28 3.60 346.3 288.7 403.8
Pickerel River (1995) 2.44 1.22 3.65 374.8 354.4 395.2
Four Mile 2.47 1.21 3.73 358.8 320.0 397.6
Woodcock 2.61 1.82 3.40 288.0 264.2 311.8
Constant 2.63 1.38 3.87 424.6 401.1 448.2
SC - 35
Table SC-8b. (continued)
Maturity by age (years) Maturity by size (mm) Lake
50% 10% 90% 50% 10% 90%
Dollars 2.65 1.83 3.47 317.5 286.1 348.9
Huron (1997) 2.67 1.68 3.66 414.8 370.9 458.7
Chesley 2.79 2.60 2.98 370.7 359.4 382.1
Centennial 2.82 2.24 3.41 373.2 358.1 388.4
Balsam (1998) 2.89 1.36 4.41 353.1 316.0 390.3
Buckhorn 2.90 2.30 3.49 353.1 334.5 371.7
Fraser 2.94 2.31 3.57 352.7 337.0 368.3
Madawaska 2.94 2.60 3.27 302.1 290.4 313.9
French River (1997) 3.09 2.18 4.00 331.6 269.8 393.4
McFarlane 3.15 0.71 5.58 394.9 378.8 411.0
French River (1993) 3.48 2.09 4.88 378.4 226.8 530.0
Chemong 4.87 2.95 6.80 359.7 348.9 370.5
South-central region male walleye maturity benchmarks.
Maturity by age (years) Maturity by size (mm)
50% 10% 90% 50% 10% 90%
Mean 2.17 1.25 3.10 341.7 305.9 377.5
Minimum 0.80 0.11 1.31 258.8 226.8 268.1
Maximum 4.87 2.95 6.80 424.6 401.1 530.0
Median 2.23 1.21 3.14 340.3 303.2 372.3
5% Percentile 1.30 0.30 1.68 288.0 247.3 309.7
25% Quartile 1.60 0.70 2.48 330.3 286.1 348.9
75% Quartile 2.61 1.67 3.57 358.8 324.1 397.6
95% Percentile 3.15 2.60 4.88 394.9 370.9 458.7
Sample Size 53
SC
- 3
6
Ta
ble
SC
-9.
Wa
lleye
re
lative
fe
cu
nd
ity (
eg
gs p
er
gra
m o
f to
tal
we
igh
t) d
ata
fo
r so
uth
-ce
ntr
al
reg
ion
wa
terb
od
ies s
am
ple
d u
sin
g
the
fa
ll w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
. O
nly
me
an
va
lue
s w
ith
a m
inim
um
sa
mp
le s
ize
3
are
in
clu
de
d.
La
ke
M
ea
n
Sta
nd
ard
Err
or
Min
imu
m
Maxim
um
Media
n
5%
Pe
rce
ntile
25
%Q
ua
rtile
75
%Q
ua
rtile
95
%P
erc
en
tile
Sa
mp
leS
ize
Big
Cle
ar
20
.30
5
.89
3
.11
5
8.0
2
8.5
0
3.6
8
5.1
5
35
.21
5
0.5
5
11
Sto
co
2
3.0
7
1.5
4
20
.43
2
7.4
8
22
.19
2
0.6
4
21
.45
2
3.8
1
26
.75
4
Bra
nd
y
30
.35
3
.45
2
5.2
3
36
.92
2
8.8
9
25
.59
2
7.0
6
32
.91
3
6.1
2
3
McK
ella
r 3
0.4
6
2.4
0
27
.27
3
5.1
7
28
.94
2
7.4
4
28
.11
3
2.0
5
34
.54
3
Mo
ira
3
4.0
1
2.2
9
11
.38
4
9.9
3
35
.02
1
9.4
8
31
.05
3
9.4
1
48
.23
1
7
No
sb
on
sin
g
39
.86
1
.47
2
5.4
5
52
.44
4
0.2
9
27
.38
3
5.8
7
46
.47
4
9.0
6
27
La
ng
4
1.2
0
4.5
4
3.3
3
77
.92
4
7.3
4
8.8
4
18
.71
5
8.8
6
76
.62
2
6
McF
arl
an
e
42
.03
2
.90
2
7.0
0
60
.54
4
2.7
7
29
.31
3
4.8
6
46
.74
5
5.9
1
11
Ka
sh
wa
ka
ma
k
44
.29
6
.07
8
.51
8
5.2
0
46
.15
9
.85
2
9.8
0
58
.93
8
3.1
7
16
Fre
nch
Riv
er
(200
1)
45
.29
1
.20
2
3.5
8
66
.33
4
6.0
9
30
.35
4
0.1
0
49
.45
5
8.9
4
53
Re
sto
ule
4
7.5
1
3.4
2
32
.78
7
0.4
6
44
.19
3
3.6
3
37
.73
5
4.7
7
68
.30
1
3
Fre
nch
Riv
er
(199
5)
50
.37
1
.97
2
2.5
9
89
.32
4
9.0
0
29
.18
4
0.9
3
58
.60
7
2.7
6
50
Mo
un
tain
(20
01)
50
.59
7
.51
4
2.4
9
65
.59
4
3.6
7
42
.61
4
3.0
8
54
.63
6
3.4
0
3
Big
Gu
ll 5
1.2
6
1.8
5
41
.67
5
7.4
8
53
.90
4
2.1
9
46
.61
5
5.3
8
56
.80
1
0
Fre
nch
Riv
er
(199
7)
52
.89
2
.21
3
0.7
9
89
.58
5
1.8
2
34
.32
4
6.8
3
58
.01
7
3.5
0
30
Ho
lde
n
53
.72
2
.09
3
9.0
5
72
.26
5
1.9
5
42
.90
4
6.9
6
58
.74
6
8.7
9
21
Lo
ng
(SU
DB
UR
Y)
54
.00
5
.08
1
4.8
9
80
.65
5
5.0
9
26
.34
4
8.2
3
64
.75
7
4.4
8
12
Pic
ke
rel R
ive
r (1
99
6)
54
.12
2
.48
3
0.6
9
69
.58
5
6.8
3
38
.76
4
4.6
6
62
.52
6
6.2
9
19
Pic
ke
rel R
ive
r (1
99
5)
54
.73
3
.36
3
1.0
5
77
.12
5
4.3
2
31
.63
4
6.4
3
63
.18
7
6.0
2
16
La
Clo
ch
e (
1996)
55
.45
4
.89
3
4.3
6
74
.14
5
8.1
3
38
.02
4
8.5
8
62
.20
7
1.2
5
7
Go
uld
5
8.4
7
4.8
0
41
.68
7
9.9
6
62
.18
4
2.4
9
44
.76
6
6.0
0
75
.45
8
Ka
ka
kiw
ag
an
da
5
9.4
3
2.6
9
45
.88
8
8.5
8
56
.87
4
6.4
6
50
.27
6
3.2
0
84
.52
2
0
Fre
nch
Riv
er
(200
0)
59
.65
1
.68
2
8.0
4
14
1.6
9
58
.68
3
5.5
3
50
.41
6
5.9
5
89
.28
9
9
Fre
nch
Riv
er
(199
4)
59
.89
1
.81
3
3.1
5
10
2.8
3
58
.55
3
7.3
2
48
.36
7
1.5
1
81
.09
7
0
Pic
ke
rel R
ive
r (1
99
7)
60
.21
4
.42
4
0.6
2
81
.24
5
7.2
8
44
.52
5
2.1
9
64
.31
8
0.6
4
9
Allu
me
tte
6
2.1
1
1.5
4
40
.91
8
5.2
8
59
.98
4
6.9
7
55
.18
6
8.9
3
80
.89
4
3
Bo
bs
63
.22
3
.34
5
2.5
0
81
.00
6
2.5
5
54
.05
5
9.4
8
63
.76
7
6.1
4
7
Cro
tch
6
4.0
4
3.4
9
55
.65
7
1.3
6
66
.98
5
5.6
7
55
.73
7
0.4
7
71
.18
5
Ja
cks
(NO
RT
H B
AY
) 6
4.8
4
3.1
2
54
.01
7
2.7
6
66
.77
5
5.8
3
63
.11
6
7.5
6
71
.72
5
Ch
esle
y
67
.56
3
.57
5
5.6
5
79
.22
6
9.6
2
56
.53
6
1.3
8
71
.90
7
7.4
4
6
Sh
arb
ot
(200
0)
67
.67
4
.48
4
9.1
9
96
.99
6
8.4
0
49
.63
5
3.2
2
73
.94
9
6.1
7
13
SC
- 3
7
Ta
ble
SC
-9.
(co
ntin
ue
d)
La
ke
M
ea
n
Sta
nd
ard
Err
or
Min
imu
m
Maxim
um
Media
n
5%
Pe
rce
ntile
25
%Q
ua
rtile
75
%Q
ua
rtile
95
%P
erc
en
tile
Sa
mp
leS
ize
Da
lrym
ple
7
2.3
4
3.6
4
57
.24
1
02
.10
7
1.2
8
60
.82
6
5.1
0
74
.58
9
2.6
3
11
Sto
rmy
73
.12
1
.87
5
2.8
7
87
.74
7
4.6
8
61
.32
6
9.2
7
76
.52
8
7.1
2
19
Mo
un
tain
(19
98)
80
.32
9
.53
6
8.0
2
10
8.3
6
72
.44
6
8.1
2
68
.51
8
4.2
5
10
3.5
4
4
Ric
e
83
.48
1
.43
5
0.1
9
12
2.4
4
82
.86
6
0.7
9
76
.53
9
1.8
5
10
6.1
3
88
Sko
ota
ma
tta
8
5.0
4
2.9
1
54
.58
1
16
.53
8
3.6
8
61
.17
7
4.3
5
95
.16
1
05
.89
2
9
So
uth
-ce
ntr
al
reg
ion
wall
eye
re
lati
ve
fe
cu
nd
ity b
en
ch
ma
rks
.
Ave
rag
es
M
ean
Min
imu
m
Maxim
um
Media
n
5%
P
erc
en
tile
25
%Q
ua
rtile
75
%Q
ua
rtile
95
%P
erc
en
tile
Re
gio
n A
ve
rag
e1
.5
4.3
6
35
.44
7
8.1
7
53
.83
3
8.8
7
46
.39
6
0.7
4
71
.98
Re
gio
n A
ve
rag
e2
.5
4.5
2
31
.58
8
3.3
8
54
.09
3
6.1
6
45
.69
6
2.1
9
75
.13
1.
All
lakes c
om
bin
ed (
n=
36).
2.
Only
lakes w
ith
sam
ple
siz
e
10 (
n=
24).
SC
- 3
8
Ta
ble
SC
-10
. W
alle
ye
fe
cu
nd
ity-t
ota
l le
ng
th r
ela
tion
sh
ips f
or
so
uth
-ce
ntr
al
reg
ion
wa
terb
od
ies s
am
ple
d u
sin
g t
he
fa
ll w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
. A
ge
om
etr
ic m
ea
n p
red
ictive
re
gre
ssio
n m
od
el
wa
s f
itte
d t
o
log
ari
thm
ically
tr
an
sfo
rme
d
(Na
pie
ria
n
log
ari
thm
s)
ind
ivid
ua
l o
bse
rva
tio
ns.
To
tal
len
gth
in
m
illim
etr
es. F
ecu
nd
ity-l
en
gth
re
latio
n1
.C
alc
ula
ted
fe
cu
nd
ity a
t sta
nd
ard
to
tal le
ng
th (
mm
)
La
ke
x 1
0-6
R2
n
40
0
45
0
50
0
55
0
60
0
65
0
Pic
ke
rel R
ive
r (1
99
7)
2.6
9
45
71
.91
0
.72
9
4
57
30
6
27
78
8
33
50
1
07
71
1
13
61
20
1
68
82
6
Ho
lde
n
2.9
9
59
6.4
8
0.9
0
21
3
49
99
4
97
47
6
81
36
9
05
64
1
17
42
8
14
91
27
Cro
tch
3
.04
4
33
.88
0
.92
5
3
55
98
5
09
33
7
01
73
9
37
71
1
22
18
0
15
58
57
Mo
ira
3
.13
1
43
.68
0
.60
1
7
20
04
1
28
97
5
40
29
4
54
30
0
71
29
8
91
59
8
Sko
ota
ma
tta
3
.19
2
78
.57
0
.78
2
9
57
18
1
83
30
2
11
66
35
1
58
14
7
20
88
22
2
69
66
5
Pic
ke
rel R
ive
r (1
99
6)
3.3
3
68
.33
7
0.8
5
19
3
25
09
4
81
45
6
84
08
9
39
96
1
25
63
0
16
40
56
Sh
arb
ot
(200
0)
3.3
6
84
.70
4
0.6
8
13
4
59
17
6
81
82
9
71
09
1
33
72
0
17
90
76
2
34
27
2
Fre
nch
Riv
er
(199
4)
3.4
8
31
.25
2
0.7
8
70
3
46
25
5
21
43
7
52
05
1
04
74
2
14
17
33
1
87
19
8
Ric
e
3.5
5
26
.48
1
0.8
1
88
4
48
38
6
80
88
9
89
37
1
38
72
9
18
88
83
2
50
89
0
Fre
nch
Riv
er
(200
1)
3.6
6
7.0
19
0
.82
5
3
23
92
8
36
83
9
54
19
2
76
83
8
10
56
85
1
41
69
9
Allu
me
tte
3
.75
6
.11
1
0.8
0
43
3
54
69
5
51
80
8
19
37
1
17
16
5
16
24
02
2
19
29
5
Bo
bs
3.7
8
4.7
57
0
.98
7
3
34
20
5
21
89
7
77
56
1
11
52
4
15
50
13
2
09
85
2
Re
sto
ule
3
.82
3
.17
6
0.9
3
13
2
73
05
4
28
09
6
40
08
9
21
02
1
28
39
2
17
42
83
Sto
rmy
3.9
0
2.5
55
0
.96
1
9
36
37
3
57
59
3
86
88
0
12
60
19
1
76
96
6
24
18
42
Big
Gu
ll 3
.94
1
.69
1
0.9
1
10
2
94
38
4
67
97
7
08
44
1
03
08
7
14
51
86
1
98
94
6
No
sb
on
sin
g
3.9
9
0.7
54
0
.81
2
7
18
55
3
29
69
5
45
22
8
66
17
6
93
67
1
12
89
51
Fre
nch
Riv
er
(199
7)
4.0
5
0.7
54
0
.87
3
0
25
62
8
41
28
0
63
23
1
92
99
4
13
22
50
1
82
84
6
Pic
ke
rel R
ive
r (1
99
5)
4.0
5
0.6
88
0
.90
1
6
23
42
0
37
72
5
57
78
8
84
99
3
12
08
75
1
67
12
4
Fre
nch
Riv
er
(199
5)
4.0
8
0.5
60
0
.81
5
0
23
47
8
37
97
3
58
38
1
86
14
9
12
28
89
1
70
38
2
Fre
nch
Riv
er
(200
0)
4.1
3
0.4
81
0
.80
9
9
26
19
1
42
58
0
65
76
4
97
44
7
13
95
34
1
94
13
4
Ka
ka
kiw
ag
an
da
4
.18
0
.38
9
0.8
9
20
2
91
91
4
77
56
7
41
76
1
10
47
4
15
89
24
2
22
06
1
Ja
cks
(NO
RT
H B
AY
) 4
.20
0
.35
1
0.9
4
5
29
53
0
48
42
1
75
36
1
11
24
44
1
62
03
0
22
67
50
Ch
esle
y
4.2
2
0.3
68
0
.78
6
3
58
42
5
89
40
9
19
72
1
37
55
0
19
86
31
2
78
51
7
McF
arl
an
e
4.5
5
0.0
25
4
0.9
1
11
1
72
81
2
95
25
4
76
74
7
35
38
1
09
23
4
15
71
95
Da
lrym
ple
4
.67
0
.02
46
0
.90
1
1
34
62
3
60
00
6
98
13
9
15
31
46
2
29
90
2
33
40
79
La
Clo
ch
e (
1996)
5.0
7
0.0
01
45
0
.93
7
1
83
72
3
34
00
5
70
12
9
24
77
1
43
81
6
21
58
87
Go
uld
5
.38
0
.00
02
33
0
.71
8
2
37
37
4
47
50
7
89
06
1
31
80
5
21
05
49
3
23
95
5
Big
Cle
ar
6.2
1
0.0
00
00
03
0
.81
1
1
48
45
1
00
65
1
93
56
3
49
74
6
00
21
9
86
44
La
ng
7
.11
0
.00
00
00
00
10
.72
2
6
46
46
1
07
36
2
27
13
4
47
36
8
30
62
1
46
76
7
1. T
wo-t
aile
d S
tudent
t-te
sts
were
em
plo
yed t
o t
est
sig
nific
ance o
f slo
pe a
nd inte
rcept
(o:
or
= 0
).
SC
- 3
9
So
uth
-ce
ntr
al
reg
ion
wall
eye
fe
cu
nd
ity-t
ota
l le
ng
th b
en
ch
ma
rks
.
F
ecu
nd
ity-l
en
gth
re
latio
n
Ca
lcu
late
d fe
cu
nd
ity a
t sta
nd
ard
to
tal le
ng
th (
mm
)
x 1
0-6
R2
n
40
0
45
0
50
0
55
0
60
0
65
0
Me
an
4
.05
2
16
.05
0
.84
2
9
29
40
4
46
08
9
69
29
6
10
07
36
1
42
42
1
19
67
14
Min
um
um
2
.69
0
.00
00
00
00
10
.60
5
4
64
6
10
06
5
19
35
7
34
97
4
60
02
1
91
59
9
Ma
xim
um
7
.11
4
57
1.9
1
0.9
8
99
5
71
81
8
33
03
1
16
63
6
15
81
47
2
29
90
2
33
40
79
Me
dia
n
3.9
4
1.6
91
0
.82
1
7
29
43
8
47
75
7
70
17
4
97
44
8
13
95
34
1
87
19
8
5%
Pe
rce
ntile
2
.99
0
.00
00
00
3
0.6
8
5
48
45
1
07
37
2
27
13
4
47
36
7
12
99
9
86
45
25
% Q
ua
rtile
3
.48
0
.36
8
0.7
8
10
2
34
78
3
77
26
5
77
89
8
61
50
1
20
87
5
15
71
95
75
% Q
ua
rtile
4
.20
3
1.2
52
0
.91
2
9
35
46
9
55
18
1
81
93
7
11
71
65
1
62
40
2
22
67
50
95
% P
erc
en
tile
6
.21
5
96
.47
0
.96
8
8
45
91
7
68
18
2
98
93
8
15
31
46
2
10
55
0
32
39
55
Sa
mp
le S
ize
2
9
SC
- 4
0
Ta
ble
SC
-11
a.
Ma
ture
fe
ma
le w
alle
ye
fa
ll go
na
doso
ma
tic i
nde
x (
GS
I is
ova
ry w
eig
ht
exp
resse
d a
s a
pe
rcen
t o
f th
e t
ota
l w
eig
ht
min
us
th
e o
va
ry w
eig
ht)
, vis
cera
l fa
t in
de
x (
VF
I is
fa
t w
eig
ht
exp
resse
d a
s a
pe
rce
nt
of
the
to
tal
we
igh
t),
an
d r
ep
rod
uctive
in
de
x
(R
is 3
h/L
) d
ata
fo
r sou
th-c
en
tra
l re
gio
n w
ate
rbo
die
s s
am
ple
d u
sin
g t
he
fa
ll w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
. O
nly
me
an
va
lue
s w
ith
a m
inim
um
sa
mp
le s
ize
3
are
in
clu
de
d.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Do
llars
2
.24
%
0.3
9%
3
.69
%
5
0.8
8%
0
.41
%
1.6
0%
3
N
o d
ata
Go
Ho
me
2
.32
%
1.6
1%
3
.82
%
21
2
.33
%
0%
5
.22
%
21
N
o d
ata
De
er
2.3
7%
0
.58
%
3.7
5%
1
0
0.9
6%
0
%
1.9
5%
1
0
0.3
8
Ro
ck I
sla
nd
2.3
9%
1
.89
%
3.2
8%
5
1
.65
%
0.6
7%
3
.68
%
5
No
da
ta
Bra
nd
y
2.4
8%
1
.74
%
3.2
8%
3
2
.17
%
0.5
4%
3
.43
%
3
No
da
ta
Cro
tch
2
.55
%
0.2
2%
4
.67
%
8
0.4
8%
0
%
1.3
2%
7
0
.42
McK
ella
r 2
.57
%
2.1
9%
3
.17
%
3
1.3
5%
0
.72
%
2.1
1%
3
N
o d
ata
Go
va
n
2.7
2%
1
.27
%
3.9
2%
8
0
.96
%
0%
2
.02
%
8
0.5
9
Go
uld
2
.83
%
1.6
4%
4
.86
%
8
0.4
0%
0
%
0.9
9%
8
N
o d
ata
Ch
esle
y
2.9
9%
2
.12
%
3.8
1%
6
0
.84
%
0.3
4%
1
.37
%
6
0.5
5
Pic
ke
rel R
ive
r (1
99
5)
3.1
3%
1
.85
%
4.8
5%
1
6
3.7
5%
0
%
7.0
7%
1
6
0.4
1
Ba
lsa
m(2
00
1)
3.4
1%
2
.07
%
4.5
7%
7
0
.39
%
0.2
9%
0
.57
%
7
0.4
3
Bu
ll 3
.43
%
2.9
6%
3
.80
%
6
1.6
6%
0
.84
%
3.3
3%
6
N
o d
ata
Wo
lfe
3
.44
%
2.3
7%
5
.28
%
9
0.8
3%
0
%
2.8
7%
9
0
.59
Ce
ce
be
3
.57
%
2.5
3%
5
.91
%
13
1
.27
%
0%
2
.44
%
8
No
da
ta
Ke
nn
eb
ec
3.6
2%
2
.43
%
5.5
3%
5
1
.75
%
1.2
3%
2
.50
%
5
No
da
ta
Mo
un
tain
(19
98)
3.7
4%
0
.02
%
6.0
7%
5
0
.68
%
0.0
2%
1
.33
%
5
0.5
1
Sto
co
3
.75
%
2.9
0%
4
.96
%
4
0.6
8%
0
.42
%
1.1
7%
4
N
o d
ata
Fre
nch
Riv
er
(199
7)
3.9
3%
1
.80
%
8.1
5%
3
0
2.4
3%
0
%
5.4
7%
3
0
0.3
1
Th
ree
Mile
3
.93
%
2.7
0%
4
.41
%
5
1.7
5%
1
.36
%
2.3
8%
5
N
o d
ata
McF
arl
an
e
4.1
2%
2
.63
%
6.1
5%
1
1
5.2
8%
3
.78
%
6.6
5%
1
1
No
da
ta
Fre
nch
Riv
er
(199
5)
4.1
2%
1
.29
%
7.6
8%
5
0
3.6
7%
0
.87
%
8.7
8%
5
0
0.4
6
Oa
k
4.1
4%
2
.13
%
5.6
3%
1
1
0.6
0%
0
.05
%
1.9
4%
1
1
No
da
ta
SC
- 4
1
Ta
ble
SC
-11
a.
(co
ntin
ue
d)
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Fre
nch
Riv
er
(200
1)
4.1
6%
2
.24
%
7.0
7%
5
3
3.6
4%
0
.43
%
7.6
5%
5
3
0.4
9
Ja
cks
(BR
AC
EB
RID
GE
) 4
.17
%
3.3
9%
5
.31
%
3
1.1
1%
0
.32
%
1.8
1%
3
N
o d
ata
Bu
ck
4.2
3%
3
.66
%
4.8
6%
3
3
.01
%
1.9
5%
4
.85
%
3
No
da
ta
Fre
nch
Riv
er
(200
0)
4.3
3%
0
.34
%
8.6
6%
1
00
3
.92
%
0%
8
.17
%
10
0
0.4
4
Ca
no
nto
4
.44
%
3.7
5%
4
.85
%
3
1.0
1%
0
.66
%
1.6
7%
3
N
o d
ata
Mo
ira
4
.45
%
1.3
4%
7
.37
%
17
1
.40
%
0.4
7%
2
.3%
1
7
0.4
3
Isa
be
lla
4.4
8%
1
.67
%
7.0
6%
3
0
.83
%
0.1
4%
1
.88
%
3
No
da
ta
Ka
ka
kiw
ag
an
da
4
.49
%
2.8
7%
6
.81
%
20
4
.04
%
2.4
9%
6
.22
%
20
N
o d
ata
La
Clo
ch
e (
1996)
4.5
2%
2
.78
%
6.2
0%
7
5
.16
%
3.4
1%
6
.63
%
7
No
da
ta
Bo
bs
4.8
2%
3
.44
%
7.1
1%
7
1
.50
%
0.3
5%
2
.78
%
7
No
da
ta
Ja
ck
4.8
6%
4
.01
%
5.4
4%
5
1
.21
%
0.8
2%
2
.23
%
5
No
da
ta
Pic
ke
rel R
ive
r (1
99
6)
4.9
4%
2
.59
%
6.6
9%
1
9
2.4
1%
0
.12
%
4.2
0%
1
9
0.4
7
Big
Gu
ll 5
.10
%
3.6
9%
7
.49
%
10
1
.45
%
0.3
8%
2
.65
%
10
0
.49
Mis
sis
sip
pi
5.2
1%
4
.17
%
6.4
9%
3
0
.28
%
0.1
1%
0
.43
%
3
No
da
ta
Ja
cks
(NO
RT
H B
AY
) 5
.22
%
4.1
8%
5
.96
%
5
2.0
5%
1
.50
%
2.8
3%
5
N
o d
ata
Ho
lde
n
5.3
2%
0
.31
%
9.2
7%
2
2
3.4
9%
1
.59
%
7.5
5%
2
2
0.3
6
Pic
ke
rel R
ive
r (1
99
7)
5.4
3%
3
.68
%
6.8
8%
9
2
.92
%
0.6
9%
5
.01
%
9
No
da
ta
Sto
rmy
5.4
8%
3
.47
%
8.7
3%
1
9
0.8
6%
0
%
2.6
0%
1
9
No
da
ta
Ba
rk
5.4
9%
2
.60
%
8.5
3%
2
0
3.4
5%
2
.11
%
4.8
2%
2
0
No
da
ta
No
sb
on
sin
g
5.5
4%
3
.48
%
8.8
3%
2
8
0.9
5%
0
%
2.6
0%
1
7
0.4
0
Hu
ron
(200
1)
5.6
1%
3
.29
%
9.2
8%
2
1
3.7
5%
0
.62
%
9.1
1%
2
1
No
da
ta
Lo
ng
(SU
DB
UR
Y)
5.7
3%
2
.89
%
7.9
3%
1
2
3.1
8%
1
.90
%
5.1
9%
1
2
0.3
3
Hu
ron
(199
7)
5.9
7%
4
.25
%
7.2
9%
3
2
.86
%
1.4
7%
4
.78
%
7
0.4
4
Da
lrym
ple
6
.00
%
3.7
7%
7
.97
%
11
0
.42
%
0%
1
.32
%
11
N
o d
ata
Co
nse
co
n
6.0
2%
4
.07
%
7.2
5%
5
0
%
0%
0
%
5
No
da
ta
Allu
me
tte
6
.10
%
2.3
2%
1
2.2
2%
4
3
4.3
7%
0
.38
%
9.4
1%
4
3
0.4
2
SC
- 4
2
Ta
ble
SC
-11
a.
(co
ntin
ue
d)
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Be
nn
ett
6
.10
%
4.7
3%
7
.55
%
4
0.1
5%
0
.01
%
0.2
9%
4
N
o d
ata
Fo
ur
Mile
6
.19
%
4.5
4%
7
.62
%
5
0.4
1%
0
.07
%
0.7
7%
5
N
o d
ata
Min
k
6.3
6%
0
.07
%
9.3
6%
1
1
1.5
1%
0
.02
%
2.8
7%
1
1
0.5
6
Fre
nch
Riv
er
(199
4)
6.4
3%
3
.21
%
10
.94
%
70
3
.63
%
0.1
9%
7
.09
%
70
0
.37
Mo
un
tain
(20
01)
6.5
6%
5
.15
%
8.0
3%
3
0
.82
%
0.2
1%
1
.21
%
3
No
da
ta
Go
od
erh
am
7
.09
%
6.5
3%
7
.88
%
3
1.2
8%
1
.04
%
1.5
7%
3
N
o d
ata
Ge
org
es
7.1
5%
4
.48
%
9.6
9%
5
0
.54
%
0.1
9%
1
.11
%
5
No
da
ta
Ch
ristie
8
.11
%
6.7
5%
9
.05
%
6
0.2
5%
0
%
0.9
6%
6
N
o d
ata
Wilb
er
8.1
3%
6
.22
%
10
.89
%
3
1.3
8%
1
.27
%
1.4
4%
3
N
o d
ata
Sko
ota
ma
tta
8
.41
%
2.6
6%
1
1.9
5%
3
0
1.4
1%
0
.39
%
3.8
5%
2
7
0.6
3
Sh
arb
ot
(200
1)
9.6
8%
8
.29
%
11
.21
%
4
2.4
6%
1
.72
%
3.2
7%
4
N
o d
ata
So
uth
-ce
ntr
al
ma
ture
fe
ma
le g
on
ad
os
om
ati
c,
vis
ce
ral
fat,
an
d r
ep
rod
uc
tiv
e i
nd
ice
s b
en
ch
ma
rks
.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at In
de
x
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Re
gio
n A
ve
rag
e1
.4
.79
%
3.2
2%
6
.46
%
11
1
.64
%
0.6
7%
2
.90
%
11
0
.46
Re
gio
n A
ve
rag
e2
.4
.87
%
2.2
2%
7
.80
%
27
2
.57
%
0.6
3%
5
.08
%
26
0
.43
1.
All
lakes c
om
bin
ed (
n=
60).
2.
O
nly
la
ke
s w
ith
sam
ple
siz
e
10 (
n=
24).
SC
- 4
3
Ta
ble
SC
-11
b.
Ma
ture
ma
le w
alle
ye
fa
ll g
on
ad
oso
ma
tic i
nd
ex (
GS
I is
te
ste
s w
eig
ht
exp
resse
d a
s a
pe
rce
nt
of
the
to
tal
we
igh
t m
inu
s t
he
te
ste
s w
eig
ht)
, vis
ce
ral
fat
ind
ex (
VF
I is
fa
t w
eig
ht
exp
resse
d a
s a
perc
en
t o
f th
e t
ota
l w
eig
ht)
, a
nd
re
pro
du
ctive
in
de
x (
R
is
3h
/L)
da
ta f
or
so
uth
-ce
ntr
al re
gio
n w
ate
rbo
die
s s
am
ple
d u
sin
g t
he
fa
ll w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
. O
nly
me
an
va
lue
s w
ith
a m
inim
um
sa
mp
le s
ize
3
are
in
clu
de
d.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Stu
rge
on
(C
ha
lk)
1.7
1%
1
.27
%
2.2
9%
1
1
1.4
0%
0
.48
%
2.8
9%
1
1
No
da
ta
Ge
org
es
1.8
6%
1
.43
%
2.3
7%
3
1
.29
%
0.8
4%
1
.58
%
3
No
da
ta
Do
llars
1
.91
%
0.1
9%
3
.36
%
33
0
.46
%
0%
1
.38
%
25
0
.41
Se
co
nd
De
po
t 2
.09
%
0.3
0%
4
.13
%
5
0.3
5%
0
%
1.0
7%
5
N
o d
ata
Ro
ck I
sla
nd
2.1
6%
1
.16
%
4.0
2%
7
0
.94
%
0.0
3%
2
.61
%
7
No
da
ta
Ste
wa
rtvill
e(2
00
1)
2.1
7%
1
.54
%
2.8
0%
5
0
.39
%
0.0
3%
1
.18
%
5
No
da
ta
Bra
nd
y
2.2
4%
1
.25
%
3.3
9%
1
0
1.1
8%
0
.12
%
2.7
9%
1
0
No
da
ta
Ho
lde
n
2.3
6%
1
.38
%
3.7
1%
9
3
2.5
6%
0
.54
%
5.9
8%
9
3
0.4
9
Ma
da
wa
ska
2
.42
%
1.5
9%
3
.12
%
12
0
.37
%
0.0
2%
1
.49
%
12
N
o d
ata
Mis
sis
sip
pi
2.4
5%
1
.78
%
2.8
6%
9
0
.11
%
0.0
2%
0
.53
%
9
No
da
ta
Ce
ce
be
2
.50
%
0.2
9%
4
.30
%
26
0
.64
%
0%
2
.33
%
20
N
o d
ata
McF
arl
an
e
2.5
0%
0
.66
%
3.2
6%
1
2
2.7
2%
0
.73
%
4.2
4%
1
2
0.4
6
Be
nn
ett
2
.52
%
1.5
2%
4
.08
%
12
0
.19
%
0.0
2%
0
.70
%
12
N
o d
ata
Go
va
n
2.5
3%
0
.50
%
5.0
6%
3
4
0.6
4%
0
%
1.9
9%
3
4
0.6
1
Co
nsta
nt
2.5
4%
0
.25
%
3.9
8%
6
0
.52
%
0%
1
.17
%
6
No
da
ta
Allu
me
tte
2
.55
%
0.3
6%
5
.68
%
60
3
.18
%
0%
7
.85
%
60
0
.50
No
sb
on
sin
g
2.5
8%
1
.40
%
4.3
9%
3
2
0.2
4%
0
%
2.0
8%
3
2
0.5
0
Ch
esle
y
2.5
8%
1
.35
%
3.8
0%
1
7
0.9
2%
0
.19
%
2.0
6%
1
7
No
da
ta
Ce
nte
nn
ial
2.6
0%
1
.35
%
3.5
6%
1
6
0.4
5%
0
%
1.2
6%
1
6
No
da
ta
Ja
cks
(NO
RT
H B
AY
) 2
.61
%
1.2
7%
3
.77
%
30
1
.93
%
0.2
5%
4
.17
%
30
0
.66
Cro
tch
2
.65
%
0.4
3%
4
.26
%
34
0
.93
%
0.0
4%
2
.68
%
18
0
.42
Gir
ou
x
2.6
8%
1
.71
%
3.2
6%
4
1
.13
%
0.3
7%
1
.90
%
4
No
da
ta
Fre
nch
Riv
er
(199
7)
2.7
0%
0
.04
%
4.6
3%
8
9
1.4
5%
0
%
4.4
9%
8
9
0.3
9
SC
- 4
4
Ta
ble
SC
-11
b.
(co
ntin
ue
d)
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Sto
rmy
2.7
2%
0
.01
%
4.5
1%
2
4
0.3
1%
0
%
1.5
1%
2
4
No
da
ta
Da
lho
usie
2
.72
%
1.7
6%
3
.87
%
8
0.5
1%
0
%
1.2
9%
8
N
o d
ata
Wo
lfe
2
.72
%
0.2
6%
4
.13
%
71
0
.63
%
0%
2
.57
%
71
0
.53
Ja
cks
(BR
AC
EB
RID
GE
) 2
.76
%
1.4
9%
3
.46
%
19
0
.86
%
0%
2
.66
%
19
0
.56
Ja
ck
2.7
6%
1
.13
%
5.4
0%
1
3
0.4
8%
0
%
1.7
2%
1
2
No
da
ta
McK
ella
r 2
.78
%
2.0
5%
4
.09
%
4
0.5
8%
0
%
1.2
3%
4
N
o d
ata
Fre
nch
Riv
er
(199
5)
2.7
8%
0
.19
%
7.7
3%
7
9
2.9
6%
0
%
7.8
9%
7
9
0.5
2
Ric
ha
rd
2.8
3%
2
.42
%
3.1
9%
3
0
.97
%
0.2
9%
1
.81
%
5
No
da
ta
Lo
ng
(SU
DB
UR
Y)
2.8
3%
0
.23
%
4.5
1%
4
0
1.5
7%
0
%
4.7
6%
4
0
0.4
0
Da
lrym
ple
2
.84
%
1.8
4%
3
.90
%
14
0
.33
%
0%
1
.77
%
13
N
o d
ata
Hu
ron
(199
7)
2.8
9%
0
.99
%
4.0
3%
6
2
.61
%
0.9
7%
5
.64
%
10
0
.53
Go
uld
2
.91
%
0.0
9%
4
.57
%
15
0
.75
%
0.0
4%
3
.38
%
15
N
o d
ata
Pic
ke
rel R
ive
r (1
99
7)
2.9
3%
2
.16
%
4.3
3%
1
3
2.5
3%
0
.95
%
3.6
5%
1
3
No
da
ta
Be
ave
r 3
.01
%
0.2
9%
5
.94
%
27
0
.16
%
0%
1
.18
%
27
0
.52
Bu
ll 3
.04
%
0.2
0%
5
.24
%
15
0
.67
%
0%
2
.72
%
15
0
.60
Ba
lsa
m(2
00
1)
3.0
5%
1
.73
%
5.1
6%
2
4
0.2
9%
0
%
1.0
7%
2
4
0.4
4
Big
Gu
ll 3
.08
%
0.3
1%
4
.87
%
77
0
.91
%
0.0
2%
2
.39
%
77
0
.53
Fre
nch
Riv
er
(200
0)
3.1
2%
0
.12
%
6.0
2%
1
63
2
.56
%
0%
9
.99
%
16
3
0.5
3
Pic
ke
rel R
ive
r (1
99
6)
3.1
3%
0
.25
%
5.7
5%
4
6
2.3
6%
0
.29
%
5.4
3%
4
6
0.5
7
Co
nse
co
n
3.1
4%
1
.94
%
4.2
9%
2
7
0%
0
%
0%
2
7
0.4
6
Sto
co
3
.16
%
2.2
7%
3
.86
%
7
0.3
3%
0
.05
%
0.9
5%
7
N
o d
ata
Go
Ho
me
3
.16
%
1.2
2%
4
.76
%
29
1
.34
%
0%
3
.44
%
29
N
o d
ata
Ch
ristie
3
.17
%
1.7
3%
5
.19
%
14
0
.28
%
0%
1
.25
%
14
N
o d
ata
Isa
be
lla
3.1
7%
1
.71
%
4.5
9%
5
2
.94
%
1.6
3%
4
.20
%
5
No
da
ta
Fre
nch
Riv
er
(199
4)
3.1
9%
0
.37
%
5.3
1%
7
3
2.8
2%
0
%
6.6
5%
7
4
0.4
3
Fre
nch
Riv
er
(200
1)
3.1
9%
0
.23
%
6.1
2%
8
7
2.2
3%
0
%
5.2
5%
8
7
0.5
3
SC
- 4
5
Ta
ble
SC
-11
b.
(co
ntin
ue
d)
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Wilb
er
3.2
1%
2
.41
%
6.3
1%
8
1
.26
%
0.2
0%
2
.24
%
8
No
da
ta
Bu
ck
3.2
2%
2
.40
%
4.4
5%
1
2
0.5
8%
0
%
2.0
9%
1
2
No
da
ta
Mo
un
tain
(19
98)
3.2
3%
0
.04
%
5.5
3%
5
2
0.3
8%
0
%
1.8
1%
5
2
0.5
8
Sa
nd
y
3.2
4%
1
.39
%
4.7
2%
1
3
0.2
2%
0
.06
%
0.7
8%
1
2
0.7
1
Fo
ur
Mile
3
.27
%
0.0
7%
4
.93
%
33
0
.47
%
0.0
3%
1
.21
%
33
0
.43
Pic
ke
rel R
ive
r (1
99
5)
3.3
4%
1
.11
%
4.3
3%
1
1
4.0
8%
2
.51
%
6.4
0%
1
1
0.4
9
No
rca
n
3.4
1%
2
.87
%
4.1
0%
3
0
.44
%
0.2
1%
0
.61
%
3
No
da
ta
Min
k
3.4
4%
0
.67
%
5.5
5%
6
4
1.3
4%
0
.30
%
3.3
2%
6
4
0.6
5
De
er
3.5
1%
1
.34
%
5.0
0%
4
6
1.0
1%
0
%
2.5
1%
4
6
0.5
5
Ba
rk
3.5
3%
0
.63
%
4.9
2%
3
3
2.9
2%
0
%
5.8
1%
3
3
No
da
ta
Ke
nn
eb
ec
3.6
0%
0
.02
%
5.5
1%
1
8
1.4
3%
0
%
3.5
2%
1
8
No
da
ta
Fe
rgu
so
ns
3.6
1%
3
.06
%
4.1
5%
3
0
.76
%
0.3
0%
1
.28
%
3
No
da
ta
Go
od
erh
am
3
.71
%
2.3
0%
4
.84
%
7
0.7
6%
0
%
2.1
4%
7
N
o d
ata
Bo
bs
3.7
3%
0
.58
%
5.7
7%
1
7
1.8
6%
0
%
4.7
6%
1
8
0.5
0
Gib
so
n
3.7
8%
2
.72
%
4.7
4%
4
1
.25
%
0.7
4%
1
.64
%
4
No
da
ta
Mo
ira
3
.78
%
2.2
3%
6
.19
%
28
0
.85
%
0%
2
.15
%
28
0
.53
Mo
un
tain
(20
01)
3.8
4%
2
.35
%
4.8
7%
1
0
1.0
9%
0
.49
%
1.9
3%
1
0
No
da
ta
La
Clo
ch
e (
1996)
3.9
0%
2
.36
%
6.5
3%
1
2
4.5
2%
1
.79
%
7.7
3%
1
2
No
da
ta
Ca
no
nto
3
.95
%
2.9
4%
5
.20
%
9
0.5
6%
0
%
1.4
4%
9
N
o d
ata
Ka
ka
kiw
ag
an
da
4
.13
%
1.7
2%
6
.26
%
30
2
.89
%
1.0
5%
6
.85
%
30
N
o d
ata
Oa
k
4.2
6%
0
.30
%
7.5
0%
4
0
0.9
6%
0
%
4.0
0%
4
0
0.6
5
Sko
ota
ma
tta
4
.31
%
2.1
3%
6
.44
%
48
1
.82
%
0.2
6%
3
.58
%
40
0
.73
Cra
ne
4
.55
%
3.4
0%
5
.46
%
5
3.9
6%
2
.04
%
7.1
8%
5
N
o d
ata
Mis
sis
sa
ga
go
n
4.6
3%
1
.59
%
6.2
7%
8
1
.11
%
0%
2
.26
%
8
No
da
ta
De
nb
igh
4
.64
%
3.2
9%
6
.76
%
6
0.8
0%
0
%
1.8
9%
6
N
o d
ata
Th
ree
Mile
4
.90
%
3.0
0%
6
.33
%
8
1.2
5%
0
%
1.8
7%
8
N
o d
ata
SC
- 4
6
Ta
ble
SC
-11
b.
(co
ntin
ue
d)
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at in
de
x
La
ke
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Sh
arb
ot
(200
1)
5.1
6%
3
.13
%
7.2
7%
5
2
.35
%
0.8
1%
4
.04
%
5
No
da
ta
So
uth
-ce
ntr
al
ma
ture
ma
le g
on
ad
os
om
ati
c,
vis
ce
ral
fat,
an
d r
ep
rod
uc
tiv
e i
nd
ice
s b
en
ch
ma
rks
.
Go
na
do
so
ma
tic I
nd
ex
Vis
ce
ral F
at In
de
x
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Mean
Min
imu
m
Maxim
um
Sam
ple
Siz
e
Re
pro
du
ctive
In
de
x
Re
gio
n A
ve
rag
e1
.3
.17
%
1.4
9%
4
.73
%
25
1
.29
%
0.3
1%
2
.92
%
24
0
.53
Re
gio
n A
ve
rag
e2
.3
.02
%
0.9
6%
4
.85
%
36
1
.34
%
0.2
0%
3
.39
%
35
0
.53
1.
All
lakes c
om
bin
ed (
n=
76).
2.
O
nly
la
ke
s w
ith
sam
ple
siz
e
10 (
n=
52).
SC
- 4
7
Ta
ble
SC
-12
. W
alle
ye
life
his
tory
pa
ram
ete
rs f
or
On
tari
o w
ate
rbo
die
s s
am
ple
d u
sin
g th
e w
alle
ye
in
de
x n
ett
ing
sta
nd
ard
199
3 t
o 2
00
1.
Pe
rce
ntile
sL
ife
His
tory
Pa
ram
ete
r (S
am
ple
Siz
e)
Me
an
(±9
5%
C.I
.)1
.S
.D.
(C.V
.)2
.M
inim
um
to
Maxim
um
M
ed
ian
5%
2
5%
7
5%
9
5%
Fe
ma
le a
du
lt m
ort
alit
y (
A%
) (n
=2
96
)2
8(2
7-3
0)
13
.6(4
8%
)7
to
89
2
5
12
1
9
34
5
7
Ma
le a
du
lt m
ort
alit
y (
A%
) (n
=2
95
)3
0(2
9-3
2)
12
.8(4
2%
)6
to
82
2
8
14
2
1
38
5
1
Fe
ma
le r
ela
tive
co
nd
itio
n
(n=
24
1)
1.0
3(1
.02
-1.0
4)
0.0
79
(8%
)0
.90
to
1.4
9
1.0
2
0.9
4
0.9
8
1.0
7
1.1
6
Ma
le r
ela
tive
co
nd
itio
n
(n=
26
5)
1.0
3(1
.02
-1.0
4)
0.0
90
(9%
)0
.87
to
1.4
8
1.0
2
0.9
2
0.9
7
1.0
7
1.1
8
Fe
ma
le a
sym
pto
tic le
ngth
(L
, m
m)
(n=
23
8)
74
4(7
33
-75
5)
84
.2(1
1%
)5
29
to
95
7
74
8
60
6
68
4
80
4
88
0
Ma
le a
sym
pto
tic le
ng
th (
L,
mm
) (n
=2
52
)6
18
(61
0-6
26
)6
4.7
(10
%)
48
5 t
o 8
14
6
16
5
08
5
72
6
54
7
38
Fe
ma
le B
rod
y c
oe
ffic
ien
t (K
, yr-1
)(n
=2
38
)0
.15
8(0
.15
4-0
.16
3)
0.0
37
7(2
4%
)0
.09
0 t
o 0
.32
7
0.1
52
0.1
10
0.1
32
0.1
80
0.2
30
Ma
le B
rod
y c
oe
ffic
ien
t (K
, yr-1
)(n
=2
52
)0
.18
0(0
.17
6-0
.18
5)
0.0
35
0(1
9%
)0
.10
5 t
o 0
.32
00
.17
60
.12
90
.15
5
0.2
00
0.2
44
Fe
ma
le O
me
ga
(,
mm
•yr-1
)(n
=2
38
)1
16
.40
(11
3.4
1-1
19
.38
)2
3.3
22
(20
%)
73
.49
to
20
0.2
7
11
4.4
5
80
.84
9
9.3
1
13
2.8
2
15
5.9
2
Ma
le O
me
ga
(,
mm
•yr-1
)(n
=2
52
)1
10
.57
(10
8.1
2-1
13
.02
)1
9.7
48
(18
%)
69
.45
to
18
1.1
9
10
9.4
1
80
.67
9
7.2
0
12
2.0
7
14
8.0
6
Pre
-ma
tura
tio
n g
row
th (
h,
mm
•yr-1
)(n
=2
78
)9
3.0
0(9
0.4
8-9
5.5
1)
21
.30
6(2
3%
)5
4.3
1 t
o 1
85.9
0
92
.07
6
1.1
3
76
.59
1
05
.95
1
29
.45
Fe
ma
le a
ge
-at-
50
% m
atu
rity
(ye
ars
) (n
=2
61
)4
.5(4
.3-4
.7)
1.5
1(3
4%
)2
.0 t
o 1
0.2
4
.2
2.5
3
.4
5.4
7
.1
Ma
le a
ge
-at-
50
% m
atu
rity
(ye
ars
) (n
=2
66
)3
.0(2
.9-3
.2)
1.1
0(3
7%
)0
.8 t
o 7
.7
2.9
1
.5
2.2
3
.7
4.9
SC
- 4
8
Ta
ble
SC
-12
. (c
on
tin
ued
)
Pe
rce
ntile
sL
ife
His
tory
Pa
ram
ete
r (S
am
ple
Siz
e)
Me
an
(±9
5%
C.I
.)1
.S
.D.
(C.V
.)2
.M
inim
um
to
Maxim
um
M
ed
ian
5%
2
5%
7
5%
9
5%
Fe
ma
le s
ize
-at-
50
% m
atu
rity
(m
m)
(n=
26
1)
44
1(4
36
-44
7)
42
.1(1
0%
)3
38
to
58
6
44
0
36
9
41
6
47
3
50
4
Ma
le s
ize
-at-
50
% m
atu
rity
(m
m)
(n=
26
6)
34
9(3
45
-35
3)
32
.9(9
%)
24
6 t
o 4
76
3
46
2
99
3
31
3
70
4
01
Fe
ma
le g
on
ad
oso
ma
tic in
de
x (
%)
(n=
15
1)
4.4
(4.2
-4.7
)1
.64
(37
%)
1.5
to
11
.1
4.2
2
.1
3.3
5
.4
7.5
Ma
le g
on
ad
oso
ma
tic ind
ex (
%)
(n=
15
5)
2.9
(2.8
-3.0
)0
.76
(26
%)
1.3
to
5.4
2
.8
1.8
2
.5
3.2
4
.5
Re
lative
fe
cu
nd
ity (
eg
gs•g
-1)
(n=
13
6)
49
.79
(47
.41
-52
.17
)1
4.0
49
(28
%)
18
.04
to
87
.98
5
0.4
7
26
.22
40
.01
5
8.6
9
72
.65
Fe
ma
le v
isce
ral fa
t in
de
x (
%)
(n=
15
1)
1.9
(1.7
-2.1
)1
.29
(68
%)
0.1
to
6.8
1
.6
0.3
0
.8
2.7
4
.4
Ma
le v
isce
ral fa
t in
de
x (
%)
(n=
15
5)
1.4
(1.2
-1.5
)0
.98
(72
%)
0.1
to
5.3
1
.1
0.2
0
.6
1.8
3
.2
Re
lative
ab
un
da
nce
(n
um
be
r•n
et-1
)(n
=4
41
)7
.5(6
.9-8
.2)
7.1
3(9
5%
)0
to
36
.1
5.3
0
.4
2.0
1
0.9
2
2.5
1.
95%
Confidence inte
rval of
the m
ean.
2.
Sta
ndard
devia
tion o
f th
e m
ean (
S.D
.) a
nd c
oeffic
ient
of
variation o
f th
e m
ean e
xpre
ssed a
s a
perc
ent
(C.V
.).
