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i New York Natural Heritage Program A Partnership between the SUNY College of Environmental Science and Forestry and the NYS Department of Environmental Conservation 625 Broadway, 5 th Floor, Albany, NY 12233-4757 (518) 402-8935 Fax (518) 402-8925 www.nynhp.org Rare Animal Status List October 2017

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i

New York Natural Heritage Program A Partnership between the SUNY College of Environmental Science and Forestry and the NYS Department of Environmental Conservation

625 Broadway, 5th Floor, Albany, NY 12233-4757 (518) 402-8935 Fax (518) 402-8925 www.nynhp.org

Rare Animal Status List October 2017

ii

Established in 1985, the New York Natural Heritage Program (NYNHP) is a program of the State University of New York College of Environmental Science and Forestry (SUNY ESF). Our mission is to facilitate conservation of rare animals, rare plants, and significant ecosystems. We accomplish this mission by combining thorough field inventories, scientific analyses, expert interpretation, and the most comprehensive database on New York's distinctive biodiversity to deliver the highest quality information for natural resource planning, protection, and management. The Program is funded by grants and contracts from government agencies whose missions involve natural resource management, private organizations involved in land protection and stewardship, and both government and private organizations interested in advancing the conservation of biodiversity.

NY Natural Heritage is housed within NYS DEC’s Division of Fish, Wildlife & Marine Resources. The program is staffed by more than 25 scientists and specialists with expertise in ecology, zoology, botany, information management, and geographic information systems.

NY Natural Heritage maintains New York’s most comprehensive database on the status and location of rare species and natural communities. We presently monitor 181 natural community types, 850 rare plant species, and 481 rare animal species and assemblages across New York, keeping track of more than 13,800 locations where these important elements of biodiversity have been recorded. The database also includes detailed information on the relative rareness of each species and community, the quality of their occurrences, and descriptions of sites. The information is used by public agencies, the environmental conservation community, developers, and others to aid in land-use decisions. Our data are essential for prioritizing those species and communities in need of protection and for guiding land-use and land-management decisions where these species and communities exist.

In addition to tracking recorded locations, NY Natural Heritage has developed models of the areas around these locations important for conserving biodiversity, and models of the distribution of suitable habitat for rare species across New York State.

NY Natural Heritage also houses iMapInvasives, an online tool for invasive species reporting and data management.

NY Natural Heritage has developed two notable online resources: Conservation Guides include the biology, identification, habitat, and management of many of New York’s rare species and natural community types; and NY Nature Explorer lists species and communities in a specified area of interest.

The program is an active participant in the NatureServe Network – an international network of biodiversity data centers overseen by a Washington D.C. based non-profit organization. There are currently Natural Heritage Programs or Conservation Data Centers in all 50 states and several interstate regions. There are also 10 programs in Canada, and many participating organizations across 12 Latin and South American Countries. Our collaboration with NatureServe and other states helps us put our information into a broader context. With NatureServe, we track the rarity of species and natural communities at global and state scales, allowing us to distinguish conservation priorities for species with just a few populations in the world to other species with a few populations in New York but many populations elsewhere. We can also pool our data to look across state and international lines. For example, New York data on rare species and natural communities along Lake Ontario have been combined with similar data from Canada to facilitate analyses of potential consequences of lake-level changes. New York information has also been combined with data from neighboring states to help us understand the significance of our best biodiversity sites relative to similar systems in southeastern Canada, New England, the Mid-Atlantic states, and other Great Lakes states.

Learn more at www.nynhp.org.

iii

New York Natural Heritage Program

Rare Animal Status List

October 2017

The most recent version of this list may be found on our website: www.nynhp.org

For more information, suggestions, or comments about this list, contact

Matthew D. Schlesinger Chief Zoologist

New York Natural Heritage Program SUNY College of Environmental Science and Forestry

625 Broadway, 5th Floor Albany, NY 12233-4757

(518) 402-8939 [email protected]

To report sightings of rare species online, visit www.nynhp.org/report-rare

Cover photos (from top to bottom and left to right): Humpback Whales (Megaptera novaeangliae), Atlantic Coast Leopard Frog (Rana kauffeldi), Green Sea Turtle (Chelonia mydas; Leah J. Akins), Inland Barrens Buck Moth (Hemileuca maia), Bog Turtle (Glyptemys muhlenbergii), Timber Rattlesnake (Crotalus horridus), Mocha Emerald (Somatochlora linearis). All photos by Matthew D. Schlesinger except as noted.

iv

TABLE OF CONTENTS

Introduction ........................................................................................................................................................ v

Taxonomy and Nomenclature ..................................................................................................................... v

State & Federal Listings ................................................................................................................................ v

Global and Subnational Conservation Status Ranks ............................................................................... vi

Heritage-tracked Species .............................................................................................................................. ix

Active Inventory Species .................................................................................................................................. 1

Mammals ......................................................................................................................................................... 1

Birds ................................................................................................................................................................. 1

Reptiles ............................................................................................................................................................ 3

Turtles .............................................................................................................................................................. 3

Amphibians ..................................................................................................................................................... 3

Ray-finned Fishes ........................................................................................................................................... 4

Lampreys ......................................................................................................................................................... 5

Gastropods ..................................................................................................................................................... 6

Bivalves ............................................................................................................................................................ 6

Malacostracans ............................................................................................................................................... 7

Insects .............................................................................................................................................................. 7

Mayflies ....................................................................................................................................................................... 7

Dragonflies and Damselflies .................................................................................................................................... 8

Stoneflies ................................................................................................................................................................... 10

Beetles, Weevils ........................................................................................................................................................ 10

Caddisflies ................................................................................................................................................................. 10

Butterflies, Skippers, and Moths ........................................................................................................................... 10

Bees, Wasps, Ants, Sawflies ................................................................................................................................... 14

Animal Assemblages .................................................................................................................................... 15

Watch List Species ........................................................................................................................................... 16

Mammals ....................................................................................................................................................... 16

Birds ............................................................................................................................................................... 16

Reptiles .......................................................................................................................................................... 17

Turtles ............................................................................................................................................................ 18

Amphibians ................................................................................................................................................... 18

Ray-finned Fishes ......................................................................................................................................... 18

Lampreys ....................................................................................................................................................... 20

Gastropods ................................................................................................................................................... 20

Bivalves .......................................................................................................................................................... 20

Insects ............................................................................................................................................................ 20

Dragonflies and Damselflies .................................................................................................................................. 20

Beetles, Weevils ........................................................................................................................................................ 21

Butterflies, Skippers, and Moths ........................................................................................................................... 21

v

INTRODUCTION The following list is the New York Natural Heritage Program’s current assessment of the status

of New York’s rarest and most imperiled animal species. We have developed this list over the years in consultation with the Department of Environmental Conservation’s Endangered Species and Nongame Units, NatureServe (www.natureserve.org), researchers, conservation organizations, and knowledgeable amateur biologists. Population locations in our database are drawn from sources including museum specimens, personal contacts, and the scientific literature, and through extensive field research. This list is dynamic, meaning it changes as new discoveries are made or populations are lost. Check www.nynhp.org for the latest version of this list. TAXONOMY AND NOMENCLATURE

Species typically have two sets of names: scientific and common. Scientific names follow standard nomenclatures specific to the different groups of animals. The scientific names of birds, for example, follow the standards set by the American Ornithologists’ Union.

Common names are standardized in some cases, such as for birds, but are not as consistent across a species’ geographic range as scientific names. Further, some species have not yet been given common names; in the list below, these species are flagged with general names, such as “A Bluet Damselfy” or “A Noctuid Moth.”

In the following lists, major species groups (taxonomic Classes) are sorted in reverse taxonomic order; that is, groups with a longer evolutionary history are listed after groups that have evolved more recently. Thus, mammals (the most recently evolved taxonomic Class) are first, followed by birds, reptiles, turtles1, amphibians, fish, and multiple Classes of invertebrates. Within each of these taxonomic Classes (e.g., birds [Aves]), species are grouped into Orders (e.g., perching birds [Passeriformes]), and then Families (e.g., warblers [Parulidae]), each of which is sorted in taxonomic order. Within Families, species are sorted alphabetically by Genus and Species.

STATE & FEDERAL LISTINGS

NY Natural Heritage tracks a selected subset of New York’s animals. The species we track are chosen based on their degree of rarity or imperilment within the state, and as new information comes in, new species are sometimes added while others are discontinued. Information on the species and communities tracked by NY Natural Heritage are used for conservation, research, and regulatory purposes.

Many of the species tracked by NY Natural Heritage are listed as “endangered” or “threatened” under the state Environmental Conservation Law (E.C.L.). Listing is a legal process that is conducted by the state agency with authority over the species in question, and for animals confers important protection requirements. See http://www.dec.ny.gov/animals/7494.html for all state-listed animals.

The DEC Division of Fish, Wildlife, and Marine Resources has jurisdiction over rare animal species listed as “endangered,” “threatened,” or “special concern” under E.C.L. §11-0535. Animals listed as endangered or threatened receive notable legal protection, as it is illegal to take or possess any of these species or their parts without a permit from DEC. Species of special concern warrant attention and consideration but current information does not justify listing them as either endangered or threatened.

1 Note that in currently accepted taxonomy, turtles are distinct from reptiles.

vi

A subset of the animal species listed under New York state law is also recognized under federal law. These species are so seriously imperiled across their entire range that they face the very real prospect of extinction. Species are listed as federally endangered or threatened by the U.S. Fish & Wildlife Service in consultation with state agencies and other experts, and the Service works closely with DEC on the protection of federally listed species in New York.

Ultimately, protection of New York’s biodiversity lies with landowners and land managers regardless of state or federal listings. How private and public landowners manage their properties will determine what species and natural communities persist into the future. This situation is both a great opportunity and a serious challenge.

State legal listings are identified with the following codes:

E endangered

T threatened

SC special concern

Federal legal listings are identified with the following codes:

E listed endangered

T listed threatened

C candidate

Another state designation is Species of Greatest Conservation Need (SGCN), which was developed as part of the state’s wildlife action plan. In the following list, SGCN are coded as follows:

Y SGCN

Y-H High-priority SGCN

P Species of Potential Conservation Need See http://www.dec.ny.gov/animals/9406.html for a list of the state’s SGCN and definitions of

the three categories.

GLOBAL AND SUBNATIONAL CONSERVATION STATUS RANKS

NY Natural Heritage’s statewide inventory efforts revolve around lists of rare species and all types of natural communities known to occur, or to have historically occurred, in the state. These lists are based on a variety of sources including museum collections, scientific literature, information from state and local government agencies, regional and local experts, and data from neighboring states.

Each rare species is assigned a rank based on its rarity, population trends, and threats. Like those in all state Natural Heritage Programs, NY Natural Heritage’s ranking system assesses rarity at two geographic scales: global and state. The global rank (G-rank) reflects the status of a species or community throughout its range, whereas the subnational rank (S-rank) indicates its status within New York. Global ranks are maintained and updated by NatureServe, which coordinates the network of Natural Heritage programs. Both global and subnational ranks are usually based on the range of the species or community, the number of occurrences, the viability of the occurrences, and the vulnerability of the species or community around the globe or across the state. As new data become available, the ranks may be revised to reflect the most current information. Subspecific taxa are also assigned a taxon rank, which indicates the subspecies’ rank throughout its range.

vii

For the most part, global and subnational ranks follow a straightforward scale of 1 (rarest/most imperiled) to 5 (common/secure), as follows:

G1, S1 Critically Imperiled—Critically imperiled globally or in the state/province because of

extreme rarity (often 5 or fewer occurrences) or because of some factor(s) such as very steep declines making it especially vulnerable to extinction or extirpation from the state/province.

G2, S2 Imperiled—Imperiled globally or in the state/province because of rarity due to very restricted range, very few populations (often 20 or fewer), steep declines, or other factors making it very vulnerable to global extinction or extirpation from the state/province.

G3, S3 Vulnerable—Vulnerable globally or in the state/province due to a restricted range, relatively few populations (often 80 or fewer), recent and widespread declines, or other factors making it vulnerable to extirpation.

G4, S4 Apparently Secure—Uncommon but not rare; some cause for long-term concern due to declines or other factors.

G5, S5 Secure—Common, widespread, and abundant globally or in the state/province. G#G#, S#S# Range Rank —A numeric range rank (e.g., S2S3) is used to indicate any range of

uncertainty about the status of the species or community. Ranges cannot skip more than one rank (e.g., SU is used rather than S1S4).

There are some additional codes:

GX, SX Presumed Extinct or Extirpated—Species is believed to be extinct in the wild (GX) or extirpated from the state/province (SX). Not located despite intensive searches of historical sites and other appropriate habitat, and virtually no likelihood that it will be rediscovered.

GH, SH Possibly Extinct or Extirpated (Historical)—Species or community occurred

historically in the state/province, and there is some possibility that it may be rediscovered. Its presence may not have been verified in the past 20-40 years. A species or community could become SH without such a 20-40-year delay if the only known occurrences in a the state/province were destroyed or if it had been extensively and unsuccessfully looked for. The SH rank is reserved for species for which some effort has been made to relocate occurrences, rather than simply using this status for all elements not known from verified extant occurrences.

GU, SU Unrankable—Currently unrankable due to lack of information or due to substantially conflicting information about status or trends.

SNA Not Applicable—A conservation status rank is not applicable because the species is not a suitable target for conservation activities.

SNR Unranked—State/province conservation status not yet assessed.

viii

Codes sometimes have qualifiers attached:

T1, T2, etc. These ranks, which like global and subnational ranks run from 1 (most imperiled) to 5

(secure), are attached to global ranks to indicate the status of a subspecies or variety

Q Indicates that the species, subspecies, or variety is in taxonomic dispute

? Inexact or Uncertain—Denotes inexact or uncertain numeric rank. (The ? qualifies the character immediately preceding it in the S-rank.)

N Nonbreeding—Conservation status refers to the non-breeding population of the species in the nation or state/province.

B Breeding—Conservation status refers to the breeding population of the species in the nation or state/province.

Examples of ranks

Global State Example Explanation

G5 S2 Devil Crawfish Common rangewide (G5), imperiled in NY (S2)

G3 S2 Bog Turtle Uncommon rangewide (G3), imperiled in NY (S2)

G5 S1S2 Swamp Darter Common rangewide (G5), uncertain whether critically imperiled (S1) or imperiled (S2) in NY

G5 S1S3 Watercress Snail Common rangewide, uncertain whether critically imperiled (S1), imperiled (S2), or uncommon (S3) in NY

GH SX Shortnose Cisco Historical rangewide (GH), extirpated from NY (SX)

GU SH Black Lordithon Rove Beetle Insufficient information for ranking the species’ rangewide status (GU), historical in NY (SH)

G4 S1? Chain Fern Borer Moth Apparently secure rangewide (G4), appears to be critically imperiled in NY (S1) but more information is needed (?)

G1G2 S1 Dwarf Wedgemussel Somewhere between critically imperiled (G1) and imperiled (G2) rangewide, critically imperiled in NY (S1)

G3G4 S1S3 Bird Dropping Moth Uncertain whether uncommon (G3) or apparently secure (G4) rangewide, uncertain whether critically imperiled (S1), imperiled (S2), or uncommon (S3) in NY

G5T2 S1 Karner Blue Butterfly The species is secure (G5) but the subspecies is imperiled rangewide (T2), the subspecies is critically imperiled in NY (S1)

G4T2T4 SH The Consort Underwing The species is apparently secure (G4) but the subspecies is either imperiled (T2), uncommon (T3), or apparently secure (T4) rangewide, the subspecies is historical in NY (SH)

G1Q S1 Bogbean Buckmoth Critically imperiled rangewide (G1) but the species is in taxonomic dispute (Q), critically imperiled in NY (S1)

ix

G5 S3B,S1N Least Bittern Common rangewide (G5), breeding populations (B) are uncommon in NY (S3) while non-breeding populations (N) are critically imperiled (S1) (in this case, overwintering birds)

G5 S2S3B,S2N Bald Eagle Common rangewide (G5), breeding populations (B) are either imperiled (S2) or uncommon (S3) in NY, while non-breeding birds (N) are imperiled in NY

HERITAGE-TRACKED SPECIES

NY Natural Heritage tracks all species listed as endangered and threatened. While we track many of the species listed as being of special concern, a subset of special concern species are currently not rare or imperiled enough to merit tracking at our precise scale. In addition, we track many species that are biologically rare and imperiled (nearly all S1 and S2 species, and some S3 species), but that have not gone through the review process necessary for state listing.

Our program keeps two lists of rare animal species: the Active Inventory List and the Watch List. Species on the Active Inventory List are ones we currently track in our database; for the most part these are the most rare or most imperiled species in the state. Species on the Watch List are those that could become imperiled enough in the future to warrant being actively inventoried, or are ones for which we do not have enough data to determine whether they should be actively inventoried. The lists are dynamic, as S-ranks change with unearthed historical information, field inventories reveal new populations or local extirpations, and state and federal listings are updated. Further, we are assessing the conservation status of new species groups, such as many marine taxa and insect pollinators, through ongoing projects that will undoubtedly add species to our lists.

NY Natural Heritage currently tracks 474 animal species (plus six animal assemblages, which are extraordinarily rare, diverse, or abundant aggregations of multiple animal species). The balance of species among taxonomic groups reflects both the degree of imperilment of various taxa and the level of available knowledge about different groups.

Active Inventory

List

Watch List

Mammals 17 12

Birds 53 48

Reptiles 15 9

Amphibians 6 4

Fish 59 50

Freshwater Snails 14 10

Freshwater Mussels 38 3

Other Non-insect Invertebrates 5 0

Dragonflies and Damselflies 65 34

Beetles 15 4

Butterflies and Skippers 29 5

Moths 123 29

Other Insects 35 0

Total 474 208

10/24/2017

CLASSCOMMON NAME

GLOBAL

RANK

STATE

RANK

New York Natural Heritage Program

Rare Animal Status - Active Inventory List

Order SGCN

FED

LISTINGSCIENTIFIC NAME

MORE

INFO

STATE

LISTING

Mammals

Rodents

Allegheny Woodrat G3G4 S1 ENeotoma magister Y-H

Rabbits and Hares

New England Cottontail G3 S1S2 SCSylvilagus transitionalis Y-H

Shrews and Moles

Least Shrew G5 SHCryptotis parva P

Bats

Silver-haired Bat G3G4 S2S3BLasionycteris noctivagans Y

Eastern Small-footed Myotis G4 S1S3 SCMyotis leibii Y

Northern Long-eared Bat G1G2 S1 T TMyotis septentrionalis Y-H

Indiana Bat G2 S1 E EMyotis sodalis Y-H

Tri-colored Bat G2G3 S1Perimyotis subflavus Y-H

Carnivores

Eastern Cougar G5THQ SX E EPuma concolor couguar

Least Weasel G5 S1Mustela nivalis P

Whales, Dolphins, and Porpoises

Northern Right Whale G1 SNA E EEubalaena glacialis Y-H

Sei Whale G3 SNA E EBalaenoptera borealis Y-H

Blue Whale G3G4 SNA E EBalaenoptera musculus Y-H

Fin Whale G3G4 S1 E EBalaenoptera physalus Y-H

Humpback Whale G4 SNA E EMegaptera novaeangliae Y

Bottlenose Dolphin G5 S3Tursiops truncatus

Sperm Whale G3G4 SNA E EPhyseter macrocephalus Y-H

Birds

Waterfowl

Trumpeter Swan G4 S1BCygnus buccinator

Harlequin Duck G4 S1NHistrionicus histrionicus Y

Ruddy Duck G5 S1Oxyura jamaicensis Y

Common Eider G5 S1B,S3?NSomateria mollissima Y

Fowls and Gallinaceous Birds

Spruce Grouse G5 S2 EFalcipennis canadensis Y-H

Grebes

Pied-billed Grebe G5 S3B,S1N TPodilymbus podiceps Y

Nightbirds

Chuck-will's-widow G5 S1BAntrostomus carolinensis

Whip-poor-will G5 S3B SCAntrostomus vociferus Y-H

Common Nighthawk G5 S2S3B SCChordeiles minor Y-H

Rails and Cranes

Black Rail G3G4 S1B ELaterallus jamaicensis Y-H

1

= NatureServe Explorer Species Account

= New York Natural Heritage Program Conservation Guide

CLASSCOMMON NAME

GLOBAL

RANK

STATE

RANK

New York Natural Heritage Program

Rare Animal Status - Active Inventory List

Order SGCN

FED

LISTINGSCIENTIFIC NAME

MORE

INFO

STATE

LISTING

Birds - Continued

Rails and Cranes - Continued

Clapper Rail G5 S2S3Rallus crepitans

King Rail G4 S1B TRallus elegans Y-H

Sandhill Crane G5 S1BGrus canadensis

Gulls, Plovers, and Shorebirds

Piping Plover G3 S3B E E,TCharadrius melodus Y-H

Upland Sandpiper G5 S3B TBartramia longicauda Y-H

Black Tern G4 S2B EChlidonias niger Y-H

Gull-billed Tern G5 S1Gelochelidon nilotica Y

Caspian Tern G5 S1Hydroprogne caspia Y

Laughing Gull G5 S1Leucophaeus atricilla Y

Black Skimmer G5 S2 SCRynchops niger Y-H

Roseate Tern G4 S1B E ESterna dougallii Y-H

Forster's Tern G5 S1Sterna forsteri Y

Common Tern G5 S3B TSterna hirundo Y

Least Tern G4 S3B TSternula antillarum Y

Loons

Common Loon G5 S4 SCGavia immer Y

Pelicans and Cormorants

Great Blue Heron G5 S5Ardea herodias (large rookeries only)

Cattle Egret G5 S2Bubulcus ibis Y-H

Little Blue Heron G5 S2Egretta caerulea Y

Snowy Egret G5 S2S3Egretta thula Y

Tricolored Heron G5 S2Egretta tricolor Y

Least Bittern G5 S3B,S1N TIxobrychus exilis Y

Yellow-crowned Night-Heron G5 S2Nyctanassa violacea Y

Glossy Ibis G5 S2Plegadis falcinellus Y

Hawks, Kites, Eagles and allies

Golden Eagle G5 SHB,S1N EAquila chrysaetos Y

Northern Harrier G5 S3B,S3N TCircus cyaneus Y

Bald Eagle G5 S2S3B,S2N THaliaeetus leucocephalus Y

Owls

Barn Owl G5 S1S2Tyto alba Y-H

Short-eared Owl G5 S2 EAsio flammeus Y-H

Woodpeckers

Red-headed Woodpecker G5 S2?B SCMelanerpes erythrocephalus Y-H

American Three-toed

Woodpecker

G5 S2Picoides dorsalis Y-H

Raptors

Peregrine Falcon G4 S3B EFalco peregrinus Y

2

= NatureServe Explorer Species Account

= New York Natural Heritage Program Conservation Guide

CLASSCOMMON NAME

GLOBAL

RANK

STATE

RANK

New York Natural Heritage Program

Rare Animal Status - Active Inventory List

Order SGCN

FED

LISTINGSCIENTIFIC NAME

MORE

INFO

STATE

LISTING

Birds - Continued

Perching Birds

Henslow's Sparrow G4 S3B TAmmodramus henslowii Y-H

Seaside Sparrow G4 S2S3B SCAmmodramus maritimus Y-H

Yellow-breasted Chat G5 S2?B SCIcteria virens Y-H

Loggerhead Shrike G4 S1B ELanius ludovicianus Y-H

Sedge Wren G5 S3B TCistothorus platensis Y-H

Bicknell's Thrush G4 S2S3B SCCatharus bicknelli Y-H

Kentucky Warbler G5 S2BGeothlypis formosa Y-H

Tennessee Warbler G5 S2BOreothlypis peregrina P

Prothonotary Warbler G5 S2BProtonotaria citrea Y-H

Bay-breasted Warbler G5 S2BSetophaga castanea Y-H

Cape May Warbler G5 S2BSetophaga tigrina Y-H

Rusty Blackbird G4 S2BEuphagus carolinus Y-H

Reptiles

Lizards and Snakes

Fence Lizard G5 S1 TSceloporus undulatus Y

Coal Skink G5 S2S3Plestiodon anthracinus Y

Eastern Wormsnake G5 S2 SCCarphophis amoenus Y

Queen Snake G5 S1 ERegina septemvittata Y-H

Timber Rattlesnake G4 S3 TCrotalus horridus Y-H

Eastern Massasauga G3 S1 E CSistrurus catenatus catenatus Y-H

Turtles

Turtles

Loggerhead G3 S1N T TCaretta caretta Y-H

Green Turtle G3 S1N T TChelonia mydas Y-H

Hawksbill Sea Turtle G3 SNA E EEretmochelys imbricata

Kemp's or Atlantic Ridley G1 S1N E ELepidochelys kempii Y-H

Leatherback G2 S1N E EDermochelys coriacea Y-H

Blanding's Turtle G4 S2S3 TEmydoidea blandingii Y-H

Bog Turtle G3 S2 E TGlyptemys muhlenbergii Y-H

Eastern Mud Turtle G5 S1 EKinosternon subrubrum Y-H

Spiny Softshell G5 S2S3 SCApalone spinifera Y-H

Amphibians

Frogs and Toads

Northern Cricket Frog G5 S1 EAcris crepitans Y-H

Atlantic Coast Leopard Frog G3G4 S1S2Lithobates kauffeldi Y-H

Eastern Spadefoot G5 S2S3 SCScaphiopus holbrookii Y

Salamanders

Tiger Salamander G5 S1S2 EAmbystoma tigrinum Y-H

3

= NatureServe Explorer Species Account

= New York Natural Heritage Program Conservation Guide

CLASSCOMMON NAME

GLOBAL

RANK

STATE

RANK

New York Natural Heritage Program

Rare Animal Status - Active Inventory List

Order SGCN

FED

LISTINGSCIENTIFIC NAME

MORE

INFO

STATE

LISTING

Amphibians - Continued

Salamanders - Continued

Hellbender G3G4 S2 SCCryptobranchus alleganiensis Y-H

Longtail Salamander G5 S2S3 SCEurycea longicauda Y-H

Ray-finned Fishes

Paddlefishes, Spoonfishes, and Sturgeons

Shortnose Sturgeon G3 S1 E EAcipenser brevirostrum Y

Lake Sturgeon G3G4 S2S3 TAcipenser fulvescens Y

Atlantic Sturgeon G3 S1 EAcipenser oxyrinchus Y-H

Paddlefish G4 SXPolyodon spathula Y-H

Gars

Shortnose Gar G5 SXLepisosteus platostomus

Mooneyes

Mooneye G5 S2 THiodon tergisus Y-H

Anchovies and Herrings

Hickory Shad G4 S2Alosa mediocris

Minnows and Suckers

Finescale Dace G5 S2S3Chrosomus neogaeus

Lake Chub G5 S2S3Couesius plumbeus Y

Silverjaw Minnow G5 S2Ericymba buccata

Streamline Chub G4 S1 SCErimystax dissimilis

Gravel Chub G4 SH TErimystax x-punctatus

Tonguetied Minnow G4 S2Exoglossum laurae Y

Brassy Minnow G5 S2Hybognathus hankinsoni

Bigeye Chub G5 S1S2Hybopsis amblops Y-H

Redfin Shiner G5 S1S2 SCLythrurus umbratilis Y

Silver Chub G5 SH EMacrhybopsis storeriana

Comely Shiner G5 S2?Notropis amoenus Y-H

Pugnose Shiner G3 S1S2 ENotropis anogenus Y

Bridle Shiner G3 S2?Notropis bifrenatus Y

Ironcolor Shiner G4 S1 SCNotropis chalybaeus Y

Bigmouth Shiner G5 S2Notropis dorsalis Y

Blackchin Shiner G5 S1Notropis heterodon Y

Blacknose Shiner G5 S2S3Notropis heterolepis Y

Silver Shiner G5 S2Notropis photogenis

Swallowtail Shiner G5 S2Notropis procne Y-H

Summer Sucker G2G3 S2Catostomus utawana Y-H

Lake Chubsucker G5 SH TErimyzon sucetta

River Redhorse G4 S1Moxostoma carinatum Y-H

Black Redhorse G5 S2 SCMoxostoma duquesnei Y

4

= NatureServe Explorer Species Account

= New York Natural Heritage Program Conservation Guide

CLASSCOMMON NAME

GLOBAL

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Ray-finned Fishes - Continued

Catfishes

Black Bullhead G5 S1Ameiurus melas Y

Salmon and Trout

Bloater G4 SXCoregonus hoyi Y-H

Kiyi G3G4 SXCoregonus kiyi

Shortnose Cisco GH SXCoregonus reighardi

Shortjaw Cisco G3 SXCoregonus zenithicus

Round Whitefish G5 S1S2 EProsopium cylindraceum Y

Perches

Western Pirate Perch G5T5 S1Aphredoderus sayanus gibbosus Y

Eastern Pirate Perch G5T5 S1S2Aphredoderus sayanus sayanus

Silversides

Inland Silverside G5 S2S3Menidia beryllina Y

Atlantic Silverside G5 S2S3Menidia menidia Y

Needlefishes

Atlantic Needlefish G5 S2S3Strongylura marina

Killfishes

Spotfin Killifish G4 S1Fundulus luciae Y

Sculpins and Scorpion Fishes

Spoonhead Sculpin G5 SH ECottus ricei

Deepwater Sculpin G5 S1 EMyoxocephalus thompsonii Y

Perch-like Fishes

Mud Sunfish G4G5 SH TAcantharchus pomotis

Banded Sunfish G5 S1 TEnneacanthus obesus Y

Northern Sunfish G5 S1 TLepomis peltastes Y-H

Eastern Sand Darter G4 S2S3 TAmmocrypta pellucida Y

Bluebreast Darter G4 S1 EEtheostoma camurum Y-H

Iowa Darter G5 S2Etheostoma exile Y

Swamp Darter G5 S1 TEtheostoma fusiforme Y

Spotted Darter G2G3 S1 TEtheostoma maculatum Y-H

Variegate Darter G5 S2Etheostoma variatum

Gilt Darter G4 SH EPercina evides Y-H

Longhead Darter G3 S2 TPercina macrocephala

Sauger G5 S1Sander canadensis Y-H

Lampreys

Lampreys

Ohio Lamprey G3G4 S2Ichthyomyzon bdellium Y

Northern Brook Lamprey G4 S2Ichthyomyzon fossor

Mountain Brook Lamprey G4 S1 SCIchthyomyzon greeleyi Y

Gastropods

5

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Gastropods - Continued

Neotaenioglossa

Campeloma Spire Snail G5 S1Cincinnatia integra P

Watercress Snail G5 S1S3Fontigens nickliniana P

Buffalo Pebblesnail G5 S1 SCGillia altilis P

Canadian Duskysnail G4 SNRLyogyrus walkeri P

Gravel Pyrg GU SHMarstonia letsoni P

Heterostropha

Fringed Valvata G5 S1 SCValvata lewisi P

Purplecap Valvata G2G3 SNAValvata perdepressa P

Mossy Valvata G5 S1 SCValvata sincera P

Basommatophora

Spindle Lymnaea G3 SNRAcella haldemani P

Bugle Fossaria GH SNRFossaria cyclostoma

Coldwater Pondsnail G2G3Q SNRStagnicola woodruffi P

Lance Aplexa G5 S2Aplexa elongata P

Banded Physa GU S1Physella vinosa P

File Rams-horn G4G5Q SHPlanorbella pilsbryi P

Stylommatophora

Chittenango Ovate Amber Snail G1 S1 E TNovisuccinea chittenangoensis Y-H

Bivalves

Freshwater Mussels

Eastern Pearlshell G4 S2Margaritifera margaritifera Y

Mucket G5 S1S2Actinonaias ligamentina Y

Dwarf Wedgemussel G1G2 S1 E EAlasmidonta heterodon Y-H

Brook Floater G3 S1 TAlasmidonta varicosa Y-H

Slippershell Mussel G4G5 S1S2Alasmidonta viridis Y-H

Threeridge G5 S1Amblema plicata Y

Alewife Floater G5 S1S2Anodonta implicata Y-H

Northern Riffleshell G2T2 SNA EEpioblasma torulosa rangiana

Snuffbox G3 SHEpioblasma triquetra

Wabash Pigtoe G5 S2Fusconaia flava Y-H

Pink Mucket G2 SH E ELampsilis abrupta

Yellow Lampmussel G3G4 S3Lampsilis cariosa Y

Wavyrayed Lampmussel G5 S1 TLampsilis fasciola Y-H

Pocketbook G5 S2S3Lampsilis ovata Y

Yellow Sandshell G5 SHLampsilis teres

White Heelsplitter G5 SHLasmigona complanata

Green Floater G3 S1S2 TLasmigona subviridis Y-H

Fragile Papershell G5 S3Leptodea fragilis

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Bivalves - Continued

Freshwater Mussels - Continued

Tidewater Mucket G3G4 S1Leptodea ochracea Y-H

Eastern Pondmussel G4 S2S3Ligumia nasuta Y

Black Sandshell G4G5 S2S3Ligumia recta Y-H

Hickorynut G4 S1Obovaria olivaria P

Round Hickorynut G4 SHObovaria subrotunda

Clubshell G1G2 S1 E EPleurobema clava Y-H

Round Pigtoe G4G5 S1Pleurobema sintoxia Y

Pink Heelsplitter G5 S2S3Potamilus alatus Y

Fat Pocketbook G2 SH E EPotamilus capax

Pink Papershell G5 SNAPotamilus ohiensis

Kidneyshell G4G5 S2Ptychobranchus fasciolaris Y

Pimpleback G5 SHQuadrula pustulosa

Mapleleaf G5 SHQuadrula quadrula

Salamander Mussel G3 SHSimpsonaias ambigua

Lilliput G5 S1Toxolasma parvum Y

Fawnsfoot G5 SHTruncilla donaciformis

Deertoe G5 S1Truncilla truncata Y-H

Paper Pondshell G5 S1Utterbackia imbecillis Y

Rayed Bean G2 S1 E CVillosa fabalis Y-H

Rainbow G5Q S2S3Villosa iris Y-H

Malacostracans

Amphipods

Allegheny Cave Amphipod G5 SNRStygobromus allegheniensis

Taconic Cave Amphipod G2G3 SNRStygobromus borealis

Piedmont Groundwater Amphipod G4T4 SNRStygobromus tenuis tenuis P

Decapods

Devil Crawfish G5 S2Cambarus diogenes P

Insects

Mayflies

G4 SNRAmeletus tertius P

G2G3 SNRBaetis rusticans P

G3G4 SNRDannella provonshai Y

G2G3 SNREpeorus punctatus Y

G1Q SNREpeorus suffusus Y

G3 SNREurylophella bicoloroides P

G2G3 SNRHeptagenia culacantha

G4 SNRHeptagenia julia

7

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Insects - Continued

Mayflies - Continued

G5 SNRNixe rusticalis P

G4 SNRProcloeon mendax P

G2G4 SNRProcloeon ozburni P

G3G4 SNRProcloeon simile P

G2G3 SNRProcloeon vicinum P

G3G4 SNRRhithrogena anomala P

G3G4 SNRRhithrogena uhari

Tomah Mayfly G2G3 S1 ESiphlonisca aerodromia Y-H

G3 SNRSiphlonurus barbaroides Y

G1G3 SNRSiphlonurus barbarus Y

G5 SNRSparbarus maculatus

Dragonflies and Damselflies

Appalachian Jewelwing G4 SH

Sparkling Jewelwing G5 SH

Southern Spreadwing G5 S2S3 P

Lyre-tipped Spreadwing G5 S2S3 Y

Seepage Dancer G4 SH

Dusky Dancer G5 S1 Y

Subarctic Bluet G5 S1 P

Atlantic Bluet G5 S1S2 Y

New England Bluet G3G4 S3 Y

Little Bluet G4 S1 T Y-H

Scarlet Bluet G3 S2 T Y-H

Pine Barrens Bluet G3 S1 T Y-H

Blackwater Bluet G5 S1 Y

Rambur's Forktail G5 S2S3 Y

Southern Sprite G5 S1 SC Y

Gray Petaltail G4 S2 SC Y-H

Azure Darner G5 SNR

Zigzag Darner G5 S1

Subarctic Darner G5 S1 P

Comet Darner G5 S2S3 Y

Cyrano Darner G5 S2S3 Y

Spatterdock Darner G4 S2 Y

Horned Clubtail G4 S1 Y

Midland Clubtail G5 S3 Y

Spine-crowned Clubtail G4 S1

Calopteryx angustipennis

Calopteryx dimidiata

Lestes australis

Lestes unguiculatus

Argia bipunctulata

Argia translata

Coenagrion interrogatum

Enallagma doubledayi

Enallagma laterale

Enallagma minusculum

Enallagma pictum

Enallagma recurvatum

Enallagma weewa

Ischnura ramburii

Nehalennia integricollis

Tachopteryx thoreyi

Aeshna septentrionalis

Aeshna sitchensis

Aeshna subarctica

Anax longipes

Nasiaeschna pentacantha

Rhionaeschna mutata

Arigomphus cornutus

Gomphus fraternus

Gomphus abbreviatus Y

8

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Insects - Continued

Dragonflies and Damselflies - Continued

Rapids Clubtail G3G4 S3Gomphus quadricolor Y

Sable Clubtail G4 S1Gomphus rogersi Y-H

Delaware River Clubtail G2T1 S1 SCGomphus septima delawarensis Y-H

Cobra Clubtail G5 S1Gomphus vastus P

Skillet Clubtail G3 S1Gomphus ventricosus Y-H

Green-faced Clubtail G3G4 S1Gomphus viridifrons Y-H

Southern Pygmy Clubtail G4 S1Lanthus vernalis P

Extra-striped Snaketail G4 S2S3 SCOphiogomphus anomalus Y

Brook Snaketail G4 S3Ophiogomphus aspersus Y

Boreal Snaketail G5 S1Ophiogomphus colubrinus Y-H

Pygmy Snaketail G3 S1 SCOphiogomphus howei Y-H

Common Sanddragon G5 S1 SCProgomphus obscurus Y

Riverine Clubtail G4 SHStylurus amnicola

Elusive Clubtail G3 SHStylurus notatus

Russet-tipped Clubtail G5 S1Stylurus plagiatus Y

Tiger Spiketail G4 S1Cordulegaster erronea Y-H

Arrowhead Spiketail G4 S3Cordulegaster obliqua Y

Mantled Baskettail G5 S2Epitheca semiaquea Y

Broad-tailed Shadowdragon G3G4 S1Neurocordulia michaeli P

Umber Shadowdragon G5 S1Neurocordulia obsoleta Y

Ringed Emerald G5 SHSomatochlora albicincta

Lake Emerald G5 S1Somatochlora cingulata P

Forcipate Emerald G5 S1Somatochlora forcipata P

Delicate Emerald G5 S1Somatochlora franklini P

Hine's Emerald G2G3 SNA ESomatochlora hineana

Incurvate Emerald G4 S1Somatochlora incurvata P

Kennedy's Emerald G5 S1S2Somatochlora kennedyi

Mocha Emerald G5 S2S3Somatochlora linearis Y

Ocellated Emerald G5 S2S3Somatochlora minor P

Ebony Boghaunter G4 S1Williamsonia fletcheri P

Ringed Boghaunter G3 SHWilliamsonia lintneri

Four-spotted Pennant G5 S1Brachymesia gravida Y

Martha's Pennant G4 S2Celithemis martha

Double-ringed Pennant G5 S1Celithemis verna Y

Seaside Dragonlet G5 S2Erythrodiplax berenice Y

Blue Corporal G5 S2S3Ladona deplanata

Golden-winged Skimmer G5 S1S2Libellula auripennis

9

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Insects - Continued

Dragonflies and Damselflies - Continued

Yellow-sided Skimmer G5 S1Libellula flavida P

Needham's Skimmer G5 S3Libellula needhami Y

Black Meadowhawk G5 S2S3Sympetrum danae P

Stoneflies

Illinois Stonefly G3 SNRAllocapnia illinoensis Y

Notched Forestfly G4 SNROstrocerca complexa

Lawrence Sallfly G3 SNRAlloperla voinae P

Scotia Sallfly G3 SNRAlloperla vostoki P

Gaspe Sallfly G3 SNRUtaperla gaspesiana P

Spiny Salmonfly G3 SNRPteronarcys comstocki P

Beetles, Weevils

Eastern Pinebarrens Tiger Beetle G3G4 SHCicindela abdominalis

Appalachian Tiger Beetle G3 S2Cicindela ancocisconensis Y-H

Northeastern Beach Tiger Beetle G3G4T2 SX T TCicindela dorsalis dorsalis

Hairy-necked Tiger Beetle G5 S1S2Cicindela hirticollis Y-H

Cobblestone Tiger Beetle G2 S1Cicindela marginipennis Y-H

New Jersey Pine Barrens Tiger

Beetle

G3T1T3 SHCicindela patruela consentanea

Northern Barrens Tiger Beetle G3T3 S1Cicindela patruela patruela Y-H

Puritan Tiger Beetle G1G2 SNA TCicindela puritana

One-spotted Tiger Beetle G4G5 SHCicindela unipunctata P

Sylvan Hygrotus Diving Beetle GU S1Hygrotus sylvanus P

American Burying Beetle G2G3 SH E ENicrophorus americanus

Black Lordithon Rove Beetle GU S1Lordithon niger

Two-spotted Lady Beetle GNR S2Adalia bipunctata Y-H

Nine-spotted Lady Beetle G2 S1Coccinella novemnotata Y-H

Three-banded Lady Beetle GNR S2S3Coccinella trifasciata Y-H

Caddisflies

A Caddisfly GH SHCeraclea albosticta

Butterflies, Skippers, and Moths

Chestnut Clearwing Moth G3G5 SHSynanthedon castaneae

Arogos Skipper G3T1T2 SH EAtrytone arogos arogos

Dusted Skipper G4G5 S2S3Atrytonopsis hianna

Brazilian Skipper G5 SHCalpodes ethlius

Mottled Duskywing G3 S1 SCErynnis martialis Y-H

Persius Duskywing G5T1T3 S1 EErynnis persius persius Y-H

Broad-winged Skipper G5T4 S3Poanes viator viator

Southern Grizzled Skipper G1G2Q SH EPyrgus wyandot Y-H

10

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Insects - Continued

Butterflies, Skippers, and Moths - Continued

Giant Swallowtail G5 SNAPapilio cresphontes

Olympia Marble G5 S1 SCEuchloe olympia P

West Virginia White G3? S3Pieris virginiensis

Checkered White G5 S1 SCPontia protodice Y

Henry's Elfin G5 S1 SCCallophrys henrici Y-H

Hessel's Hairstreak G3G4 S1 ECallophrys hesseli Y-H

Frosted Elfin G3 S1S2 TCallophrys irus Y-H

Bog Elfin G3G4 S1Callophrys lanoraieensis

Red-banded Hairstreak G5 SUCalycopis cecrops

Appalachian Azure G4 S1S3Celastrina neglectamajor

Silvery Blue G5T3T4 SHGlaucopsyche lygdamus

lygdamus

White-m Hairstreak G5 SUParrhasius m-album

Karner Blue G5T2 S1 E EPlebejus melissa samuelis Y-H

Edwards' Hairstreak G5 S3S4Satyrium edwardsii

Northern Oak Hairstreak G4G5T4 S2S4Satyrium favonius ontario Y-H

Northern Metalmark G3G4 S1Calephelis borealis Y-H

Tawny Emperor G5 S2S4Asterocampa clyton

Gorgone Checkerspot G5 S1Chlosyne gorgone P

Jutta Arctic G5 S1Oeneis jutta Y

Tawny Crescent G4G5T1 SH SCPhyciodes batesii batesii

Hoary Comma G5 S3Polygonia gracilis

Regal Fritillary G3 SH ESpeyeria idalia

Pin-striped Slug Moth G4G5 S1Monoleuca semifascia P

Stinging Rose Caterpillar Moth G4 S1Parasa indetermina P

A Geometer Moth G3 S1S2Apodrepanulatrix liberaria

Projecta Gray G4 SUCleora projecta

A Geometrid Moth G4 S1Digrammia denticulata P

Three-lined Angle Moth G4 SNADigrammia eremiata

Honey-streak G4G5 SUDigrammia mellistrigata

Broad-lined Catopyrrha G3G4 S1S2Erastria coloraria

The Little Beggar G4 SHEubaphe meridiana

A Geometrid Moth G4 S1Euchlaena madusaria P

Blueberry Gray G4 S1S3Glena cognataria P

Buchholz's Gray G3G4 SHHypomecis buchholzaria

Umber Moth G4 SUHypomecis umbrosaria

Black-dotted Ruddy GNR S1Ilexia intractata

11

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Insects - Continued

Butterflies, Skippers, and Moths - Continued

Dark-ribboned Wave GNR SULeptostales rubromarginaria

Woolly Gray G4 SHLycia ypsilon

Jack Pine Looper G5 S1Macaria marmorata P

Barrens Metarranthis Moth G1G3 SHMetarranthis apiciaria

A Geometrid Moth G4 SHNemoria bifilata

A Geometrid Moth GU S1Nepytia pellucidaria

Chalky White G4 SNAScopula purata

Barrens Itame G3G4 S1S3Speranza exonerata P

Faded Gray Geometer GU S1Stenoporpia polygrammaria

Dimorphic Gray G4? SHTornos scolopacinaria

Melsheimer's Sack Bearer G4 S1Cicinnus melsheimeri Y

Dot-lined White G5 SHArtace cribraria

Orange-striped Oakworm Moth G5 SUAnisota senatoria

Spiny Oakworm Moth G5 SUAnisota stigma

Regal Moth G4G5 S1Citheronia regalis Y

Pine Devil G4G5 S1Citheronia sepulcralis

Imperial Moth G5T5 SUEacles imperialis imperialis

Inland Barrens Buckmoth G5T5 S1S2 SCHemileuca maia maia Y

Coastal Barrens Buckmoth G5T3 S2 SCHemileuca maia ssp. 5

Bogbean Buckmoth G1Q S1 EHemileuca sp. 1 Y-H

Graceful Clearwing G3G4 SUHemaris gracilis

Gordian Sphinx G4G5 S1S3Sphinx gordius

A Hand-maid Moth G3G4 S1S3Datana ranaeceps Y

A Prominent Moth G3G4 S1S2 SCHeterocampa varia P

Plain Schizura G3G4 SUSchizura apicalis

Four-lined Chocolate G4 SUArgyrostrotis quadrifilaris

Quiet or Sweet Underwing G3 SHCatocala dulciola

Herodias or Pine Barrens

Underwing

G3T3 S1S2 SCCatocala herodias gerhardi Y

Jersey Jair Underwing G4?T4? S1S2 SCCatocala jair ssp. 2

Precious Underwing G4T2 SHCatocala pretiosa pretiosa

An Underwing Moth G5 SHCatocala umbrosa

Packard's Lichen Moth G5 SUCisthene packardii

Pine Tussock Moth G4 SUDasychira pinicola

Shadowy Arches G4 S1S2Drasteria adumbrata P

Anna Tiger Moth G5 SUGrammia anna

Phyllira Tiger Moth G4 SHGrammia phyllira

12

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Insects - Continued

Butterflies, Skippers, and Moths - Continued

Yellow-spotted Graylet G4 SUHyperstrotia flaviguttata

Two-striped Cord Grass Moth G3G4 S1S3Macrochilo bivittata

Richard's Fungus Moth G4 SUMetalectra richardsi

Black Fungus Moth G4 SHMetalectra tantillus

A Tussock Moth G3G4 SHOrgyia detrita

Chocolate Renia G4 SURenia nemoralis

Orange Holomelina G5 SUVirbia aurantiaca

Black-eyed Zale G4 SUZale curema

A Zale Moth G4 S1Zale largera

Pine Barrens Zale G3G4 SUZale lunifera

Pine Barrens Zanclognatha G4 S1S2Zanclognatha martha P

A Notodontid Moth G4 S1Paectes abrostolella P

Coastal Heathland Cutworm G4T3 S1S3Abagrotis nefascia benjamini P

Well-marked Cutworm G5 S1Abagrotis orbis P

Barrens Dagger Moth G3G4 S1Acronicta albarufa

Doll's Merolonche G3G4 SHAcronicta dolli

Oblique Dart G4 S1Agrotis obliqua P

An Ear Moth GUT1Q S1Amphipoea erepta ryensis P

Burgess's Apamea G4 SUApamea burgessi

Irregular Apamea GU S1Apamea inordinata P

Sand Wainscot Moth G4 SUApamea lintneri

Straight Lined Mallow Moth G4 SUBagisara rectifascia

Bird Dropping Moth G3G4 S1S2Cerma cora

Waxed Sallow G3G4 S1S3Chaetaglaea cerata Y

A Noctuid Moth G3G4Q S1Chytonix ruperti

Sensitive Chytonix G4 S1S3Chytonix sensilis Y

The Pink Streak G3G4 SUDargida rubripennis

Pink Star Moth G4 S1Derrima stellata

Switchgrass Dart G4G5 S2S3Dichagyris acclivis P

Fringed Dart G4 S1Eucoptocnemis fimbriaris Y

A Dart Moth G5T5 SHEuxoa adumbrata thanatologia

Fawn Brown Dart G4 S2S3Euxoa pleuritica P

Violet Dart G4 SUEuxoa violaris

Marsh Fern Moth G4 S1S3Fagitana littera P

A Noctuid Moth G4 SHFishia yosemitae

Hairy Hydraecia G4 S1S3Hydraecia stramentosa

Pine Pinion Moth G4T1T3 S1 ELithophane lepida lepida P

13

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Insects - Continued

Butterflies, Skippers, and Moths - Continued

Thaxter's Pinion Moth G4 SULithophane thaxteri

Pale Green Pinion Moth G5 S1Lithophane viridipallens P

Black-bordered Lemon Moth G5 S1Marimatha nigrofimbria P

An Apamea Moth GU SHMelanapamea mixta

Gray Woodgrain G4G5 S1S3Morrisonia mucens P

Bridgham's Brocade G4 SUOligia bridghamii

A Notodontid Moth G4 S1?Orthodes obscura Y

A Borer Moth GH SHPapaipema aerata

Pitcher Plant Borer Moth G4 SUPapaipema appassionata

Yellow Stoneroot Borer G2G4 SHPapaipema astuta

Aweme Borer Moth G1 SHPapaipema aweme

Golden Borer Moth G2G4 SHPapaipema cerina

Seaside Golden Borer Moth G4 S1Papaipema duovata

Heracleum Stem Borer Moth G4? SHPapaipema harrisii

A Borer Moth G4 SHPapaipema marginidens

Maritime Sunflower Borer Moth G3 SHPapaipema maritima

Culvers Root Borer G3 SHPapaipema sciata

Ostrich Fern Borer Moth G3G4 S1S3Papaipema sp. 2 nr. pterisii

Chain Fern Borer Moth G4 S1?Papaipema stenocelis P

A Noctuid Moth G4 SNAPhotedes enervata

Spartina Borer Moth G3G4 SUPhotedes inops

Thaxter's Sallow G4 SHPsaphida thaxterianus

Pink Sallow G3 S2Psectraglaea carnosa Y

Annointed Sallow Moth GU SXPyreferra ceromatica

Aureolaria Seed Borer G3G4 SUPyrrhia aurantiago

Slender Flower Moth G4Q S1Schinia bifascia P

Spinose Flower Moth G4 SUSchinia spinosae

Golden Aster Flower Moth G4 S2Schinia tuberculum P

Variable Sallow G5 S1Sericaglaea signata

Hairy Artesta G5 S1S3Sideridis artesta

Toothed Apharetra G4 S2S4Sympistis dentata

Scribbled Sallow Moth G4 S1Sympistis perscripta P

Dune Sympistis G4 SUSympistis riparia

A Dart Moth G4 S1S2Xestia homogena

Acadian Swordgrass Moth G4 S1S2Xylena thoracica P

Bees, Wasps, Ants, Sawflies

Rusty-patched Bumble Bee G1 SHBombus (Bombus) affinis Y-H

14

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Insects - Continued

Bees, Wasps, Ants, Sawflies - Continued

Yellowbanded Bumble Bee G5 S1Bombus (Bombus) terricola Y-H

Ashton's Cuckoo Bumble Bee G4 SHBombus (Psithyrus) ashtoni Y-H

Fernald's Cuckoo Bumble Bee G5 SHBombus (Psithyrus) fernaldae

Indiscriminate Cuckoo Bumble

Bee

G4G5 SHBombus (Psithyrus) insularis

Northern Amber Bumble Bee G4G5 S1Bombus (Subterraneobombus)

borealis

Y-H

Yellow Bumble Bee G4? S1Bombus (Thoracobombus)

fervidus

Y-H

American Bumble Bee G3G4 S1Bombus (Thoracobombus)

pensylvanicus

Y-H

Black and Gold Bumble Bee G4G5 S1Bombus auricomus P

Animal Assemblages

GNR S3Anadromous Fish Concentration Area

GNR SNRBat Colony

GNR S3Colonial Waterbird Nesting Area

GNR SNRGull Colony

GNR SNRRaptor Winter Concentration Area

GNR SNRSeal Haul-out Site

GNR S3S4Waterfowl Winter Concentration Area

15

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Mammals

Rodents

Eastern Fox Squirrel G5 S3Sciurus niger

Carnivores

Gray Wolf G5 SX E ECanis lupus

Canada Lynx G5 SX T TLynx canadensis

Even-toed Hoofed Mammals

Moose G5 S3S4Alces americanus Y

Whales, Dolphins, and Porpoises

Minke Whale G5 SNRNBalaenoptera acutorostrata

Harbor Porpoise G4G5 S4 SCPhocoena phocoena Y-H

Pygmy Sperm Whale G4 SNRNKogia breviceps

Blainsville's Beaked Whale G4 SUMesoplodon densirostris

Gervais's Beaked Whale G3 SUMesoplodon europaeus

True's Beaked Whale G3 SUMesoplodon mirus

Cuvier's Beaked Whale G4 SUZiphius cavirostris

Birds

Waterfowl

Northern Pintail G5 S1B,S3NAnas acuta Y

American Wigeon G5 S3Anas americana

Northern Shoveler G5 S2Anas clypeata

Green-winged Teal G5 S3Anas crecca

Blue-winged Teal G5 S2S3B,SNRNAnas discors Y

American Black Duck G5 S3B,SNRNAnas rubripes Y-H

Common Goldeneye G5 S3,SNRNBucephala clangula Y

Red-breasted Merganser G5 S3Mergus serrator

Rails and Cranes

American Coot G5 S3Fulica americana

Common Moorhen G5 S3S4Gallinula galeata

Gulls, Plovers, and Shorebirds

American Oystercatcher G5 S3Haematopus palliatus Y

Wilson's Phalarope G5 S1BPhalaropus tricolor

Frigatebirds, Boobies, Cormorants and allies

Double-crested Cormorant G5 S3Phalacrocorax auritus

Pelicans and Cormorants

American White Pelican G4 S1BPelecanus erythrorhynchos

Great Egret G5 S4Ardea alba Y

American Bittern G4 S4 SCBotaurus lentiginosus Y

Black-crowned Night-Heron G5 S3Nycticorax nycticorax Y

Hawks, Kites, Eagles and allies

Osprey G5 S4B SCPandion haliaetus

16

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Birds - Continued

Hawks, Kites, Eagles and allies - Continued

Cooper's Hawk G5 S4 SCAccipiter cooperii

Northern Goshawk G5 S3S4B,S3N SCAccipiter gentilis Y

Sharp-shinned Hawk G5 S4 SCAccipiter striatus

Red-shouldered Hawk G5 S4B SCButeo lineatus Y

Owls

Northern Saw-whet Owl G5 S3Aegolius acadicus

Long-eared Owl G5 S2S3B,SNRNAsio otus Y

Woodpeckers

Black-backed Woodpecker G5 S3?Picoides arcticus

Raptors

Merlin G5 S3?BFalco columbarius

Perching Birds

Saltmarsh Sharp-tailed Sparrow G4 S3BAmmodramus caudacutus Y-H

Grasshopper Sparrow G5 S3B SCAmmodramus savannarum Y-H

Vesper Sparrow G5 S3B SCPooecetes gramineus Y-H

Clay-colored Sparrow G5 S3S4BSpizella pallida

Olive-sided Flycatcher G4 S3BContopus cooperi Y-H

Yellow-bellied Flycatcher G5 S3BEmpidonax flaviventris

Acadian Flycatcher G5 S3BEmpidonax virescens

Philadelphia Vireo G5 S3BVireo philadelphicus

Gray Jay G5 S3Perisoreus canadensis Y-H

Boreal Chickadee G5 S3Poecile hudsonicus

Horned Lark G5 S3S4B SCEremophila alpestris Y-H

Brown Thrasher G5 S3S4BToxostoma rufum Y-H

Gray-cheeked Thrush G5 SNACatharus minimus

Red Crossbill G5 S2S3Loxia curvirostra

White-winged Crossbill G5 S2S3Loxia leucoptera

Northern Parula G5 S3S4BSetophaga americana

Cerulean Warbler G4 S3?B SCSetophaga cerulea Y

Palm Warbler G5 S2S3BSetophaga palmarum

Blackpoll Warbler G5 S3BSetophaga striata

Golden-winged Warbler G4 S3B SCVermivora chrysoptera Y-H

Boat-tailed Grackle G5 S3BQuiscalus major

Blue Grosbeak G5 SNAPasserina caerulea

Reptiles

Lizards and Snakes

Five-lined Skink G5 S3Plestiodon fasciatus

Eastern Hog-nosed Snake G5 S3 SCHeterodon platirhinos Y-H

17

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Reptiles - Continued

Lizards and Snakes - Continued

Short-headed Gartersnake G4 S3Thamnophis brachystoma Y

Copperhead G5 S3Agkistrodon contortrix Y

Turtles

Turtles

Spotted Turtle G5 S3 SCClemmys guttata Y-H

Wood Turtle G3 S3 SCGlyptemys insculpta Y-H

Common Map Turtle G5 S3Graptemys geographica Y

Diamondback Terrapin G4 S3Malaclemys terrapin Y

Eastern Box Turtle G5 S3 SCTerrapene carolina Y-H

Amphibians

Frogs and Toads

Boreal Chorus Frog G5 S2S3Pseudacris maculata P

Western Chorus Frog G5 S2S3Pseudacris triseriata Y

Salamanders

Marbled Salamander G5 S3 SCAmbystoma opacum Y

Wehrle's Salamander G4 S3Plethodon wehrlei

Ray-finned Fishes

Gars

Longnose Gar G5 S3Lepisosteus osseus

Eels

American Eel G4 S2S3Anguilla rostrata Y-H

Anchovies and Herrings

Bay Anchovy G5 S3Anchoa mitchilli Y

Blueback Herring G3G4 S3 SCAlosa aestivalis Y

Gizzard Shad G5 S3Dorosoma cepedianum

Minnows and Suckers

Redside Dace G3G4 S3Clinostomus elongatus

Satinfin Shiner G5 S3Cyprinella analostana

Striped Shiner G5 S3Luxilus chrysocephalus

Hornyhead Chub G5 S3Nocomis biguttatus

Sand Shiner G5 S3Notropis stramineus

Mimic Shiner G5 S3Notropis volucellus

Quillback G5 S3S4Carpiodes cyprinus

Longnose Sucker G5 S3Catostomus catostomus Y

Silver Redhorse G5 S3Moxostoma anisurum

Golden Redhorse G5 S3Moxostoma erythrurum

Greater Redhorse G4 S3S4Moxostoma valenciennesi

Catfishes

Stonecat G5 S3Noturus flavus

18

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Ray-finned Fishes - Continued

Catfishes - Continued

Tadpole Madtom G5 S3Noturus gyrinus

Brindled Madtom G5 S3S4Noturus miurus

Salmon and Trout

Cisco or Lake Herring G5 S3Coregonus artedi Y

Atlantic Salmon G5 S3Salmo salar Y

Mudminnows and Pikes

Eastern Mudminnow G5 S3Umbra pygmaea

Perches

Trout-perch G5 S3Percopsis omiscomaycus

Cods

Burbot G5 S3Lota lota

Atlantic Tomcod G5 S3Microgadus tomcod Y-H

Silversides

Brook Silverside G5 S3Labidesthes sicculus

Killfishes

Mummichog G5 S3Fundulus heteroclitus Y

Pipefishes and Sticklebacks

Brook Stickleback G5 S3Culaea inconstans

Blackspotted Stickleback G5 SUGasterosteus wheatlandi

Ninespine Stickleback G5 SUPungitius pungitius Y

Lined Seahorse GNR SUHippocampus erectus Y-H

Northern Pipefish G5 S3Syngnathus fuscus Y-H

Sculpins and Scorpion Fishes

Grubby G5 S3Myoxocephalus aenaeus

Perch-like Fishes

Bluespotted Sunfish G5 S3Enneacanthus gloriosus

Redbreast Sunfish G5 S3Lepomis auritus

Green Sunfish G5 S3Lepomis cyanellus

Greenside Darter G5 S3Etheostoma blennioides

Rainbow Darter G5 S3Etheostoma caeruleum

Banded Darter G5 S3Etheostoma zonale

Logperch G5 S3Percina caprodes

Channel Darter G4 S3Percina copelandi

Blackside Darter G5 S3Percina maculata

Shield Darter G5 S3Percina peltata

Freshwater Drum G5 S4Aplodinotus grunniens

Naked Goby G5 SUGobiosoma bosc

Seaboard Goby GNR SUGobiosoma ginsburgi

Flatfishes, Flounders, and Soles

19

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Ray-finned Fishes - Continued

Flatfishes, Flounders, and Soles - Continued

Winter Flounder G5 S3?Pseudopleuronectes americanus Y-H

Hogchoker G5 S3Trinectes maculatus

Lampreys

Lampreys

Silver Lamprey G5 S3Ichthyomyzon unicuspis

American Brook Lamprey G4 S3Lethenteron appendix

Gastropods

Neotaenioglossa

Piedmont Elimia G5 S3Elimia virginica

Sharp Hornsnail G5 S3Pleurocera acuta

Globe Siltsnail G4 S3Birgella subglobosus P

Slender Walker G5 S3Pomatiopsis lapidaria

Basommatophora

Graceful Fossaria G5Q S3Fossaria exigua

Pygmy Fossaria G5 S3Fossaria parva

A Fossaria Snail G5Q S3Fossaria rustica

Swamp Lymnaea G5 S3Lymnaea stagnalis

Glass Physa G5 SUPhysa skinneri

Vernal Physa G4 SNRPhysa vernalis

Bivalves

Freshwater Mussels

Spike G5 S3S4Elliptio dilatata

Creek Heelsplitter G5 S3S4Lasmigona compressa

Lake Floater GU SUPyganodon lacustris

Insects

Dragonflies and Damselflies

Superb Jewelwing G4 S3Calopteryx amata

American Rubyspot G5 S3Hetaerina americana Y

Emerald Spreadwing G5 S3Lestes dryas

Blue-fronted Dancer G5 S3Argia apicalis

Blue-tipped Dancer G5 S3Argia tibialis

Taiga Bluet G5 S3Coenagrion resolutum

Double-striped Bluet G5 S3Enallagma basidens

Boreal Bluet G5 S3Enallagma boreale

Turquoise Bluet G5 S3Enallagma divagans

Big Bluet G5 S3Enallagma durum

A Bluet Damselfly G4Q S3Enallagma vernale

Citrine Forktail G5 S3Ischnura hastata

20

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Insects - Continued

Dragonflies and Damselflies - Continued

Lilypad Forktail G5 S3Ischnura kellicotti

Mottled Darner G4G5 S4Aeshna clepsydra

Swamp Darner G5 S3Epiaeschna heros

Mustached Clubtail G4G5 S2S3Gomphus adelphus

Harpoon Clubtail G4 S3Gomphus descriptus

Northern Pygmy Clubtail G4G5 S3Lanthus parvulus

Riffle Snaketail G5 S2S3Ophiogomphus carolus

Maine Snaketail G4G5 S3Ophiogomphus mainensis

Zebra Clubtail G4G5 S3Stylurus scudderi

Arrow Clubtail G5 S3Stylurus spiniceps

Petite Emerald G5 S3Dorocordulia lepida

Spiny Baskettail G5 S3Epitheca spinigera

Uhler's Sundragon G5 S3Helocordulia uhleri

Stygian Shadowdragon G5 S3Neurocordulia yamaskanensis

Brush-tipped Emerald G5 S3Somatochlora walshii

Banded Pennant G5 S3Celithemis fasciata

White Corporal G5 S3Ladona exusta

Bar-winged Skimmer G5 SNALibellula axilena

Great Blue Skimmer G5 S3Libellula vibrans

Elfin Skimmer G4 S3Nannothemis bella P

Saffron-winged Meadowhawk G5 S3S4Sympetrum costiferum

Ruby Meadowhawk G5 S3Sympetrum rubicundulum

Beetles, Weevils

Ghost Tiger Beetle G3G4 S3Cicindela lepida

Margined Tiger Beetle G5 S3Cicindela marginata P

Eastern Red-bellied Tiger Beetle G5 S3Cicindela rufiventris

Festive Tiger Beetle G5T5 S3Cicindela scutellaris rugifrons

Butterflies, Skippers, and Moths

Dion Skipper G4 S3Euphyes dion

Falcate Orangetip G4G5 S3S4Anthocharis midea

Pink-edged Sulphur G5 S3S4Colias interior

Great Purple Hairstreak G4G5 SNAAtlides halesus

Hackberry Emperor G5 S3S4Asterocampa celtis

Black-waved Flannel Moth G5 SULagoa crispata

Smaller Parasa G4 SUParasa chloris

Brown-bordered Geometer G4 S2S4Eumacaria madopata Y

Sharp Green Hydriomena G4 SUHydriomena pluviata

21

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Insects - Continued

Butterflies, Skippers, and Moths - Continued

A Geometrid Moth G4 SUIdaea productata

Dusky Mid-summer Nepytia G4G5 SUNepytia sp. 1

Spring Canker Moth G4 SUPaleacrita merriccata

Imperial Moth G5T3T4 SNREacles imperialis pini Y

Wild Cherry Sphinx G3G5 SUSphinx drupiferarum

Unexpected Cycnia G4 SNACycnia inopinatus

Long-winged Doryodes G4 SUDoryodes grandipennis

Atlantic Graphic Moth G4T4 SUDrasteria graphica atlantica

A Tiger Moth G5 S3Grammia figurata

Thin-lined Owlet G4 SNAIsogona tenuis

Texas Mocis G5 SUMocis texana

A Noctuid Moth G3G4 S2S3Phoberia ingenua P

Gray-banded Zale G4 S3S4Zale squamularis

A Dart Moth G4 SUAgrotis stigmosa

An Apamea Moth G4 SUApamea relicina

A Noctuid Moth G4 S3S4Aplectoides condita

Speyer's Cucullia G4 SUCucullia speyeri

A Noctuid Moth G4 SUFeltia manifesta

Lemmer's Noctuid Moth G3G4 SNALithophane lemmeri

Footpath Sallow Moth G5 SUMetaxaglaea semitaria

Holly Sallow G5 SNAMetaxaglaea violacea

A Moth G5T4 S3Panthea furcilla australis

Northern Flower Moth G3G4 SUSchinia septentrionalis

The Maroonwing G4 S2S3Sideridis maryx P

Barrens Xylotype G4 S3S4Xylotype capax

22

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