Penyerapan Hara Oleh Akar Tumbuhan

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  • 8/10/2019 Penyerapan Hara Oleh Akar Tumbuhan

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    PM PSLP PPSUB MALANG 2011

    PENYERAPAN

    UNSUR HARA

    OLEH

    AKAR TUMBUHAN

    Penyunting:Soemarno

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    PENYERAPAN UNSUR HARA

    OLEH

    AKAR TUMBUHAN

    Cetakan : IMalangProgram Pascasarjana, Universitas Brawijaya, Malang

    Viii : 60 hlm : 20 cm x 25 cm

    Penyunting: Soemarno

    Tata Letak :

    Grafis & Desain Sampul : Soemarno

    Hak Cipta pada penulis.

    Cetakan Pertama Agustus 2011

    Penerbit : PPSUB

    Jl. Mayjen MT. Haryono 165, Malang

    Tel. 0341-571260

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    KATA PENGANTAR

    Dengan memanjatkan puji syukur kehadirat Allah swt, atas segala

    karunia-Nya, penyusunan buku Bahan Kajian mengenai PENYERAPAN

    UNSUR HARA OLEH AKAR TUMBUHAN ini dapat diselesaikan.

    Buku memori ini menyajikan data, informasi dan konsep-konsep

    tentang Pengelolaan Hara Tanaman yang relevan dengan filosofi

    Sustainable Crop Management.

    Data dan infromasi yang disajikan dalam tulisan ini dihimpun dari

    beragam pengalaman dan dari berbagai sumber yang dianggap valid,

    termasuk hasil-hasil pengkajian yang tersebar di berbagai referensi .

    Dengan telah selesainya Buku ini, perkenankanlah kami mengucapkan

    terima kasih yang sebesar-besarnya kepada semua pihak yang telah

    membantu, terutama kepada rekan-rekan sejawat yang telah secara

    tulus ikhlas memberikan informasi yang ada kaitannya dengan

    penyusunan buku ini.

    Semoga informasi yang terkandung dalam buku ini dapat dimanfaatkansebagaimana mestinya bagi pengembangan pembelajaran pada program

    pascasarjana di Universitas Brawijaya.

    Malang, Agustus 2011

    Penyunting

    Soemarno

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    DAFTAR ISI

    Pendahuluan .. 1

    Unsur Hara Tanaman .. 2

    Sistem Perakaran Tanaman .. 7

    Penyerapan air dari tanah .. 10Penyerapan Hara Tanaman .. 19

    Mekanisme Penyediaan Unsur Hara .. 22

    Mekanisme Penyerapan Hara .. 29

    Mikorizha .. 34Serapan Hara Tanaman (Nutrient Uptake) 41

    Hubungan Kandungan N dengan Hasil .. 47

    Efisiensi Pupuk N pada Padi .. 48

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    KETERSEDIAAN

    UNSUR HARA

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    ZONE UJUNG AKAR

    Plants absorb water through the entire surface - roots, stems and

    leaves. However, mainly the water is absorbed by roots. The area

    of young roots where most absorption takes place is the root hairzone. The root hairs are delicate structures which get continuously

    replaced by new ones at an average rate of 100 millions per day.

    The root hairs lack cuticle and provide a large surface area. They

    are extensions of the epidermal cells. They have sticky walls by

    which they adhere tightly to soil particles. As the root hairs are

    extremely thin and large in number, they provide enormous

    surface area for absorption. They take in water from theintervening spaces mainly by osmosis.

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    PERTUKARAN ION

    Ion exchange theory

    Both cations and anions have a tendency to getadsorbed on the surfaces of the cell walls, and

    exchange with ions present in the soil solution. This

    process of exchange between the adsorbed ions andions in solution is known as ion exchange.

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    SIKLUS UNSUR HARA

    Biogeochemical cycle of mineral nutrients in

    agricultural ecosystems

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    Types of roots:

    Seminal root - from a seed

    Adventitious root - from a

    stemFirst order lateral root - from

    a seminal or adventitious

    root

    Second order laterals, etc. -

    from first order laterals,

    which in turn produce third

    order laterals, and so on . . .Feeder roots - fine, relatively

    short-lived roots that

    acquire nutrients and water

    in the topsoil

    Primary roots - from primary

    growth by the apical

    meristem

    Secondary roots - mature,

    thicker "woody" roots with

    bark and additional vascular

    tissue

    Coarse roots - may live for along time and have roles in

    transport and mechanical

    support

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    KETERSEDIAAN

    UNSUR HARA

    Iron Uptake and TranslocationIron is primarily taken up by plants as the ferrous (Fe2+) iron.

    However, because most agricultural soils contain iron in the

    ferric (Fe3+) form, plant must somehow first solubilise the Fe3+

    and then reduce it to Fe2+so that it can pass through the root

    hair plasma membrane (plasmalemma).

    In dicotyledoneous species (ie most crops), iron uptake is an

    active, energy, requiring process.

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    THE UPTAKE OF WATER FROM THE SOIL TO THE

    LEAVES

    The cytoplasm of root hair cells is usually hypertonic to thesurrounding soil water. Hence, water enters the root hair cells by

    osmosis. The root hair cells are now hypotonic to the adjacent root

    cortex cells. Water then diffuses into these cells until it reaches the

    cortex region. Water flows through the parenchyma cells (cortex)

    until it reaches the endodermis through either the apoplastic pathway

    or the symplastic pathway . The Casparian strips (impermeable to

    water) prevent water from moving through the cell wall of theendodermal cells. The water concentration gradient which exists

    across the cortex creates a transpiration pull to direct water into the

    xylem vessels. Water finally reaches the mesophyll cells in theleaves where it will be lost through the stomata into the surroundings

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    Ion homeostasis: plants feel better

    with proper controlThe model plant species, thale cress (Arabidopsis thaliana,

    right) and rice (Oryza sativa, left), are displayed as a

    chimera. (A) OsHKT2;1 functions as a transporter for Na+uptake. (B) QSO2, a flavoprotein quiescin sulphydryl

    oxidase, is a new component in ion homeostasis at the root

    symplastxylem interface. (C) Downregulation of

    OST2/AHA1 H+-ATPase activity is a prerequisite for ABA-

    induced stomatal closure. ABA, abscisic acid; OST2/AHA,

    OPEN STOMATA 2.

    http://www.nature.com/embo

    r/journal/v8/n8/fig_tab/7401040_F1.html