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ORIGIN AND HISTORY OF LIFE
Origin of Life
Darwin’s theory is based around the idea of a common ancestor that all living organisms can be traced back to.
This ancestor is called the Last Universal Common Ancestor or LUCA.
Origin of Life
Since Darwin, advances in chemistry, evolutionary biology, paleontology, microbiology and other fields of science, have helped develop new hypothesis about the origin of life.
There is now a body of scientific evidence that life began 3.5-4 billion years ago and developed in 4 stages.
Origin of Life
Stage 1- Organic Monomers Simple organic molecules evolved from inorganic
monomers. Ex: Amino Acids and Nucleotides Stage 2- Organic Polymers
Organic monomers joined to form polymers. Ex. DNA, RNA, and proteins
Stage 3- Protocells Polymers became enclosed by membranes to
form the first cell precursors. Stage 4- Living Cells
Protocells began self replicating and gained other cellular properties.
Stage 1- Monomers
The Oparin- Haldane hypotheis, also known as the “primordial soup hypothesis” proposed that the lack of oxygen in the atmosphere gave ammonia (NH3) and Methane (CH4) strong reducing capability. Therefore, the redox reactions in the early atmosphere could have powered the chemical evolution of organic monomers from inorganic molecules.
Stage 1- Monomers
In 2008 a group of scientist re-tested the Miller-Urey experiment and found a greater variety of organic materials than Miller reported.
The iron-world hypothesis, proposed by Gunter Wachterschauser, stated that thermal vents emitted all the elements necessary for organic monomers to synthesize.
Stage 1- Monomers
Some scientist believe that the early organic monomers could have been brought to Earth by comets and meteorites.
Stage 2- Polymers
The Iron-sulfur World Hypothesis Research has found that organic
monomers will react in the presence of iron-nickel sulfides found at thermal vents.
Stage 2- Polymers
Protein-First Hypothesis Sidney Fox found that amino acids will
polymerize in the presence of dry heat. He proposed that when amino acids floating in the ocean were exposed to the sun, they formed proteionoids, small polypeptides that have some catalytic activity.
Stage 2- Polymers
RNA- First Hypothesis This hypothesis suggests that only RNA
was needed to progress towards the first cell(s).
In a study by Cech and altman, they discovered RNA can act as both a substrate and an enzyme. Based on their findings and the fact that some viruses have RNA, it would seem that RNA could have carried out processes of life associated with DNA and proteins today.
Stage 3-Protcells
Protocells are precursors to true cells. The formation of an outer membrane
was essential to regulation and maintenance of cellular functions.
The plasma membranes of today’s cells are formed from a phospholipid bilayer. Early membranes were likely composed of fatty acids, which are smaller than phospholipids but still have a hydrophobic tail and hydrophilic head.
Stage 3- The protocell
When put in water, fatty acids will assemble into small spheres call micelles with heads pointing out and tails pointing in.
If conditions are right, then they can form a bilayer called a vesicle.
Stage 3- The protocell
The first cells would have evolved some way to obtain nutrients.
Because they were formed by thermal vents, it’s probable that they carried out chemosynthesis (the synthesis of organic molecules from inorganic molecules and nutrients) before they evolved the ability to break down organic materials.
Stage 4- Self- Replication In modern cells, DNA directs protein
synthesis and information flows from DNA to RNA to protein.
It is possible that this sequence developed in stages. RNA-first Hypothesis- RNA would have first
regulated protein synthesis until DNA could evolve
Protein- first hypothesis- proteins developed first and then once a membrane was evolved could DNA and RNA evolve.
History of Life
Relative Dating of Fossils Even before the theory of evolution,
scientist were trying to age the strata layer
Geologists used index fossils (fossils used to identify deposits made at the same time) to relatively date the age of the layer
Absolute Dating Rely on radiometric techniques to assign
actual dates to fossils. Most common technique is Carbon-14
dating. Carbon-14 is a rare isotope of carbon that
has a half life of 5,730 years Only accurate for fossils up to 100,000.
The Precambrian Time
The precambrian time comprises about 87% of the geologic timescale.
This is when life began.
The first modern cells were probably prokaryotes, appearing approximately 3.5 bya. The first identifiable fossils are complex
prokaryotes The oldest were discovered in west Australia
date back 3.46 bya.
The Precambrian Time
Eukaryotic Cells Arise Eukaryotes originated around 2.1 bya
once oxygen was added to the atmosphere.
The nucleus may have developed by an invagination of the plasma membrane.
The endosymbiotic theory states that a nucleated cell engulfed prokaryotes, which then became organelles.
Evidence to support this theory Present day mitochondria and
chloroplasts have relatively the same size range as bacteria.
Mitochondria and chloroplasts have their own DNA and make some proteins.
Mitochondria and cholorplasts divide by binary fission.
Outer membrane of mitochondria resemble eukaryotic cells and the inner resemble bacterial cells
Multicellularity Arises Multicellular protists fossils have been
found in Canada dating back to 1.4 BYA These gave rise to the invertebrates,
whose fossils date back to 630-545 mya.
The Paleozoic Era
The paleozoic era lasted about 300 million years.
There were 3 major mass extinctions during this time period. 57% of marine species Over 50% coastal species Over 50% of corals
The Paleozoic Era
The Cambrian Period (542 mya) Referred to as the Cambrian Explosion All of today’s animals can trace their
ancestry back to this time. Animals had a hard exoskeleton.
The Paleozoic Era
Life began to move onto land around 500 mya.
Algae started invading damp areas on land during the Ordovician period.
Insects entered the fossil records during the Carboniferous period.
Vertebrates descent began during the Orovician period with the jawless fishes.
The Mesozoic Era
Carboniferous Period 359-299 mya Permian Period 299-251 mya Triassic Period 251-199 mya Jurassic Period 199-145 mya Cretaceous 145-65 mya
The Mesozoic Era
Reptiles can be traced back to The permian period.
The average size of dinosaurs were about the size of a crow, although there were a few giant species.
At the end of the Cretaceous period, dinosaurs became extinct.