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Module Name Introduction to Biological Oceanography Module Number MNF-bioc-101 Person in Charge Prof. Dr. Johannes Imhoff Phone: +49-(0)431-600-4450, E-mail: [email protected] Semester / Duration 1. semester / one semester Regular Cycle annual in winter semester Status Compulsory Study Programme Master of Science in Biological Oceanography Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size Introduction to Biological Oceanography (Lecture) Prof. Dr. Martin Wahl Prof. Dr. Julie LaRoche Prof. Dr. Ulrich Sommer Prof. Dr. Ulf Riebesell Prof. Dr. Johannes Imhoff 3 hrs per week / 40 students Credit Points / Workload 6 ECTS / 180 hours Prerequisites A bachelor’s degree in a biological discipline. Completion Module MNF-bioc-102 Following Module MNF-bioc-201 Educational Objectives On completion of this module students should be able to discuss and link key concepts in biological oceanography and fish ecology. They should have an understanding of the importance of functional groups of organisms both in shaping the food web including nekton as well as in elemental fluxes. Students should have enough knowledge to able to read and critically judge current literature on the topics covered. Content of Teaching This module will provide a broad overview of the functioning of marine ecosystems and the interactions between organismal groups that determine the cycling of bio-reactive elements in the ocean. Topics to be covered include: Physicochemical conditions in the ocean : large and small scale heterogeneity. Functional groups : micro-organisms, phytoplankton, zooplankton, benthos animals, algae, fishes, sea birds, mammals. Ecophysiology : light and photosynthesis, physiology of picoplankton, primary production, nutrients, microbial loop. Populations and communities : distribution, growth, age structure and demography, interactions, food webs. Biogeochemical cycles : classification of elements and their residence times, sources and sinks of elements, linking c to N, Si, P and Fe, microbiology of C-, N- and S-cycle. Diversity : patterns, significance and loss. Global Change : ocean acidification, global warming and “The Future Ocean”. Examination A written and graded examination will cover all topics of this module. Literature Sommer, U. 2005: Biologische Meereskunde; 2. Auflage, Springer Berlin. Lalli, C.M. & Parsons, T.R. 1993. Biological Oceanography: An Introduction. Open University, Pergamon Press. Additional current literature and lecture notes will be distributed during the semester. Additional Information None.

Module Name Introduction to Biological Oceanography · Additional Information The final note will combine the lecture and exercise exams. Module Name Introduction to Marine Geology

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Module Name Introduction to Biological Oceanography Module Number MNF-bioc-101

Person in Charge Prof. Dr. Johannes Imhoff Phone: +49-(0)431-600-4450, E-mail: [email protected]

Semester / Duration 1. semester / one semester

Regular Cycle annual in winter semester

Status

Compulsory

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Introduction to Biological Oceanography (Lecture)

Prof. Dr. Martin Wahl Prof. Dr. Julie LaRoche Prof. Dr. Ulrich Sommer Prof. Dr. Ulf Riebesell Prof. Dr. Johannes Imhoff

3 hrs per week / 40 students

Credit Points / Workload 6 ECTS / 180 hours

Prerequisites A bachelor’s degree in a biological discipline.

Completion Module MNF-bioc-102

Following Module MNF-bioc-201

Educational Objectives

On completion of this module students should be able to discuss and link key concepts in biological oceanography and fish ecology. They should have an understanding of the importance of functional groups of organisms both in shaping the food web including nekton as well as in elemental fluxes. Students should have enough knowledge to able to read and critically judge current literature on the topics covered.

Content of Teaching

This module will provide a broad overview of the functioning of marine ecosystems and the interactions between organismal groups that determine the cycling of bio-reactive elements in the ocean. Topics to be covered include: Physicochemical conditions in the ocean: large and small scale heterogeneity. Functional groups: micro-organisms, phytoplankton, zooplankton, benthos animals, algae, fishes, sea birds, mammals. Ecophysiology: light and photosynthesis, physiology of picoplankton, primary production, nutrients, microbial loop. Populations and communities: distribution, growth, age structure and demography, interactions, food webs. Biogeochemical cycles: classification of elements and their residence times, sources and sinks of elements, linking c to N, Si, P and Fe, microbiology of C-, N- and S-cycle. Diversity: patterns, significance and loss. Global Change: ocean acidification, global warming and “The Future Ocean”.

Examination A written and graded examination will cover all topics of this module.

Literature

Sommer, U. 2005: Biologische Meereskunde; 2. Auflage, Springer Berlin. Lalli, C.M. & Parsons, T.R. 1993. Biological Oceanography: An Introduction. Open University, Pergamon Press. Additional current literature and lecture notes will be distributed during the semester.

Additional Information None.

Module Name Practical Course in Biological Oceanography Module Number MNF-bioc-102

Person in Charge Dr. Uwe Piatkowski Phone: +49-(0)431-600-4571, E-mail: [email protected]

Semester / Duration 1. semester / one semester

Regular Cycle annual in winter semester

Status

Compulsory

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Practical Course in Biological Oceanography (Practical)

Prof. Dr. Ulrich Sommer Prof. Dr. Martin Wahl Prof. Dr. Andreas Oschlies Prof. Dr. Peter Reijnders PD Dr. Avan Antia Prof. Dr. Thorsten Reusch

Exercises in Biological Oceanography (Exercise)

12 hrs per week / 30 students

1 hr per week / 30 students

Credit Points / Workload 10 ECTS / 300 hours

Prerequisites A bachelor’s degree in a biological discipline.

Completion Module MNF-bioc-101

Following Module MNF-bioc-201

Educational Objectives

Students should leave this module with skills that enable them to make measurements of key variables and parameters of marine ecosystems as well as be able to interpret the results they obtain. They will gain experience in taking samples on board a research ship and using standard oceanographic equipment.

Content of Teaching

This module consists largely of practical courses introducing the main methods used in marine biological research. Students will work in small groups on experimental and methodological aspects of research as well as gain experience in field research on board a ship. Topics to be covered are: Protists Benthos Zooplankton Top predators Biogeochemistry Fish biology

Examination Written protocols or oral presentations (pass / fail) are required and a written examination (100%) will be used to grade participants of this module.

Literature Detailed methodological instructions and accompanying notes will be distributed during the practical course.

Additional Information The Practical will be conducted in two groups of 15 students each.

Module Name Doing Science Module Number MNF-bioc-110

Person in Charge Prof. Dr. Martin Wahl Phone: +49-(0)431-600-4577, E-mail: [email protected]

Semester / Duration 1. semester / one semester

Regular Cycle annual in winter semester

Status

Compulsory

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Key Scientific Skills (Lecture)

Prof. Dr. Douglas Wallace PD Dr. Avan Antia Prof. Dr. Martin Wahl Dr. Mark Lenz Prof. Dr. Arne Körtzinger Dr. habil. Iris Werner

Key Scientific Skills (Exercise)

1 hr per week / 35 students 2 hrs per week / 35 students

Credit Points / Workload 6 ECTS / 180 hours

Prerequisites A bachelor’s degree and participation in a master study programme in a scientific discipline.

Completion Module None.

Following Module The skills taught in this module will be drawn upon in the modules MNF-bioc-210 and MNF-bioc-302.

Educational Objectives This module will equip students to pursue independent scientific research, be able to critically read and judge scientific literature and communicate their results lucidly in oral and verbal form.

Content of Teaching

This module aims at teaching and practicing specific skills necessary for participating in scientific research. Topics to be covered are: Recognition of scientifically interesting questions, formulation of hypotheses, gathering data and designing experiments. Analysis of results and graphic presentation. Written and Oral Scientific Communication. Critical review of scientific literature. Writing a proposal.

Examination The grade will comprise a written exam (75%) and a protocol or oral presentation (25%).

Literature Doing Science - Design, Analysis and Communication of Scientific Research von Ivan Valiela, Oxford, University Press, 2001. Quinn, G.P. and Keough, M.J. Experimental design and data analysis for biologists, Cambridge University Press.

Additional Information This module will consist of mostly interactive modes of learning including periodic home assignments, giving short talks and refereeing individual topics.

Module Name Introduction to Chemical Oceanography Module Number MNF-choc-101

Person in Charge Prof. Dr. Douglas Wallace Phone: +49-(0)431-600-4200, E-mail: [email protected]

Semester / Duration 1. semester / one semester

Regular Cycle annual in winter semester

Status

Compulsory

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Chemical Oceanography (Lecture)

Prof. Dr. Douglas Wallace

Chemical Oceanography (Exercise)

1 hr per week / 50 students 2

hrs per week / 50 students

Credit Points / Workload 5 ECTS / 150 hours

Prerequisites A bachelor’s degree or equivalent in a scientific discipline. Basic knowledge of chemistry.

Completion Module None.

Following Module None.

Educational Objectives Students will gain demonstrable abilities to evaluate the role of ocean chemistry in major elemental cycles and be able to use these to understand interdisciplinary concepts and principles associated with them.

Content of Teaching

Topics to be covered are: Basic concepts and principles in marine chemistry. Major elemental cycles. Chemical interactions (river-ocean, sediment-ocean, atmosphere-ocean). Air-sea gas exchange.

Examination Periodic exercises (graded; 40% of final mark) with pass as prerequisite for graded written exam (60% of final mark)..

Literature Recommendations for textbooks and relevant literature will be made during the course.

Additional Information The final note will combine the lecture and exercise exams.

Module Name Introduction to Marine Geology Module Number MNF-geol-101

Person in Charge Prof. Dr. Ralph Schneider Phone: +49-(0)431-880-1457, E-mail: [email protected]

Semester / Duration 1. semester / one semester

Regular Cycle annual in winter semester

Status

Compulsory

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Introduction to Marine Geology (Lecture)

Prof. Dr. Jan Behrmann Prof. Dr. Dirk Nürnberg Prof. Dr. Klaus Wallmann Dr. Christian Hensen

2 hrs per week / 35 students

Credit Points / Workload 3 ECTS / 90 hours

Prerequisites A bachelor’s degree or similar level.

Completion Module None.

Following Module None.

Educational Objectives The objective of this course is to provide the basic knowledge on Marine Geology. The course focuses on all aspects of Marine Geology and is the basis for all more advanced courses in biological oceanography.

Content of Teaching

The topics of the lectures include: General introduction to marine geology, Geophysics and ocean morphology, Oceanic sediments and microfossils in relationship to the modern ocean system, Degradation of organic matter, Dissolution and precipitation of carbonate minerals, Evolution of marine biogeochemical cycles, Geological resources, Reading global change within oceanic sediments: Quantifying paleo-envrionmental change by using proxies, Gimme Age!!! Age control, stratigraphy, Approaches to Paleoceanography.

The topics of the seminars/laboratory exercises include: Long-term ocean floor observatories, Modeling early diagenesis, Modeling marine biogeochemical cycles, Ocean bathymetry, Biogeochemical fluxes across the sediment/water interface, Modern methods in paleoclimatic research: From sediment core studies to the understanding of our climate (part 1), Modern methods in paleoclimatic research: From sediment core studies to the understanding of our climate (part 2), Field course: Glacial and postglacial geology of NW Europe, Field course: Foraminifera in German tidal flats, Field course: Ancient pelagic sediments, Introduction to mass spectrometry, Geo-archaeology, Carbonate mounds, corals: Warm and cool water carbonates, sclerospongies, Venting, cold seeps, fluid flow, Coastal geology, Particle transport dynamics.

Examination Written examination (100%). Internal examiner. Relevant tables and calculator are allowed as aids.

Literature

Robert. A. Berner (1980), Early Diagenesis: A Theoretical Approach, Princeton University Press. Bernard P. Boudreau (1997), Diagenetic Models and Their Implementation. Modelling Transport and Reactions in Aquatic Sediments, Spinger-Verlag.

Additional Information A field course entitled ‘Glacial and postglacial geology of NW Europe’ accompanies this module.

Module Name Introduction to Physical Oceanography Module Number MNF-ozgr-151

Person in Charge Prof. Dr. Martin Visbeck Phone: +49-(0)431-600-4100, E-mail: [email protected]

Semester / Duration 2. semester / one semester

Regular Cycle annual in summer semester

Status

Compulsory

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Introduction to Physical Oceanography (Lecture)

Prof. Dr. Martin Visbeck Prof. Dr. Claus Böning

3 hrs per week / 80 students

Credit Points / Workload 5 ECTS / 150 hours

Prerequisites Basic knowledge in physics.

Completion Module None.

Following Module None.

Educational Objectives Understanding of the basic concepts used in Physical Oceanography, including a description of the large scale Ocean circulation and water mass properties.

Content Of Teaching Topography of the oceans, physical properties of sea water and sea-ice, water mass properties, thermal and haline startification, wind driven currents, geostrophic currents, thermo-haline circulation, regional oceanography, waves and tides.

Examination Written exam (100%).

Literature

Pickard, G.L. and Emery, W.J. 1990. Descriptive Physical Oceanography - An Introduction. Pergamon Press, 5th edition, 320 pp. Bearman, G. (Ed.) 1989. Waves, tides and shallow-water processes. Pergamon Press, Oxford (Open Univ.), reprinted with corrections 1991, 1995, 1997, 187 pp. Bearman, G. (Ed.) 1989. Ocean circulation. Pergamon Press, Oxford (Open Univ.), reprinted with corrections 1998, 238 pp. Bearman, G. (Ed.) 1998. The ocean basins: their structure and evolution. Pergamon Press, Oxford (Open Univ.), 2nd edition, 185 pp. Tomczak, M. and Godfrey, J.S. 1994. Regional Oceanography: An Introduction. Pergamon Press, 422 pp. Pond, S., and Pickard, G.L. 1983. Introductory Dynamical Oceanography, Butterworth-Heinemann, reprinted with corrections 1986, 1989, 329 pp. Gill, A.E. 1982. Atmosphere – Ocean Dynamics. International Geophysics Series, Vol. 30, Academic Press, 662 pp. Peixoto, J.P. and Oort, A.H. 1992. Physics of Climate. Springer-Verlag New York, Inc., 520 pp. Wang, C., Xie, S.-P., Carton, J. A. (eds) 2004. Earth’s Climate: The Ocean – Atmosphere Interaction. Geophysical Monograph 147, American Geophysical Union, 405 pp.

Additional Information None.

Module Name Advanced Course in Biological Oceanography Module Number MNF-bioc-201

Person in Charge Prof. Dr. Ulrich Sommer Phone: +49-(0)431-600-4400, E-mail: [email protected]

Semester / Duration 2. semester / one semester

Regular Cycle annual in summer semester

Status

Compulsory

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Advanced Biological Oceanography (Lecture, all students) Part A: Marine Molecular and Chemical Ecology

Prof. Dr. Thorsten Reusch Prof. Dr. Martin Wahl Prof. Dr. Frank Melzner Dr. Christophe Eizaguirre Dr. Inken Kruse Dr. Florian Weinberger Part B: Plankton Ecology and Pelagic BiogeochemistryProf. Dr. Ulrich Sommer Prof. Dr. Ulf Riebesell Dr. Nicole Aberle-Malzahn Part C: Microbial Ecology, Microbial Interactions and Marine BiotechnologyProf. Dr. Johannes Imhoff Prof. Dr. apl. Julie LaRoche Dr. Antje Labes Dr. Jörg Süling __________________________

Part A: Marine Molecular and Chemical Ecology (Practical)

Part A: Marine Molecular and Chemical Ecology (Exercise / Tutorial) ___________________________________________________________

Part B: Plankton Ecology and Pelagic Biogeochemistry (Practical)

Part B: Plankton Ecology and Pelagic Biogeochemistry (Exercise / Tutorial) ____________________________________________________________

Part C: Microbial Ecology, Microbial Interactions and Marine Biotechnology (Practical)

Part C: Microbial Ecology, Microbial Interactions and Marine Biotechnology (Exercise / Tutorial)

3 hr per week / 30 students 3 hrs per week / 15 students

1 hr per week / 15 students

3 hrs per week / 15 students

1 hr per week / 15 students

3 hrs per week / 15 students

1 hr per week / 15 students

Credit Points / Workload 11 ECTS / 330 hours

Prerequisites MNF-bioc-101 and MNF-bioc-102 or equivalent modules.

Completion Module None.

Following Module None.

Educational Objectives

Part A: Marine Molecular and Chemical Ecology On completion of this course, the students should have an in-depth understanding of stress reactions, chemical interactions and evolutionary processes in marine ecosystems. They should be able to understand the design of experiments / sampling schemes, and the concept ofmoelcular markers. They will learn to conduct experiments, retrieve and evaluate data and put their findings into the context of the scientific literature on chemical ecology, population genetics and evolutionary biology.

Part B: Plankton Ecology and Pelagic Biogeochemistry On completion of this course, the students should have an in-depth understanding of plankton ecology and pelagic biogeochemistry. They should be able to understand the design of experiments and sampling schemes, conduct experiments, retrieve and evaluate data and put their findings into the context of the scientific literature on pelagic ecology and biogeochemistry.

Part C: Microbial Ecology, Microbial Interactions and Marine Biotechnology On completion of this course, the students should have an in-depth understanding of marine microbial ecology and its contribution to biogeochemical cycles. They should be able to understand the design of experiments and sampling schemes, conduct experiments, retrieve and evaluate data and put their findings into the context of the scientific literature on marine.

Content of Teaching

Part A: Marine Molecular and Chemical Ecology: Lectures will cover the following aspects: Stress ecology & transcription analysis a) Stress, interaction between stressors, biotic modulation of stress b) quantitative genetics of stress genes in fucoid algae and shore crabs c) gene expression responses of shore crabs to ocean acidification Chemical Ecology: Main principles and regulation of a) predator-prey- and host-pathogen interactions b) substrate recognition c) anti-fouling defences Population genetics & evolution: a) gene flow & dispersal estimates b) marine speciation, phylogeography & hybridisation c) selection & adaptation, rapid evolutionary processes, marine genomics d) practical: DNA sequence analysis of immune genes in coastal fish, microsatellite genotyping of an invasive jellyfish

We aim at dividing the entire group into smaller groups of 3-5- students who will work on one continual project in one of the lecture topics for 3 weeks. At the end of the course, time will be given to mutually exchange information among groups. Additionally, practical courses are aimed at teaching laboratory skills in addressing research questions on selected topics listed above A weekly tutorial will update students and lecturers on progress made and clarify open questions.

Part B: Plankton Ecology and Pelagic Biogeochemistry Lectures will cover the following aspects: a) seawater carbonate system b) carbon and nutrient cycling c) nutrient limitation d) competition e) grazing

Additionally, practical courses are aimed with a mesocosm experiment for an intergrated study of the lecture topics. A weekly tutorial will update students and lecturers on progress made and clear outstanding questions.

Part C: Microbial Ecology, Microbial Interactions and Marine Biotechnology Lectures will cover the following aspects: a) ecophysiology of microbial groups and their genetic analysis in the environment b) secondary metabolites of marine microorganisms c) marine biotechnological applications

Additionally, practical courses are aimed at teaching laboratory skills in addressing research questions on selected topics listed above A weekly tutorial will update students and lecturers on progress made and clarify open questions.

Examination Per part: WE or P or OP (30%) Lecture: written exam (40%).

Literature

Part A: Lalli C. M. & Parsons T. R. (1997). Biological Oceanography: An Introduction. Butterworth-Heinemann Oxford, 314 pp. Parsons T.R., Takahashi M. & Hargrave B. (1984). Biological Oceanographic Processes. Pergamon Press Oxford, 330 pp. Ocean Biogeochemistry, (2003), M.J.R. Fasham (ed.) Springer Verlag, Berlin. ISBN 3-540-42398-2

Part B: Lalli C. M. & Parsons T. R. (1997). Biological Oceanography: An Introduction. Butterworth-Heinemann Oxford Townsend, C.R., Harper, J.L. & Begon M.E. (2002) Essentials of Ecology. 2nd edition. Blackwell, Oxford – or any other standard textbook of ecology. R.T. Paine (1994) Marine Rocky Shores and Community Ecology: An Experimentalist’s Perspective. Ecology Institute, Oldendorf Further literature recommendations will be communicated in the course

Part C: Pitcher, T.J. and P.J.B. Hart. 1992. Fisheries Ecology. Chapman & Hall, London. 414 pp. Helfman, G.S, Bruce, B. and E.F. Douglas. 1997. The Diversity of Fishes. Blackwell Science. 528 pp. Nelson, J.S. 2006. Fishes of the World. Wiley.

Additional Information

This Module comprises of an overarching lecture for all students and 3 sets of practical courses with tutorials. Students must choose two of the three sets: Part A: Chemical Ecology and Fish Ecology and/or Part B: Plankton Ecology and Pelagic Biogeochemistry and/or Part C: Microbial Ecology, Microbial Interactions and Marine Biotechnology.

Module Name Biological Modelling and Biostatistics Module Number MNF-bioc-220

Person in Charge Prof. Dr. Andreas Oschlies Phone: +49-(0)431-600-1936, E-mail: [email protected]

Semester / Duration 2. semester / one semester

Regular Cycle annual in summer semester

Status

Compulsory

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Biological Modelling and Biostatistics (Lecture)

Prof. Dr. Andreas Oschlies

Biological Modelling and Biostatistics (Exercise)

Fundamentals of Biostatistics (Lecture)

Dr. Mark Lenz

Fundamentals of Biostatistics (Exercise)

1 hrs per week / 30 students

1 hr per week / 30 students

1 hrs per week / 30 students

1 hr per week / 30 students

Credit Points / Workload 6 ECTS / 180 hours

Prerequisites A bachelor’s degree and participation in a master study programme in a scientific discipline. Basic knowledge of MATLAB.

Completion Module None.

Following Module MNF-bioc-341

Educational Objectives

Modelling: The class shall - educate the students in quantitative environmental and Earth system science, - strengthen the students' quantitative and computational skills, - provide understanding of a variety of forward and inverse modelling approaches, and - develop an understanding of the creation and application of numerical models.

Biostatistics: The class is intended to impart introductory knowledge and basic concepts of biostatistics to biology students. To consolidate their theoretical knowledge, it will be taught together with applications using real and virtual data sets from marine ecology and the statistical software package STATISTICA 6.1 (learning-by-doing principle). The course will enable students to formulate scientific models and derive hypothesis from them which are testable in experimental or observational studies. They will learn about basic concepts in experimental design, data collection and processing, and the identification of the appropriate statistical method to analyse a given set of data. Lectures and exercises about the most common statistical procedures will supplement this module.

Content of Teaching

Modelling: The unit will be delivered through a combination of lectures and computer-based accompanying assignments. The coarse will teach how to use relatively simple mathematical methods to understand in a quantitative manner how planet Earth and its inhabitants interact. Modelling concepts will be presented and applied to simple aquatic ecosystems. Students will learn how to use computer-based interactive modelling platforms to build, apply and analyse simple numerical models.

Biostatistics: The theoretical part of the course is structured into following chapters: 1. Selected Literature 2. Patterns, Models, Hypotheses 3. Populations, Samples, Variables 4. Parameters of Location and Spread 5. Hypothesis Testing 6. Assumptions of Statistical Tests 7. Parametric and Non-parametric Tests for two Samples 8. One- and Multifactorial Analysis of Variance, Post-hoc Tests, Planned Comparisons 10. Correlations 11. Simple and Multiple Regression 12. Multivariate Statistics for Community Analysis: ANOSIM, MDS

The applied part of the course using STATISTICA provides insight into following topics: 1. Processing Data 2. Plotting Data 3. Basic Statistics 4. General Linear Models

Examination Graded protocol or homework

Literature Literature references will be provided in the individual lectures. The course is based on two manuscripts, one covers the theory, the other SAS applications; both will be issued as handouts to the students on a chapter-by-chapter basis.

Additional Information None.

Module Name Current Topics in Marine Biogeochemistry I Module Number MNF-bioc-231

Person in Charge Prof. Dr. Ulf Riebesell Phone: +49-(0)431-600-4444, E-mail: [email protected]

Semester / Duration 2. semester / one semester

Regular Cycle annual in summer semester

Status

Compulsory elective

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Current Topics in Biogeochemistry I (Lecture)

Prof. Dr. Ulf Riebesell Prof. Dr. Arne Körtzinger

Current Topics in Biogeochemistry I (Seminar)

1 hrs per week / 30 students

2 hrs per week / 30 students

Credit Points / Workload 4 ECTS / 120 hours

Prerequisites MNF-bioc-101, MNF-bioc-102, MNF-choc-101 Basic knowledge of elemental cycles, chemical and biological processes in the ocean.

Completion Module None.

Following Module None.

Educational Objectives

This module serves two purposes: In the lectures students will be taught the basics of marine biogeochemical cycling, with particular focus on the carbon, nitrogen and iron cycles. In the seminar series students will be exposed to current research topics, new developments and novel scientific concepts in the area of marine biogeochemistry. On successful completion of this module, students will be familiar with biogeochemical processes involving the ocean’s major bioactive elements. They will have a basic understanding of the physical, chemical, and biological processes driving fluxes between the different marine carbon pools, between land, ocean and atmosphere. Students will learn about the dynamics of the marine carbon, nitrogen and iron cycle on time scales from seconds (seawater carbonate system), to months (seasonal cycles), to years (inter-annual variability), to tens of thousands of years (glacial/interglacial periods) and will be able to identify feedback mechanisms in the climate system. They will be aware of the present and projected future impacts of human activities on marine ecosystems and biogeochemistry. The seminar series will provide students with the opportunity to, improve their competence to critically evaluate on-going research, participate in scientific discussions, select their own research questions, formulate testable hypotheses, and select state-of-the-art methodologies. By experiencing the interdisciplinary nature of marine sciences, students will develop the ability to place results in a specific area into the larger context of understanding the role of the ocean in the earth system.

Content Of Teaching

On-going research in marine biogeochemistry will be presented and critically discussed. Current topics will be from a broad area of marine sciences with relevance to marine biogeochemistry, including molecular biology and genetics, physiology, ecology, to marine chemistry, isotope geochemistry, atmospheric chemistry, to ecosystem and biogeochemical modelling.

Examination Written exam (100%).

Literature Relevant literature will be given out during the module.

Additional Information None.

Module Name Current Topics in Marine Ecology I Module Number MNF-bioc-232

Person in Charge Prof. Dr. Ulrich Sommer Phone: +49-(0)431-600-4400, E-mail: [email protected]

Semester / Duration 2. semester / one semester

Regular Cycle annual in summer semester

Status

Compulsory elective

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Current Topics in Marine Ecology I (Lecture)

N.N.

Current Topics in Marine Ecology I (Seminar)

Prof. Dr. Ulrich Sommer Prof. Dr. Martin Wahl

1 hrs per week / 30 students

2 hrs per week / 30 students

Credit Points / Workload 4 ECTS / 120 hours

Prerequisites MNF-bioc-101, MNF-bioc-102, MNF-choc-101 Basic knowledge of marine organisms and ecology.

Completion Module None.

Following Module MNF-bioc-332

Educational Objectives

The students will get knowledge and understanding of the current marine ecological research activities (pelagic and benthic) at IFM-GEOMAR and their scientific background. The goal of the lecture is to convey a basic understanding of and current research topics in ecological processes and patterns in selected marine pelagic ecosystems.

Content Of Teaching

In this module current activities marine plankton and benthos research will be presented and discussed in the context of the actual theoretical framework. The lecture will cover patterns and mechanisms of pelagic and benthic food web organisation from costal to off-shore and surface to deep ecosystems.

Examination Graded written exam (100%).

Literature Literature will be given during the module.

Additional Information The module will be given on a weekly basis.

Module Name Current Topics in Fish Ecology and Aquaculture Module Number MNF-bioc-233

Person in Charge Prof. Dr. Reinhold Hanel Phone: +49-(0)431-600-4556, E-mail: [email protected]

Semester / Duration 2. semester / two weeks

Regular Cycle annual in summer semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Fish Ecology (Excursion)

Prof. Dr. Reinhold Hanel

Fish Ecology (Exercise)

Faunistics and Ecology of the Mediterranean Sea (Seminar)

3 hrs per week / 8 students

3 hrs per week / 8 students

1 hr per week / 8 students

Credit Points / Workload 4 ECTS / 120 hours

Prerequisites None.

Completion Module None.

Following Module None.

Educational Objectives

On completion of this module, students will have gained a basic knowledge on biodiversity and ecology of the littoral zone of the Mediterranean Sea, with special focus on fish, macrophytes and macrozoobenthos. They will have an understanding of different functional groups of organisms, their interactions and the principal abiotic factors shaping their environment.

Content of Teaching

This module will give insight into the flora and fauna of the Mediterranean Sea, the role of environmental parameters and different methods to explore the littoral zone. Topics to be covered include: Taxonomy and ecology of key species of different marine littoral habitats. Ecology and behaviour of Mediterranean fish species. Mediterranean benthos: Taxonomy, habitats and life forms.

Examination A graded protocol (100%) will cover all topics in this module.

Literature

Among others: Hofrichter, R. 2002. Das Mittelmeer – Fauna, Flora, Ökologie. Teile 1-3. Spektrum Verlag. Riedl, R. 1984. Fauna und Flora des Mittelmeers. Verlag Paul Parey, Hamburg – Berlin.

Additional Information This module will take place as a 2-weeks block course in Calvi, Corsica.

Module Name Element cycles in the ocean - Stoffkreisläufe im Meer (500040)

Module Number MNF-bioc-250

Person in Charge PD Dr. Hermann W. Bange Phone: +49-(0)431-600-4204, E-mail: [email protected], Homepage: www.ifm-geomar.de/

Semester / Duration 2. semester / one semester

Regular Cycle annual, in summer semester

Status

optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Marine biogeochemical cycles (Lecture)

PD Dr. Hermann W. Bange

2 hr per week / 20 students

Credit Points / Workload 3 ECTS / 90 hours

Prerequisites Basic knowledge in chemistry, physics, and biology

Completion Module None.

Following Module None.

Educational Objectives The goal of this lecture is to gain a deeper understanding of the marine biogeochemical cycles in the water column and their interactions with the atmosphere.

Content of Teaching

(i) Evolution of biogeochemical cycles (ii) Basic principles and concepts (iii) Nitrogen cycle (incl. N2 fixation, nitrification, denitrification,

anammox) (iv) Phosphorus cycle (v) Sulphur cycle (vi) Silicon cycle (vii) Trace metal cycles (focus on iron) (viii) Coupling of biogeochemical cycles

Examination A graded oral exam.

Literature

1) “Earth System Science – From biogeochemical cycles to global change” ed. MC Jacobson et al., Academic Press, 2000.

2) “Biogeochemistry – An analysis of global change”,2. Auflage, WH Schlesinger, Academic Press, 1997.

3) “Ocean Biogeochemical Dynamics”, JL Sarmiento andN Gruber, Princeton University Press, 2006.

4) “Introduction to Marine Biogeochemistry”, 2nd edition,SM Libes, Academic Press, 2009

Additional Information

This lecture is interdisciplinary. Students interested in chemical oceanography, biological oceanography,marine microbiology and Earth system science are welcome. The lecture will be given regularly every week. Please check UnivIS for exact dates.

Module Name Stoffkreisläufe im Meer (500040) Module Number MNF-bioc-250

Person in Charge PD Dr. Hermann W. Bange Phone: +49-(0)431-600-4204, E-mail: [email protected], Homepage: www.ifm-geomar.de/

Semester / Duration 2. semester / one semester

Regular Cycle annual, in summer semester

Status

optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Marine biogeochemical cycles (Lecture)

PD Dr. Hermann W. Bange

2 hr per week / 20 students

Credit Points / Workload 3 ECTS / 90 hours

Prerequisites Basic knowledge in chemistry, physics, and biology

Completion Module None.

Following Module None.

Educational Objectives The goal of this lecture is to gain a deeper understanding of the marine biogeochemical cycles in the water column and their interactions with the atmosphere.

Content of Teaching

(i) Evolution of biogeochemical cycles (ii) Basic principles and concepts (iii) Nitrogen cycle (incl. N2 fixation, nitrification, denitrification,

anammox) (iv) Phosphorus cycle (v) Sulphur cycle (vi) Silicon cycle (vii) Trace metal cycles (focus on iron) (viii) Coupling of biogeochemical cycles

Examination A graded oral exam.

Literature

1) “Earth System Science – From biogeochemical cycles to global change” ed. MC Jacobson et al., Academic Press, 2000.

2) “Biogeochemistry – An analysis of global change”,2. Auflage, WH Schlesinger, Academic Press, 1997.

3) “Ocean Biogeochemical Dynamics”, JL Sarmiento andN Gruber, Princeton University Press, 2006.

4) “Introduction to Marine Biogeochemistry”, 2nd edition,SM Libes, Academic Press, 2009

Additional Information

This lecture is interdisciplinary. Students interested in chemical oceanography, biological oceanography,marine microbiology and Earth system science are welcome. The lecture will be given regularly every week. Please check UnivIS for exact dates.

Module Name Biogeochemistry of Marine Sediments I Module Number MNF-bioc-251

Person in Charge Prof. Dr. Tina Treude Phone: +49-(0)431-600-2837, E-mail: [email protected],

Semester / Duration 2. semester / one semester

Regular Cycle annual, in summer semester

Status

optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Biogeochemistry of Marine Sediments I (Lecture)

Prof. Dr. Tina Treude

2 hrs per week / 30 students

Credit Points / Workload 4 ECTS / 120 hours

Prerequisites A bachelor’s degree in a biological, geological or chemical discipline.

Completion Module None.

Following Module MNF-bioc-356.

Educational Objectives

In this module students will learn basic microbial reactions that are connected to biodegradation, element cycling, and synthesis in marine sediments. After completion of the module students should be familiar with the principles of microbial redox reactions and able to interpret for example geochemical gradients that are shaped through biological activity. Furthermore the module offers an interdisciplinary basis to understand biogeochemical processes in benthic environments.

Content of Teaching

This module provides a comprehensive overview of microbial processes at the interface between the marine biosphere and geosphere. Topics will include redox reactions in oxygenated and reduced sediments (e.g. nitrate reduction, sulfate reduction, methanogenesis, and methane oxidation), element cycling, microbial photosynthesis, chemosynthesis, nitrogen fixation, fermentation, and isotope fractionation. The subject will be taught with examples from a variety of marine settings reaching from coastal to deep-sea as well as from polar to tropical environments.

Examination Manuscript 100%

Literature Canfield et al.: “Advances in Marine Biology: Aquatic Geomicrobiology”, Elsevier, ISBN: 0-12-026147-2; Konhauer: “Introduction to Geomicrobiology”, Blackwell, ISBN: 0-632-05454-9; Schulz et al.: “Marine Geochemistry”, Springer, ISBN: 3-540-66453-X

Additional Information This lecture is interdisciplinary and addresses students from the fields of biological oceanography, geochemistry, and microbiology. The lecture will be given regularly every week.

Module Name Marine Animal Physiology I Module Number MNF-bioc-252

Person in Charge Prof. Dr. Frank Melzner Phone: +49-(0)431-600-4274, E-mail: [email protected]

Semester / Duration 2. semester / one semester

Regular Cycle annual in summer semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Marine Animal Physiology I (Lecture)

Prof. Dr. Frank Melzner

1 hr per week / 40 students

Credit Points / Workload 2 ECTS / 60 hours

Prerequisites A bachelor’s degree in a biological discipline.

Completion Module MNF-bioc-355

Following Module None.

Educational Objectives The goal of this module is to introduce the students to the physiological diversity of marine invertebrate and vertebrate metazoans. Special attention will be given to physiological challenges associated with the specific abiotic environment of the Baltic Sea.

Content Of Teaching

This module will provide a broad overview on general principles in marine animal physiology. This includes: principles of cell biology and thermodynamics, metabolism & excretion, respiration and circulation, ion- and osmoregulation, response to key abiotic factors (oxygen, salinity, and temperature). In addition, special physiological adaptations to life in extreme habitats will be discussed (e.g. cold seeps, hydrothermal vents, mud flats, polar oceans, deep sea etc.)

Examination A written and graded examination (100%) will cover all topics in this module.

Literature Schmidt – Nielsen: Animal physiology: Adaptation and Environment, Cambridge University Press, 5th edition. Additional current literature and lecture notes will be distributed during the lecture.

Additional Information None.

Module Name How to Write and Publish a Scientific Paper Module Number MNF-bioc-253

Person in Charge Prof. Dr. Tina Treude Phone: +49-(0)431-600-2837, E-mail: [email protected]

Semester / Duration 2. semester / one semester

Regular Cycle

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

How to write and publish a scientific paper (Seminar)

Prof. Dr. Tina Treude

1 hr per week / 25 students

Credit Points / Workload 2 ECTS / 60 hours

Prerequisites A bachelor’s degree in a natural science discipline.

Completion Module None.

Following Module None.

Educational Objectives

In this module students will be introduced into the process of manuscript publishing in peer-reviewed scientific journals. The goal of the module is to provide insights into the fun and frustration of paper writing, important rules on manuscript structuring and scientific language as well as how to deal with the reviewing process. After completion of the module students should be familiar with the general principles of a successful publishing process.

Content of Teaching

This module will address different stages of manuscript writing and publishing answering basic questions such as: When are my data ready for publishing? Where should I publish? How do I structure the manuscript? How to present the data? What is my message? What are the Does and Don’ts of scientific writing? How to deal with the reviewers and editors? How to manage my coauthors? The module will be an interactive seminar.

Examination Graded homework 100%.

Literature Day: “How to write and publish a scientific paper”, Oryx Press, ISBN: 1-57356-165-7; Day: “Scientific English”, Oryx Press, ISBN: 0-89774-989-8; San Francisco Edit: http://www.sfedit.net/newsletters.htm

Additional Information

This module will be given in 2 hour-seminars every second week. For participation a subscription is necessary. Deadline is April 4th. First come, first serve. For subscription please contact Tina Treude via email. The module will start End of April. Please check the dates online or contact Tina Treude.

Module Name Mechanisms of biomineralization Module Number MNF-bioc-255

Person in Charge Prof. Dr. Frank Melzner Phone: +49-(0)431-600-4274, E-mail: [email protected]

Semester / Duration 2.-4. semester / one semester

Regular Cycle Summer and winter semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Mechanisms of biomineralization (Seminar)

Prof. Dr. Frank Melzner

2 hr per week / 40 students

Credit Points / Workload 3 ECTS / 90 hours

Prerequisites A bachelor’s degree in a biological discipline.

Completion Module MNF-bioc-355

Following Module None.

Educational Objectives The goal of this module is to introduce the students to the physiological diversity of marine invertebrate and vertebrate metazoans. Special attention will be given to physiological challenges associated with the specific abiotic environment of the Baltic Sea.

Content Of Teaching

This module will provide a broad overview on general principles in marine animal physiology. This includes: principles of cell biology and thermodynamics, metabolism & excretion, respiration and circulation, ion- and osmoregulation, response to key abiotic factors (oxygen, salinity, and temperature). In addition, special physiological adaptations to life in extreme habitats will be discussed (e.g. cold seeps, hydrothermal vents, mud flats, polar oceans, deep sea etc.)

Examination oral presentation (100%)

Literature Schmidt – Nielsen: Animal physiology: Adaptation and Environment, Cambridge University Press, 5th edition. Additional current literature and lecture notes will be distributed during the lecture.

Additional Information None.

Module Name Mechanisms of biomineralization Module Number MNF-bioc-255

Person in Charge Prof. Dr. Frank Melzner Phone: +49-(0)431-600-4274, E-mail: [email protected]

Semester / Duration 2.-4. semester / one semester

Regular Cycle Summer and winter semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Mechanisms of biomineralization (Seminar)

Prof. Dr. Frank Melzner

2 hr per week / 40 students

Credit Points / Workload 3 ECTS / 90 hours

Prerequisites A bachelor’s degree in a biological discipline.

Completion Module MNF-bioc-355

Following Module None.

Educational Objectives The goal of this module is to introduce the students to the physiological diversity of marine invertebrate and vertebrate metazoans. Special attention will be given to physiological challenges associated with the specific abiotic environment of the Baltic Sea.

Content Of Teaching

This module will provide a broad overview on general principles in marine animal physiology. This includes: principles of cell biology and thermodynamics, metabolism & excretion, respiration and circulation, ion- and osmoregulation, response to key abiotic factors (oxygen, salinity, and temperature). In addition, special physiological adaptations to life in extreme habitats will be discussed (e.g. cold seeps, hydrothermal vents, mud flats, polar oceans, deep sea etc.)

Examination oral presentation (100%)

Literature Schmidt – Nielsen: Animal physiology: Adaptation and Environment, Cambridge University Press, 5th edition. Additional current literature and lecture notes will be distributed during the lecture.

Additional Information None.

Module Name Trophodynamic Interactions Module Number MNF-bioc-262

Person in Charge Prof. Dr. Maarten Boersma Phone: +49-(0)4725-819-3350, E-mail: [email protected]

Semester / Duration 2. semester / ten days

Regular Cycle annual in summer semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Trophodynamic Interactions (Practical)

Prof. Dr. Maarten Boersma Prof. Dr. Karen Wiltshire Dr. Arne Malzahn Dr. Nicole Aberle-Malzahn

3 hrs per week / 20 students

Credit Points / Workload 5 ECTS / 150 hours

Prerequisites A bachelor’s degree in a biological discipline.

Completion Module None.

Following Module None.

Educational Objectives

This module aims at broadening the students’ view on food webs in marine systems and provides an overview of the key functional groups and trophic interactions in the plankton. The students will gain knowledge of the transfer of energy between trophic levels as well as stoichiometric constraints of this transfer. The students will learn how to develop and plan experiments. Finally, this course provides information on advanced food web concepts and, enables the students to discuss and link this knowledge to current research topics and relevant publications in the field of trophic ecology.

Content of Teaching

This module will provide an overview of key groups and species in the plankton of the North Sea (phytoplankton, microzooplankton, mesozooplankton) and of trophodynamic interactions between these organisms. By conducting microcosm grazing experiments in small groups simulating various abiotic conditions the students will gain knowledge of aspects i.e. energy transfer, stoichiometric needs and constrains (i.e. nutrient and light limitation light) and the transfer of limitation signals onto higher trophic levels.

Examination Graded homework or manuscript or oral presentation.

Literature

Sommer U. 2005: Biologische Meereskunde; 2. Auflage, Springer Berlin, Lalli, C.M. & T.R. Parsons. 1993. Biological Oceanography: an Introduction. Open University, Pergamon Press Additional current literature and lecture notes will be distributed during the semester.

Additional Information This module will take place as an 10-days block course from the 27.09.-06.10.2010 at the “Biologische Anstalt Helgoland” (Alfred Wegener Institute).

Module Name Sea Bird Ecology Module Number MNF-bioc-264

Person in Charge PD Dr. Stefan Garthe Phone: +49-(0)4834-604-116, E-mail: [email protected]

Semester / Duration 2. semester / seven days

Regular Cycle annual in summer semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Sea Bird Ecology (Practical)

PD Dr. Stefan Garthe 4 hrs per week / 12 students

Credit Points / Workload 4 ECTS / 120 hours

Prerequisites A bachelor’s degree in a biological discipline.

Completion Module None.

Following Module None.

Educational Objectives

On completion of this course students should have aquired an advanced knowledge of key aspects of the life history of seabirds. This includes an understanding of seabird behaviour, distribution, diet and habitat choice. Also, students should have understood the use of seabirds to indicate changes in the marine environment.

Content of Teaching

This course will have two main goals. One is to provide an overview of key aspects of the life history of seabirds, with practical elements studying seabird behaviour, seabird distrubution at sea, diet and feeding ecology as well as habitat choice. Secondly, the students will learn with which methods seabird ecology may be studied, including observations and experiments in the field and in the lab. Also, data collected during the course will be analysed and written up in a protocol.

Examination A protocol (in groups, pass / fail) and a written and graded examination (100%) which will cover all topics in this module.

Literature Literature and lecture notes will be distributed before the course.

Additional Information This module will take place as an 7-days block course around the end of May / beginning of June at the “Forschungs- und Technologie-Zentrum Westküste” in Büsum.

Module Name Zooplankton Ecology Module Number MNF-bioc-265

Person in Charge Prof. Dr. Sigrid Schiel Phone: 0471/4831-1303, E-mail: [email protected]

Semester / Duration 2. semester / 7 days

Regular Cycle annual in summer semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Zooplankton Ecology (Lecture, Practical & Seminar)

Prof. Dr. Sigrid Schiel

4 hr per week / 15 students

Credit Points / Workload 4 ECTS / 120 hours

Prerequisites A bachelor’s degree in a biological discipline.

Completion Module None.

Following Module None.

Educational Objectives This course will introduce students to the significance of present and anticipated changes in pelagic food web structure on energy transfer and carbon export.

Content of Teaching The students will acquire knowledge of standard methodologies in zooplankton feeding ecology. They will perform, analyse and present results of laboratory experiments and discuss current research topics.

Examination An oral presentation at the end of the course and a written protocol provided after completion of the course (pass / fail).

Literature

Frost B (1972) Effects of size and concentration of food particles on the feeding behavior of the marine planktonic copepod Calanus pacificus. L&O 17:805-815 Besiktepe S, Dam HG (2002) Coupling of ingestion and defecation as a function of diet in the calanoid copepod Acartia tonsa. MEPS 229:151-164 Further material will be distributed upon inscription.

Additional Information This course will take place as a block course in the lecture free time following the lecture period in the summer semester.

Module Name Advanced course in Polar Ecology Module Number MNF-bioc-266

Person in Charge Prof. Dr. Dieter Piepenburg Phone: +49-(0)431-600-1264, E-mail: [email protected]

Semester / Duration 2. / 3. semester / one semester

Regular Cycle annually in summer / winter semester

Status

optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Polar Ecology (Lecture)

Prof. Dr. Dieter Piepenburg Prof. Dr. Manfred Bölter PD Dr. Iris Werner

Polar Ecology (Seminar) Prof. Dr. Dieter Piepenburg Prof. Dr. Manfred Bölter PD Dr. Iris Werner

1 hrs per week / 15 students 2 hrs per week / 15 students

Credit Points / Workload 3 ECTS / 90 hours

Prerequisites A bachelor’s degree in a biological discipline and participation in a master study programme in a scientific discipline.

Completion Module None.

Following Module None.

Educational Objectives

The objective of this module is to introduce students to the biology and ecology of polar environments in different regions. After completion of the module, students should have a sound knowledge on diversity, habitats, life cycles, feeding ecology and adaptations in biology, physiology and behaviour of marine and terrestrial polar organisms.

Content of Teaching

The module provides in-depth information about dominant sympagic, pelagic and benthic organisms (including micro-organisms) and their specific environments in the polar regions of both hemispheres. The course will also cover topics of land-sea interactions, e.g. in estuaries and shelf ecosystems. Besides presenting the current knowledge on the general biology and ecology of these organisms, special topics such as threats, population status and conservation issues of habitats for individual species are also addressed with special respect to warming and subsequent men-made changes. The course is a combination of a series of lectures and a literature seminar with oral presentations of the students.

Examination The grade is based on a final written exam (100%) A “pass” for an oral presentation during the course constitutes the prerequisite for participating in the final exam.

Literature Hempel, G., Hempel, I. (eds) 2009: Biological studies in polar Oceans – exploration of life in icy waters. Wirtschaftsverlag NW, Verlag für neue Wissenschaft, Bremerhaven. Seminar papers will be distributed during the course.

Additional Information This module will also offer a one-day excursion, either to the Alfred-Wegener-Instutute of Polar und Marine Research in Bremerhaven (AWI), or the Hamburg Ice Tank Facilitiy (HSVA)

Module Name Identification and taxonomy of marine invertebrates

Module Number MNF-bioc-267

Person in Charge Prof. Dr. Martin Wahl Phone: +49-(0)431-600-4500, E-mail: [email protected]

Semester / Duration 2nd semester / one semester

Regular Cycle annual in summer semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Marine Phyla and Communities (Lecture)

various lecturers Identification of marine animal taxa (Practical) R. Schuett & Dr. H. Rumohr

1 hr per week / 20 students 3 hr per week / 20 students

Credit Points / Workload 5 ECTS / 150 hours

Prerequisites MNF-bioc-101 and 102

Completion Module

Following Module None.

Educational Objectives The objective of this course is to enable students to identify marine animals

Content Of Teaching

This course will provide knowledge of the morphology of marine animals from the Baltic Sea and develop the capacity of correct and scientific identification of marine invertebrate and some selected vertebrate taxa. A further important aspect is which life histories are possible and are realized based on a given bauplan.

Examination Graded homework or manuscript 100%

Literature None.

Additional Information A major portion of the identification keys is in German since no English translations are available yet.

Module Name Multidisciplinary Oceanographic Research Module Number MNF-bioc-301

Person in Charge Prof. Dr. Ulf Riebesell Phone: +49-(0)431-600-4581, E-mail: [email protected]

Semester / Duration 3. semester / one semester

Regular Cycle annual in winter semester

Status

Compulsory

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Advanced Laboratory Course (Practical)

PD Dr. Avan Antia Prof. Dr. Ulrich Sommer Prof. Dr. Martin Wahl Prof. Dr. apl. Julie LaRoche Prof. Dr. Reinhold Hanel Prof. Dr. Ulf Riebesell

IFM-GEOMAR Colloquium (Colloquium)

Prof. Dr. Arne Körtzinger

Lecture Series “The Future Ocean” (Colloquium)

Prof. Dr. Klaus Wallmann

Research Cruise or Field Course (Practicals)

Prof. Dr. S. Schiel Dr. Heye Rumohr

Tutorial & Thesis Proposal (Exercise)

Prof. Dr. Ulrich Sommer Prof. Dr. Martin Wahl Prof. Dr. Ulf Riebesell Prof. Dr. apl. Julie LaRoche Prof. Dr. Reinhold Hanel

2 hrs per week / 30 students

2 hrs per week / 50 students

1 hrs per week / 50 students

9 days / 10 students 2 hrs per week / 30 students

1 hr per week / 30 students

Credit Points / Workload 10 ECTS / 300 hours

Prerequisites All the compulsory modules of the first and second semesters of MNF-bioc-… or equivalent course content.

Completion Module None.

Following Module None.

Educational Objectives

Students will gain an insight into the ideas that drive major multidisciplinary research projects. They should be able to link applied and fundamental research especially at the interfaces of economic and legal aspects of maritime issues. This module aims at encouraging students to think laterally between scientific disciplines.

Content Of Teaching

This Module will offer an overview into multidisciplinary oceanographic research from a wide range of topics. These will cover all aspects of oceanography as well as economic and legal aspects of maritime research as part of the lecture series centred in the Research Network "The Future Ocean" of the Christian Albrecht’s University at Kiel. A weekly interdisciplinary colloquium at the IFM-GEOMAR will form the second half of the lecture series. These cover topics of interest to all disciplines covers at the IFM-GEOMAR and are held by internal as well as invited speakers.

Examination A written thesis proposal. (pass / fail)

Literature Selected literature will be recommended during the Colloquia series.

Additional Information The final mark will consist of a weighted mean of the individual course modules.

Module Name Summer School or Internship Module Number MNF-bioc-310

Person in Charge Prof. Dr. Tina Treude Phone: +49-(0)431-600-2837, E-mail: [email protected]: www.ifm-geomar.de/~ttreude

Semester / Duration 3. semester / one semester

Regular Cycle annual in winter semester

Status

Compulsory

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Summer School (Practical)

N.N.

Internship (Project)

N.N.

- / 30 students

- / 30 students

Credit Points / Workload 5 ECTS / 150 hours

Prerequisites Successful completion of the compulsory requirements of the master courses in Biological Oceanography of the first and second semester.

Completion Module None.

Following Module None.

Educational Objectives

On completion of this module students will have a more firm basis for career choice either in fundamental science or applied oceanography in a variety of settings. Students will also have the opportunity to demonstrate skills in communication, application of theoretical knowledge and lateral thinking in a practical setting.

Content Of Teaching Students will choose between participation in an intensive 2-week summer school or an equivalent to 150 hours internship at a maritime company or office. This can be fulfilled in agreement with the company or office as part time practical.

Examination Summer School: protocol (100%), Internship: protocol (100%).

Literature Relevant literature will be distributed within the course.

Additional Information A ship cruise might be part of the Summer School.

Module Name Current Topics in Marine Biogeochemistry II Module Number MNF-bioc-331

Person in Charge Prof. Dr. Ulf Riebesell Phone: +49-(0)431-600-4581, E-mail: [email protected]

Semester / Duration 3. semester / one semester

Regular Cycle annual in winter semester

Status

Compulsory elective

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Current Topics in Marine Biogeochemistry II (Lecture)

Prof. Dr. Ulf Riebesell Prof. Dr. Arne Körtzinger

Current Topics in Marine Biogeochemistry II (Exercise)

Prof. Dr. Ulf Riebesell Prof. Dr. Arne Körtzinger

Current Topics in Marine Biogeochemistry II (Seminar)

Prof. Dr. Ulf Riebesell Prof. Dr. Arne Körtzinger

1 hrs per week / 30 students

2 hr per week / 20 students

2 hrs per week / 25 students

Credit Points / Workload 5 ECTS / 150 hours

Prerequisites MNF-bioc-101, MNF-bioc-102, MNF-choc-101. Basic knowledge of elemental cycles, and chemical and biological processes in the ocean.

Completion Module MNF-bioc-231

Following Module None.

Educational Objectives

Students will obtain a broad overview of current research topics, new methodologies, novel scientific concepts, and latest developments in numerical modelling in the area of marine biogeochemistry. Upon successful completion of this module, students will have improved their competence to critically evaluate on-going research, participate in scientific discussions, select their own research questions, formulate testable hypotheses, and select state-of-the-art methodologies. By experiencing the interdisciplinary nature of marine sciences, students will develop the ability to place results in a specific area into the larger context of understanding the role of the ocean in the earth system.

Content Of Teaching

On-going research in marine biogeochemistry will be presented and critically discussed. Current topics will be from a broad area of marine sciences with relevance to marine biogeochemistry, including molecular biology and genetics, physiology, ecology, to marine chemistry, isotope geochemistry, atmospheric chemistry, to ecosystem and biogeochemical modelling.

Examination Graded oral presentation (100% of final mark).

Literature Relevant literature will be given out during the module.

Additional Information None.

Module Name Current Topics in Marine Ecology II Module Number MNF-bioc-332

Person in Charge Prof. Dr. Martin Wahl Phone: +49-(0)431-600-4577, E-mail: [email protected]

Semester / Duration 3. semester / one semester

Regular Cycle annual in winter semester

Status

Compulsory elective

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Current Topics in Marine Ecology II (Lecture)

Prof. Dr. Martin Wahl Prof. Dr. Ulrich Sommer

Current Topics in Marine Ecology II (Exercise)

Prof. Dr. Martin Wahl Prof. Dr. Ulrich Sommer

Current Topics in Marine Ecology II (Seminar)

Prof. Dr. Martin Wahl Prof. Dr. Ulrich Sommer

1 hrs per week / 30 students

2 hrs per week / 30 students

2 hr per week / 30 students

Credit Points / Workload 5 ECTS / 150 hours

Prerequisites MNF-bioc-101, MNF-bioc-102 Basic knowledge of marine organisms and ecology.

Completion Module MNF-bioc-351

Following Module None.

Educational Objectives The students will get knowledge and understanding of the current marine ecological research activities at IFM-GEOMAR and their scientific background and an insight into currently important questions, approaches, models and theories in benthic ecology.

Content Of Teaching In this module current activities marine plankton and benthos research will be presented and discussed in the context of the actual theoretical framework. The latter will be the content of the lecture part.

Examination Graded oral presentation (100% of final mark).

Literature None.

Additional Information None.

Module Name Current Topics in Fish Ecology Module Number MNF-bioc-334

Person in Charge Dr. Catriona Clemmesen, Prof. Dr. Thorsten Reusch Phone: +49-(0)431-600-4558, E-mail: [email protected] Phone: +49-(0)431-600-4550, E-mail: [email protected]

Semester / Duration 3. semester / one semester

Regular Cycle annual in winter semester

Status

Compulsory elective

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Fish Systematic, Biology and Evolution (Lecture)

Prof. Dr. Reinhold Hanel

Biodiversity of Fish (Lecture)

Dr. Rainer Froese

Current Topics in Fish Ecology (Seminar)

Dr. Catriona Clemmesen Prof. Dr. Thorsten Reusch Dr. Uwe Piatkowski

1 hrs per week / 25 students

2 hrs per week / 25 students

2 hr per week / 25 students

Credit Points / Workload 5 ECTS / 150 hours

Prerequisites

Compulsory Modules of the first and second semester of the master programme in Biological Oceanography or equivalent. Course content of MNF-bioc-201 Part A (Chemical ecology and fish ecology) are required. Firm knowledge of the basic concepts of fisheries ecology and aquaculture.

Completion Module None.

Following Module None.

Educational Objectives Students will be presented with information on current research topics in fish ecology, fisheries biology, evolutionary ecology and aquaculture.

Content Of Teaching

Basics in fish systematics, biology, biodiversity and evolution and on current topics in fish ecology. Students will learn to critically evaluate primary literature, extract information, present the content in a brief and concise way and will have to lead and structure scientific discussions.

Examination Graded oral presentation (50% of final mark) and written exam (50% of final mark).

Literature None. (will follow)

Additional Information

Module Name Advanced Biological Modelling Module Number MNF-bioc-341

Person in Charge Prof. Dr. Andreas Oschlies Phone: +49-(0)431-600-1936, E-mail: [email protected]

Semester / Duration 3. semester / one semester

Regular Cycle annual in winter semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Advanced Biological Modelling (Lecture)

Prof. Dr. Andreas Oschlies

Advanced Biological Modelling (Exercise)

2 hrs per week / 15 students

2 hrs per week / 15 students

Credit Points / Workload 5 ECTS / 150 hours

Prerequisites MNF-bioc-220 or equivalent. Basic knowledge of MATLAB.

Completion Module None.

Following Module None.

Educational Objectives

The class shall educate in different modelling approaches in environmental and Earth system science, strengthen the students' quantitative and computational skills, and the students are supposed to learn how to develop, set up, run, and analyse simple numerical models.

Content Of Teaching

The unit will be delivered through a combination of lectures and computer-based accompanying assignments. Students will use higher-level programming languages to manipulate numerical models provided by the organizers. We will discuss typical model errors and provide strategies for error minimization. At the end of the course, students will develop their own simple models to address a scientific problem of their choice.

Examination Graded protocol

Literature Literature references will be provided in the individual lectures.

Additional Information None.

Module Name Research Cruise / Field Course Module Number MNF-bioc-349

Person in Charge Dr. Heye Rumohr Phone: +49-(0)431-600-4524, E-mail: [email protected]

Semester / Duration 3. semester / 2 weeks

Regular Cycle annual prior start of winter semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Research Cruise / Field Course (Practical)

Dr. Heye Rumohr

2 weeks / 10 / 30 students

Credit Points / Workload 4 ECTS / 120 hours

Prerequisites All the compulsory modules of the first and second semesters of MNF-bioc-… or equivalent course content.

Completion Module None.

Following Module None.

Educational Objectives

Students will gain an insight into the ideas that drive major multidisciplinary research projects. They should be able to link applied and fundamental research especially at the interfaces of economic and legal aspects of maritime issues. This module aims at encouraging students to think laterally between scientific disciplines.

Content Of Teaching

This Module will offer an overview into multidisciplinary oceanographic research from a wide range of topics. These will cover all aspects of oceanography as well as economic and legal aspects of maritime research as part of the lecture series centred in the Research Network "The Future Ocean" of the Christian Albrecht’s University at Kiel. A weekly interdisciplinary colloquium at the IFM-GEOMAR will form the second half of the lecture series. These cover topics of interest to all disciplines covers at the IFM-GEOMAR and are held by internal as well as invited speakers.

Examination A research report (pass / fail)

Literature Selected literature will be recommended during the Tutorials.

Additional Information None.

Module Name Climate-relevant trace gases in the ocean - Klimarelevante Spurengase im Ozean (500053)

Module Number MNF-bioc-350

Person in Charge PD Dr. Hermann W. Bange Phone: +49-(0)431-600-4204, E-mail: [email protected], Homepage: www.ifm-geomar.de/

Semester / Duration 3. semester / one semester

Regular Cycle annual, in winter semester

Status

optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Climate relevant trace gases in the ocean (Lecture)

PD Dr. Hermann W. Bange

2 hr per week / 20 students

Credit Points / Workload 3 ECTS / 90 hours

Prerequisites Basic knowledge in chemistry, physics, and biology

Completion Module None.

Following Module None.

Educational Objectives

The goal of this lecture is to gain a deeper understanding of the distribution and biogeochemical pathways of climate relevant trace gases in the ocean. Additionally the students will gain insights into the mechanisms of air-sea gas exchange as well as the role of the ocean as source or sink of atmospheric trace gases and the implications for the atmosphere (greenhouse effect, ozone hole).

Content of Teaching

(i) significance of oceanic trace gases for greenhouse effect and ozone hole

(ii) chemical and physical properties of dissolved gases (iii) models of air-sea gas exchange (iv) methods to measure dissolved trace gase (incl. guided lab

tour) (v) marine biogeochemistry of selected trace gases (N2O, CH4,

DMS, COS, CO, halocarbons, H2, NH3, OVOCs)

Examination A graded oral exam.

Literature

1) “Earth System Science – From biogeochemical cycles to global change” ed. MC Jacobson et al., Academic Press, 2000.

2) “Biogeochemistry – An analysis of global change”,2. Auflage, WH Schlesinger, Academic Press, 1997.

3) „Chemie der Atmosphäre – Bedeutung für Klima und Umwelt“, TE Graedel and PJ Crutzen, Spektrum Akademischer Verlag, 1994.

Additional Information

This lecture is interdisciplinary. Students interested in chemical oceanography, biological oceanography,marine microbiology and Earth system science are welcome. The lecture will be given regularly every week. Please check UnivIS for exact dates.

Module Name Klimarelevante Spurengase im Ozean (500053) Module Number MNF-bioc-350

Person in Charge PD Dr. Hermann W. Bange Phone: +49-(0)431-600-4204, E-mail: [email protected], Homepage: www.ifm-geomar.de/

Semester / Duration 3. semester / one semester

Regular Cycle annual, in winter semester

Status

optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Climate relevant trace gases in the ocean (Lecture)

PD Dr. Hermann W. Bange

2 hr per week / 20 students

Credit Points / Workload 3 ECTS / 90 hours

Prerequisites Basic knowledge in chemistry, physics, and biology

Completion Module None.

Following Module None.

Educational Objectives

The goal of this lecture is to gain a deeper understanding of the distribution and biogeochemical pathways of climate relevant trace gases in the ocean. Additionally the students will gain insights into the mechanisms of air-sea gas exchange as well as the role of the ocean as source or sink of atmospheric trace gases and the implications for the atmosphere (greenhouse effect, ozone hole).

Content of Teaching

(i) significance of oceanic trace gases for greenhouse effect and ozone hole

(ii) chemical and physical properties of dissolved gases (iii) models of air-sea gas exchange (iv) methods to measure dissolved trace gase (incl. guided lab

tour) (v) marine biogeochemistry of selected trace gases (N2O, CH4,

DMS, COS, CO, halocarbons, H2, NH3, OVOCs)

Examination A graded oral exam.

Literature

1) “Earth System Science – From biogeochemical cycles to global change” ed. MC Jacobson et al., Academic Press, 2000.

2) “Biogeochemistry – An analysis of global change”,2. Auflage, WH Schlesinger, Academic Press, 1997.

3) „Chemie der Atmosphäre – Bedeutung für Klima und Umwelt“, TE Graedel and PJ Crutzen, Spektrum Akademischer Verlag, 1994.

Additional Information

This lecture is interdisciplinary. Students interested in chemical oceanography, biological oceanography,marine microbiology and Earth system science are welcome. The lecture will be given regularly every week. Please check UnivIS for exact dates.

Module Name Microbiology of Extreme Marine Habitats Module Number MNF-bioc-352

Person in Charge Prof. Dr. Johannes Imhoff Phone: +49-(0)431-600-4433, E-mail: [email protected]

Semester / Duration 3. semester / one semester

Regular Cycle annual in winter semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Microbiology of Extreme Marine Habitats (Lecture)

Prof. Dr. Johannes Imhoff

1 hr per week / 30 students

Credit Points / Workload 2 ECTS / 60 hours

Prerequisites A bachelor’s degree a biological discipline. Basics in marine microbiology.

Completion Module None.

Following Module GAME Programme and every research module of the master programme.

Educational Objectives Understanding the content of the lecture.

Content Of Teaching Presentation of extreme habitats, their properties, their microbes, the characteristics of microbes from extreme habitats.

Examination A written and graded examination (100% of final mark).

Literature None.

Additional Information None.

Module Name Marine Microbiology Module Number MNF-bioc-353

Person in Charge Prof. Dr. Johannes Imhoff Phone: +49-(0)431-600-4433, E-mail: [email protected]

Semester / Duration 2. or 3. semester / one semester

Regular Cycle every winter and summer semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Marine Microbiology (Seminar)

Prof. Dr. Johannes Imhoff 2 hrs per week / 10 students

Credit Points / Workload 2 ECTS / 60 hours

Prerequisites A bachelor’s degree in a biological discipline. Basics in marine microbiology.

Completion Module None.

Following Module None.

Educational Objectives Presentation of literature and own results.

Content of Teaching Discussion of new developments in Marine Microbiology.

Examination A written and graded examination (100% of final mark).

Literature None.

Additional Information None.

Module Name Functional morphology and physiology of marine organisms

Module Number MNF-bioc-354

Person in Charge Prof. Dr. Frank Melzner, Prof. Dr. Martin Wahl Phone: +49-(0)431-600-4274, E-mail: [email protected] Phone : +49-(0)431-600-4577, E-mail: [email protected]

Semester / Duration one semester

Regular Cycle annual in winter semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Functional morphology and physiology of marine organisms (Seminar) Prof. Dr. Frank Melzner Prof Dr. Martin Wahl

2 hrs per week / 30 students

Credit Points / Workload 2 ECTS / 60 hours

Prerequisites A bachelor’s degree in a biological discipline.

Completion Module None.

Following Module None.

Educational Objectives

In this seminar, students will study basic principles of marine animal physiology and functional anatomy in an interactive fashion. The seminar aims at developing an understanding of how form and function of marine invertebrates ensure ecological / evolutionary success. We will investigate key species from all important marine phyla / classes that occur in North- and Baltic Sea. Students will learn how to gather information on species from the primary literature and to prepare oral presentations and scientific essays (short review papers) for publication in internet databases.

Content of Teaching Specific adaptations (physiology, morphology, anatomy) of key marine taxa from North and Baltic Sea will be discussed. Introductions will be given by M. Wahl and F. Melzner, followed by student presentations and class discussions.

Examination 50% oral presentation, 50% semester work (preparation of an illustrated scientific essay (review paper) on a particular species of interest for publication in an internet database of Baltic and North Sea species).

Literature Relevant literature will be given out during the module.

Additional Information None.

Module Name Marine Animal Physiology Module Number MNF-bioc-355

Person in Charge Prof. Dr. Frank Melzner Phone: +49-(0)431-600-1500, E-mail: [email protected]

Semester / Duration one semester

Regular Cycle annual in winter semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Marine Animal Physiology (Lecture)

Prof. Dr. Frank Melzner

2 hrs per week / 30 students

Credit Points / Workload 4 ECTS / 120 hours

Prerequisites A bachelor’s degree in a biological discipline.

Completion Module None.

Following Module None.

Educational Objectives

In the lecture, basic principles of marine animal physiology will be taught in an interactive fashion. Topics include: osmo - and ion regulation, excretion, thermoregulation, neuro - and muscle physiology, behavioural physiology, physiology of organisms in extreme habitats (deep sea, hydrothermal vents, polar seas, cold smokers) etc. The lecture will build on topics covered in Marine Animal Physiology I (i.e. cellular physiology, metabolic biochemistry, oxygen transport systems, ventilation & circulation). The seminar will discuss interesting new research and synthesis papers published in integrative – physiological journals. Each student will deliver an oral presentation on a specific topic of relevance. Presented publications and the presentation itself will be critically evaluated by the class.

Content of Teaching On-going research in marine biogeochemistry will be presented and critically discussed. Current topics will be from a broad area of marine sciences with relevance to marine animal physiology, including molecular biology and genetics, ecology to marine chemistry.

Examination homework (100%)

Literature Relevant literature will be given out during the module.

Additional Information None.

Module Name Biogeochemistry of Marine Sediments II Module Number MNF-bioc-356

Person in Charge Prof. Dr. Tina Treude Phone: +49-(0)431-600-2837, E-mail: [email protected], Homepage: www.ifm-geomar.de/~ttreude

Semester / Duration 3. semester / one semester

Regular Cycle annual, in winter semester

Status

optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Biogeochemistry of Marine Sediments II (Lecture)

Prof. Dr. Tina Treude

1 hr per week / 30 students

Credit Points / Workload 2 ECTS / 45 hours

Prerequisites MNF-bioc-251 or a solid background in biogeochemical cycling

Completion Module None.

Following Module None.

Educational Objectives

The goal of this lecture is to gain a deeper understanding of the complexity of microbial and geochemical processes in marine sediments. Whereas the SS lecture (MNF-bioc-251) concentrated on single processes this follow-up lecture will focus on their synthesis. Additionally the students will gain some insights into important methods in benthic biogeochemistry.

Content of Teaching

The focus of this lecture will be on biogeochemical processes in complex benthic systems including biofilms and microbial mats, organic-rich sediments, extreme environments, permeable sediments, sulphur based food chains as well as symbioses with animals. Additionally some essential methods will be introduced to (1) identify microbial organisms and communities, (2) study microbial activity, and (3) follow microbial food webs in benthic biogeochemical systems.

Examination A short exam (graded) at the end of the semester.

Literature Canfield et al.: “Advances in Marine Biology: Aquatic Geomicrobiology”, Elsevier, ISBN: 0-12-026147-2; Konhauer: “Introduction to Geomicrobiology”, Blackwell, ISBN: 0-632-05454-9; Schulz et al.: “Marine Geochemistry”, Springer, ISBN: 3-540-66453-X

Additional Information

This lecture is interdisciplinary and addresses students from the fields of biological oceanography, geochemistry, and microbiology. The lecture will be given regularly every week. Please check UnivIS for exact dates.

Module Name Early Earth Life, Extreme Environments and Astrobiology

Module Number MNF-bioc-357

Person in Charge Prof. Dr. Tina Treude Phone: +49-(0)431-600-2837, E-mail: [email protected],

Semester / Duration 3. semester / one semester

Regular Cycle annual in winter semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Early Earth Life, Extreme Environments and Astrobiology (Seminar)

Prof. Dr. Tina Treude

3 hrs per week / 20 students

Credit Points / Workload 4 ECTS / 120 hours

Prerequisites A bachelor’s degree in a biological, geological or chemical discipline.

Completion Module None.

Following Module None

Educational Objectives The main goal of this seminar is to discuss recent hypotheses on how life and biogeochemical cycles developed on Earth or could develop on other planets. Students will gain practice in interpreting controversially discussed hypotheses.

Content of Teaching

Studies on early Earth life, modern extreme environments and astrobiology have one goal in common: they search for life under extreme conditions. In this seminar students will be introduced into: (1) interdisciplinary studies that try to reconstruct how life and biogeochemical cycles developed on Earth, and (2) systems that could serve as models for life development outer space. The majority of the environments discussed will be marine, some of them also limnic or terrestrial. In the first hour of each seminar an introduction lecture will be given by the lecturer. In the second hour, students will give presentations of selected literature prepared in homework. In the third hour, the literature will be discussed (sometimes accompanied by exercises).

Examination Oral presentation (pass / fail) of selected literature prepared in homework.

Literature

Schlesinger et al: “Biogeochemistry”, Elsevier, ISBN: 978-0-08-044642-4; Gilmour & Sephton: “Astrobiology”, Cambridge Open University, ISBN: 0-521-54621-4; Canfield et al.: “Advances in Marine Biology: Aquatic Geomicrobiology”, Elsevier, ISBN: 0-12-026147-2; Konhauer: “Introduction to Geomicrobiology”, Blackwell, ISBN: 0-632-05454-9 Selected literature will be provided during the course.

Additional Information

This lecture is interdisciplinary and addresses students from the fields of biological oceanography, geology, and microbiology. The lecture will be given regularly every week. Please check UnivIS for exact dates. For participation a subscription is necessary. Deadline is October 17th. For subscription please contact Tina Treude via email.

Module Name Phytoplankton: from genome to ecology Module Number MNF-bioc-358

Person in Charge Prof. Dr. Julie LaRoche Phone: +49-(0)431-600-4212, E-mail: [email protected]

Semester / Duration 3. semester / one semester

Regular Cycle annual in winter semester

Status

Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Phytoplankton: from genome to ecology (Lecture)

Prof. Dr. Julie LaRoche

2 hr per week / 15 students

Credit Points / Workload 2 ECTS / 60 hours

Prerequisites A bachelor’s degree a biological discipline.

Completion Module None.

Following Module None.

Educational Objectives

The educational objective is to familiarize the students with modern marine genomics and how it is applied to phytoplankton research. Familiarize the students with basic methods of bioinformatics and major databases such as NCBI, SWISSPROT, sequence homology (BLAST) searches.

Content Of Teaching

Journey through the major advances in marine genomics including all of the major phytoplankton groups: cyanobacteria (Prochlorococcus and Synechococcus), photosynthetic diazotrophs, diatoms and coccolithophorids. Advances made with metagenomics approaches, including the global ocean survey (GOS) dataset collected by Craig Venter. The genomic information will be related in the context of marine phytoplankton physiology, element cyclingm and marine biogeochemistry.

Examination A written and graded examination (100%) will cover all topics in this module.

Literature None.

Additional Information None.

Module Name Modern aspects of meteorology and oceanography: Carbon cycling in a changing climate

Module Number MNF-bioc-359

Person in Charge

Prof. Dr. Tina Treude Phone: +49-(0)431-600-2837, E-mail: [email protected] Prof. Dr. Birgit Schneider Phone: +49-(0)431-880- 3254, Email: [email protected]. Dr. Kirstin Krüger Phone: +49-(0)-431-600-4062, Email: [email protected]

Semester / Duration 3. semester / compact course

Regular Cycle Annually at the end of the winter semester

Status Optional

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Carbon cycling in a changing climate (Seminar) Prof. Dr. Tina Treude Prof. Dr. Birgit Schneider Kirstin Krüger

Compact course: 5 days, 7 hrs per day/ 20 students

Credit Points / Workload 5 ECTS / 125 hours

Prerequisites A bachelor’s degree in a biological, geological, physical or chemical discipline.

Completion Module None.

Following Module None

Educational Objectives

This interdisciplinary seminar should give an introduction into the role of carbon (including the greenhouse gases CO2 and methane) in the climate-system, with a focus on the coupling of atmosphere, ocean and solid earth. The seminar will teach the basics of the earth’s climate history, with an emphasis on the cycling of carbon through sediments, ocean, atmosphere and biosphere. Climate change issues will be discussed interactively with the students along the current fourth IPCC assessment.

Content of Teaching

The seminar is divided into interactive lectures connected to a student part with presentations and exercises. The student part includes studying and presenting recent articles on modern aspects of carbon in the climate-system. Within teams the students should learn to analyze and discuss scientific papers and IPCC chapters in more detail. Finally, a synthesis paper will be developed by all participants together.

Examination Oral presentation of literature (graded).

Literature John Houghton, Global Warming: The Complete Briefing, Cambridge University Press Additional literature and lecture notes will be distributed.

Additional Information Formal subscription to this course and participation in a planning meeting is mandatory; please check UnivIS for more information

Module Name Master Thesis Module Number MNF-bioc-401

Person in Charge Prof. Dr. Ulrich Sommer Phone: +49-(0)431-600-4400, E-mail: [email protected]

Semester / Duration 4. semester / one semester

Regular Cycle annual in summer semester

Status

Compulsory

Study Programme Master of Science in Biological Oceanography

Classes Class Title (Teaching Form) Lecturers Contact Time / Group Size

Master Thesis (Project)

Prof. Dr. Ulrich Sommer Prof. Dr. Martin Wahl Prof. Dr. Ulf Riebesell Prof. Dr. Thorsten Reusch Prof. Dr. Johannes Imhoff Prof. Dr. apl. Julie LaRoche

- / 30 students

Credit Points / Workload 30 ECTS / 900 hours

Prerequisites Successful completion of all course work within the Masters Program in Biological Oceanography.

Completion Module None.

Following Module None.

Educational Objectives Students will need here to demonstrate their ability to conduct scientific research in an independent manner, apply their theoretical knowledge to a practical or conceptual scientific question and be able to communicate their results in a lucid manner.

Content of Teaching Students are required to submit a master thesis consisting of original, independent scientific work. The thesis will be accompanied by an oral presentation and defence.

Examination Written thesis (75%) and thesis defence (25%)

Literature None.

Additional Information None.