Maldanid Key All OSEANOGRAFI UNIVERSITAS DIPONEGORO

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D. L. O'DonohueRev. February, 1994

MALDANIDS KNOW N FROM CITY OF SAN DIEGO OCEAN MONITORING PROGRAM

Family M aldanidaeSubfamily Clymenurinae

Cb menura gracilis Hartman, 1969Subfamily Euclymeninae

Cbjmenella complanata Hartman, 1969 — .[Euclymene grossi newporti Berkeley and Berkeley, 1941 J - X€Mc\\j ^€^€ jEuclym eninae sp. A SCAMIT ~ Îsocirrus longiceps (Moore, 1923) <o * €. €. v\ <\ c\ Praxillella gracilis (Sars,1861)

f

Praxilklla pacifica Berkeley, 1929Subfamily Lumbriclymeninae

Notoproctus pacificus (Moore, 1906)Praxillura maculata M oore, 1923

Subfamily NicomachinaeNicotnache lumbricalis (Fabricius, 1780)Petaloproctus neoborealis H artman, 1969

Subfamily MaldaninaeMaldane sarsi M almgren, 1865Metasychis disparidentata (M oore, 1904) fide L ight, 1991

Subfamily Rhodininae

Rhodine bitorquota Moore, 1923

KEY TO TH E MALDA NIDS COLLECTED OFF POINT LOMA

la . Cephalic plaq ue presen t (Euclymeninae or M aldaninae) 6lb . Cephalic plaq ue absent (Lumbriclymeninae, Nicomachinae or Rhodininae) 2

2a. Rostrate neuro setae in doub le row s from setiger 5 ; conspicuous collars directed forward onsetigers 2 and 3 (Fig. 1) Rhodine bitorquata

2b . Rostrate neuros etae in single row s 3

3a. Acicular neu ros etae pres ent in first 6-9 setigers (Fig. 2) Praxillura maculata3b . Acicular neu ros etae restricted to first 3-4 setigers 4

4a. Acicular neuro setae restricted to first 3 setigers; anterior end short, thick an d rou nde d;wh ole specimen ha s cup-like anal funnel, its margin w ith a circlet of 14-21 short teeth(Fig. 3) Nicomache lumbricalis

4b. Acicular neu rose tae restricted to first 4 setigers 5

5a. Prostomium broadly roun ded and produced ; white dorsal internotopodial ban ds onred-b row n base color; anal plaqu e forms flat plain with a nal pore do rsal to plaq ue (Fig. 4)

Notoproctus pacificus5b. Prostom ium broad ly truncate an d comp ressed; anal plaq ue large, foliose and produ ced

ven trally (Fig. 5) Petaloproctus neoborealis



6a. Cephalic plaqu e with m edian keel low or absent 76b . Cephalic plaque with well developed (high) med ian keel; lateral lobes of cephalic pla que

smo oth; anal plaq ue broa dly o val, wi th ma rgin en tire except for lateral notches (Fig. 6)Maldane sarsi

7a. Cephalic plaqu e with out m edia n keel; lateral lobes of cephalic pla qu e not crenulate 87b . Cephalic plaq ue wi th low keel; lateral lobes of cephalic plaq ue strongly crenulate; ana l

plaq ue divid ed into two sm ooth lobes; enlarged do rsal lobe petaloid in shap e, ven tralm argin forms d eep funnel-like p ocket (Fig. 7) Metasychk disparidentata

8a. Cephalic plaqu e develop ed, lateral lobes w ith pro min ent flange (see Fig. 11); rostrate unciniin anterior neu ropo dia (larger specimens m ay app ear to have acicular uncini) 9

8b . Cephalic plaqu e flat, smo oth wi th a dee p transverse groov e me dially, lateral lobes with outflange; acicular spines in an terior neu rop odia (Fig. 8) ClymeneUa complanata

9a. Lateral lobes wi th or with out flange, lateral notches absent 109b. Lateral lobes w ith well develop ed flange, lateral notch es prese nt (see Fig. 11) 11

10a. Lateral lobes w ith poo rly develope d flange; cephalic plaq ue long, sloping posteriorly; 0

shap ed gland ular area absent from setiger 8; anal pore surrounded by about 30-33 trimcren ulation s (Fig. 9) Isocirrus longiceps

10b. Lateral lobes w ith dev elope d flange; large 0 shap ed glan dular area o n setiger 8 (visiblewith m ethyl green stain); anal pore su rround ed by ab out 20 cirri (Fig. 10)

Ctymenura gracilis

11a. Palp ode pro lon ged into a digita te extensio n (Figs. 11-12) Praxillella gracilis1 lb . Palpode not prolonged; methyl green staining pattern used to distinguish amo ng remaining

species (Staining process needs to be d one w ith con centrated stain. Leave in stain for at least30-60 min.) setigers 4-8 stain da rk gre en (Figs. 12-13) PraxiUella pacificaven tral stripes from set. 9-14 (Fig. 14) Eucly me ninae s p. Asimilar to P . pacifica bu t with w hite girdle o n set. 8 (Fig. 15) Eticlymene grossi newporti

Figure 1. Rhodine bitorquata : anteriorend , lateral view



F*»M F»Oft1Ac0t W

/ W r . / P ^ r . ?lf\o&& ^ 6 « N ) r

MALDA NIDS KNOWN FROM CITY OF SAN DII p ^ . 5fT- M> iTM £ACi e c u Ah 6DUA«

5 ^ OtfCJN' ' * VoobU J2o^S

Fami ly Ma dan idae (* e f c , t t ' / M W W A «Subfamily Clymenunnae I A , £ / j L r pLA<t06> A f t ^ r

C/ym enuraraaV /s Hartm an, 1969 I W ?©>»• »^nH

? -

Subfami ly Euclymeninae AvL * £ * » » * » S ^ T A S ;

Clymenella complanata Hartm an, 1969 ^ £ ,̂ N A e

Euclymene grossi nezoporti Berkeley an d F ̂ ft - pt^c i0 6 A ^ C A i T

Euclymeninae sp . A SCAMIT K u t P^f t .Of pn *S *^ T

Isocirrus longiceps (Moo re, 1923) U ^ C '^ t i-4 /»fcVic P© * ^Praxz7 k//ac raa7 fs(Sars,1 861 ) /••• w/flCAi Û2-'M E> „ _ , _ . hW

Praxillella paafica Berkeley, 1929 J P / p c s r PLACM* VW OH A < ^ rSubfamily Lumbriclymeninae

Notoproctus pacificus (M oore, 1906) MAUD<WIA)A£

Praxilluramaculata M oore, 1923 AiO f/poST. P iA O .0 0 p<tf<»LA/r

Subfamily Nicom achinae fitsjO^ po(2-£A*-

Nicomache lumbricalis (F abricius, 1780)Petaloproctus neoborealis H artm an, 1969

Su bfa mily M alda nina e AACCy|V\ej\JffJA6 •

MaW rmesflrsfMalmgren,1865 ftislT/poST. p t ^ £ * W > * « n

Metasychis disparidentatt£(Moore, 1904) f . c ~-p fl_/vi«/* A*'Subfamily Rhodininae ***

1 D

Rhodine bitorquata Moore, 1923

KEY TO THE M ALDANIDS COLLECTED OFF POINT LOMA

la. Cephalic plaque present (Euclymeninae or M aldaninae) 6l b . Ce phalic plaq ue absent (Lumbriclymeninae, Nicom achinae or Rho dininae) 2

2a. Rostrate neuro setae in double rows from setiger 5; consp icuous collars directed forward onsetig ers 2 an d 3 (Fig. 1) Rliodine bitorquata

2b. Rostrate neu roseta e in single rows 3

SrfAU sp ec if ic ^ vJi~ ' ,r i 'V 4 M^CsC-l. w.<fti (; < . '% r»tu,:«s:•-.-;.' [ttfi**6• WLQ

3a. Ac icular neu rose tae pre sen t in first 6-9 setigers (Fig. 2) Praxillura maculata

3b. Ac icular neu ros etae restricted to firsj 3-4 setigers :.. 4

4a. Acicular neu roseta e restricted to first 3 setigers; anterio r end short, thick and r ou nd ed;wh ole specim en ha s cup-like anal funnel, its m argin w ith a circlet of 14-21 short teeth(Fig. 3) Nicomaclie lumbricalis

4b. Acicu lar neu rose tae restricted to first 4 setigers 5

~ <

5a. Prosto m ium broadly roun ded and produ ced; wh ite dorsal interriotopodial ban ds onred -br ow n base color; anal plaque forms flat plain wit h anal pore d orsal to plaqu e (Fig. 4)

Notoproctus pacificus 1

5b. Pros tom ium broadly truncate and compressed; anal plaqu e large, foliose and produ ced V ^ve ntr al ly (Fig. 5) :'. Petaloproctus neoboreali



Figure 2. Praxilluramaculataanterior end, lateral view

acicular spine

single rowof rostrateuncini

B.

Figure 3. Nicomache luTnbricalis : A. anterior end, lateral view;B. posterior end, ventral view



\ f f qis

f; f̂&e*-

B.

Figure 4 . Notoproctus pacificus : A . anterior end, lateral view;

B. posterior end , lateral view

^ V ^ - . o êc^

B.

Figures . Petahproctus neoborealis : A . anterior end, lateral view;B. posterior end, lateral view; C anal plaqu e, posterior view



RRowe20NOV95

Separating Maldane sarsi Metasychisdisparidentatus

Maldane sarsi

An terior end formed into acephalic plaque with smoothlateral and posterior margins

\

Posterior end formed into ananal plaq ue which is flat withslightly raised smooth tocrenulate margins

•Anterior-margins

crenulate

Metasychis disparidentatus

posterior end

anterior

m arg ins can-be crenulated

Posterior end withlarge petaloiddorsal margin &deep funnel-likepocket formed by

ventral m argin



A. B.

Figure 6. Maldanesarsi A. anterior view; B. anal plaque,posterior view

B.

Figure 7. Metasychis disparidentata A. anterior end, lateral andventral view; B. anal plaque, lateral and dorsal view



transverse groove

B.

Figure 8. Ch/menella complanata : A. anterior end, lateral view; B. cephalic plaquewith transverse groove; C posterior end, lateral view

B.

Figure 9. Isocirrus longiceps : methyl green staining pattern A. anteriorsegments 1-8, lateral view; B. posterior end , lateral view



pr^

middorsal notch only , no lateralnotches

neurosetae of first 3 setigers consist1-3 modified rostrate uncini: mainfang plu s 4-5 teeth, no beard

setiger 3 with stain aroundparapodial tori

setiger 4 solid (medium ) green

setigers 5-8 with dark bandinganteriorly on each setiger,banding stops after set. 8

Figure 10. Clymenura gracilis : methyl green stainingpattern ; anterior segments 1-8, lateral view



palpode prolonged intodigitate extension

Figure 11. Praxillella gracilisanterior end, dorsal view

setiger 2 with lightgreen semicircles onventrum

setigers 4-8 more or lessuniformly dark

no stripes on ventrum

12. typical PraxiUella pygidium Figure 13. Praxillella padfica : m ethyl greenstaining pattern A. anterior segments 1-10,

lateral view; B. anterior segments 1-2,ventral view.



dark band on setiger 8,thereafter on ly parapodial •tori stain

stripes extend from setiger 9through setiger 13/14 ;thereafter only areas aroundparapodial tori remainstained

light band ing anterioron setigers 1-3

solid, light banding onsetigers 4 and 5

A B

Figure 14. Euclymeninae s p. A: methy l greenstaining patter n A. anterior segmen ts 1-11,

lateral view; B. anterio r segme nts 1-11, ventra lview

A B

Figure 15. Euclymene grossanexvporti : methygree n stain ing pa ttern A. anterior segments 1-10

lateral view; B. a nterior segm ents 7-10, ventralview



Cor^pAR\5oiO of

Sucly fr ieze ro^sa. yyetspcr

BMP

setiger 2 with lightgreen semicircles onventrum

setigers 4-8 m ore or lessuniformly dark

fflscicte

no stripes on ventrum

Figure 13. Praxillella patifica : methyl greenstaining pattern A. anterior segments 1-10,

lateral view; 6 . anterior segm ents 1-2,ventral view.

light banding anteriorlyon setigers 1-3

Figure 15. lEuclymene grossdyiavporti}. methyl green staining pa ttern A. anterior segments 1-10,lateral view; B. an terior segments 7-10, ventral

view



A x i o t h e l l a r u b r o c x n c t a ( Jo h n so n , 1901)Maldan idae , Euc lymen inae SCAMIT Vol. 6 No.

SCAMIT Code: HYP 44 (in part) Date examined: 13 July 1987Voucher by: Leslie H. Harris (AH

SYNONYMY: Clymenella rubrocincta Johnson, 1901

LITERATURE: Arwidsson, 1907 (placed into unnamed new genus)Monro, 1937 (put into Axiothella) ~Berkeley and Berkeley, 1941 (Axiothella)Hartman, 1969 (Axiothella)Clark and Dawson, 96 3 (discusses generic confusion)Harris, 1985 (SCAMIT Newsletter, vol. 3, no. 12)Imajima and Shiraki, 1982

DIAGNOSTIC CHARACTERS:

1.

2.

3.

Nuchal organs and carina extend 2/3 length of cephalicplaque; slight lateral notches, deeper mid-dorsalnotch; proboscis smooth.

Usually thick fold on anterior margin of setiger(pseudocollar).

Neuropodial uncini of first three setigers with 4 teetabove main fang, no beard; number varies with size,holotype (162 x 3.5mm has 4 uncini in 1st setiger, 6uncini in 2nd setiger, 7/8 uncini in 3rd setiger; anoth

specimen (40 x 1.5mm has 4 uncini in each of first thsetigers. Neurosetae in following setigers with 5-6teeth above main fang, beared, 12 to 24 in number.

Notosetae consist of bilimbate capillaries, hirsutecapillaries, and simple capillaries.

Nephridial pores on setigers 7-8-9.

Specimens have 18 setigers and 2 pre-anal asetigeroussegments.

Pygidium with callus ring and anal funnel rimmed with18-30 filaments alternating long and short, the mid-ventral filament longest.

Methyl green staining pattern (Fig. 1)- cephalic plaque, head light.- presetal portions of setigers 1-8 dark, postsetal

portions light; bands of setigers 4-8 darkest.- no color after setiger 8 except for slight trace's

around tori.



Axiothella rubrocincta (Johnson, 1901)Maldanidae, Euclymeninae SCAMIT Vol. No. 5

RELATED SPECIES AND CHARACTER DIFFERENCES:

1. Axiothella rubrocincta complexa Berkeley and Berkeley,

1941: 3-4 lateral notches on each side of cephalicrim; low collar on anterior margin of 4th setiger.

2. Clymenura gracilis Hartman, 1969: Cephalic rim smooth,only mid-dorsal notch; rounded ventral projectionof glandular band on setiger 8; papillated proboscis;5-6 asetigerous pre-anal segments.

ADDITIONAL NOTES:

Because specimens under 0.75mm in width may only have1-2 uncini in the first three neuropqdia, A. rubrocincta

is easily mistaken for other species -=which have acicularhooks. Great care must be taken in examination ofsmall specimens.

REMARKS: A. rubrocincta has been one of the commonest maldanidsreported in environmental studies, occurring fromthe intertidal zone through canyon and basin depths.Examination of specimens, however, revealed fourundescribed and three described euclymenid speciesmisidentified under this name. In reality, A. rubrocinctais found in shallow water, rarely deeper than 20-30meters.

DISTRIBUTION:

Western Canada through the Mexican border. Reported fromintertidal, shelf, basin and canyon depths, but in centraland southern California, found mostly in bays, mudflats,and shallow shelf. Recorded from Japan, 130-295 meters.



a rubrocincta (Johnson, 1901)SCAMIT Vol. 6 No.

Figure 1. Axiothella rubrocincta (Johnson, 1901). Methyl green



Euclymeninae sp. A SCAMITMaldanidae SCAMIT Vol. 6 No. 5

SCAMIT Codes: HYP 44 (in part), Date examined: 13 July 1987

LACO 49, SCCWRP 54 Voucher by: Leslie H. Harris (AHF)

SYNONYMY: Euclymeninae sp. C Harris

LITERATURE: Harris, 1985 (SCAMIT Newsletter, vol. 3, no. 12)


1. Nuchal organs and carina extend 3/4 - 4/5 length ofcephalic plaque; slight lateral and mid-dorsal notches,posterior margin smooth.

2. Presence of thick fold (pseudocollar)i

on 4th setigervariable, depends on degree of contraction.

3. Neurosetae of first three setigers single acicularspines, bent at tips. Neurosetae of following setigersrostrate uncini with 5 teeth above main fang plusaccessory denticles (see additional remarks).

4. Notosetae consist of narrow-edged limbate capillariesand laterally hirsute th-inner capillaries.

5. Nephridial pores on setigers 7-8-9 usually obscure.

6. Complete specimens have up to 27 setigers (numbervaries with size); no asetigerous pre-anal segments.

7. Pygidium with pronounced callus ring (flange) andanal funnel with up to 27 filaments (number varieswith size); midventral filament longest, others alternate1-2 short and 1 long. No anal cone, except as artifactof contraction.

8. Staining pattern (Fig. 1)- palpode and inside of flange dark, rest of plaque

light.- presetal portions of setigers 1-8 dark, postsetal

portions light; bands on setigers 6-7-8 darkest.- lateral and ventral stripes extend from setiger 9

through 13/14; thereafter only area around parapodialtori stained. No dorsal stripe, slight triangularextension on setiger 9.



Euclymeninae sp. A SCAMIT

haidanidae SCAMIT Vol. 6 No.


1. Euclymeninae sp. B: Different staining pattern,

especially in possession of a dorsal stripe throughsetigers 11-12, darkest bands on setigers 2-3-4, darkcarina; two asetigerous pre-anal segments; anal filamesubequal in length.

2. Clymenella californica Blake and Kudenov, 1974: Laterventral and dorsal stripes through setiger 14, pigmentareas around posterior tori extend and form encirclingbands; one asetigerous pre-anal segment; anal filamentsubequal; anal cone may be present. ;

3. Euclymene campanula Hartman, 1969: 19 setigers; 7-8

asetigerous pre-anal segments; posterior segmentsstrongly campanulate.

4. Euclymene grossa newporti Berkeley and Berkeley, 1941:19 setigers; two asetigerous pre-anal segments; postermargin of cephalic plaque serrated; no stripes instain pattern.

5. Praxillella pacifica Berkeley, 1929: Different- stainipattern (Fig. 2) : setigers 4-8 more-or-less uniformlydark, no great contrast*in presetal and postsetalportions of setigers, no stripes; anal cone present,

usually projecting from anal funnel.

6. Praxillella gracilis (Sars, 1861): Stain patternand pygidium as in P. pacifica; palpode prolongedinto digitate extension.

REMARKS: Euclymeninae sp. A and B belong in a generic complexthat is the most poorly defined in the family MaldanidaThe characters used to separate such genera as ClymeneVerrill, 1873, Axiothella Verrill, 1900, EuclymeneVerrill, 1900, and Praxillella Verrill, 1881, areconsidered inadequate by some authors because of theirvariability or because the same characters are used .on the species level within these genera. Addingto the difficulty of distinguishing between generaand assigning species is the vagueness of originaland even emended descriptions, plus the practice ofdescribing animals from incomplete specimens.

The taxa listed under Related Species all have similarcephalic plaques, acicular spines in the first threeneuropodia, and anal funnels rimmed with many filaments



eninae sp. A SCAMIT

SCAMIT Vol. 6 N

(continued)

Euclymene delineata Moore, 1923 and E. reticulataMoore, 1923, have not been included because they wer

described from anterior fragments; until examinationof the holotypes they should be considered nomendubia. Euclymeninae sp. A and B have been previouslyidentified as all of these, and as Axiothella rubroc(Johnson, 1901), which has modified uncini in' thefirst three neuropodia.

ONAL NOTES:

Juveniles may have modified uncini instead of simple

acicular hooks in the first three neuropodia becausethe type of uncini change with size in some species.The stripe pattern is constant in adults, but inspecimens under 0.75mm in width, the stripes becomespotty and may disappear entirely leaving only pigmenareas around the tori. The stain pattern of setigers1-9 remains th<= same regardless of size.

Euclymeninae A is one of the two most abundant maldanin shelf depths throughout southern California insoft sediments.



Euclymeninae sp. A SCAMITMaldanidae SCAMIT Vo l. 6 No. 5

Figure 1. Euclymeninae sp. A SCAMIT. Methyl green stainingpattern. A, anterior end, lateral view; B, anteriorend, ventral view.



Euclymeninae sp. B SCAMITMaldanidae SCAMIT Vol. 6 No.

SCAMIT Code: HYP 44 (in part) Date examined: 13 July 1987

Voucher by: Leslie H. Harris AHF)

SYNONYMY: Euclymeninae sp. D Harris

LITERATURE: Harris, 1985 (SCAMIT Newsletter, vol. 3, no . 12)


1. Nuchal organs and carina extend 3/4 - 4/5 length ofcephalic plaque; slight lateral and mid-dorsal notches,posterior margin smooth.

2. Presence of thick fold (pseudocollar)-on 4th setigervariable, depends on degree of contraction.

3. Neurosetae of first three setigers single acicularspines, bent at tips. Neurosetae of following setigersrostrate uncini with five teeth above main fang plusaccessory denticles. Juveniles may have modifieduncini instead of acicular spines.

4. Notosetae consists of narrow-edged limbate capillariesand laterally hirsute thinner capillaries.

5. Nephridial pores on setigers 7-8-9, usually difficultto distinguish except in reproductive specimens.

6. Complete specimens have up to 21 setigers (number'varies with size) and two asetigerous pre-anal segments

7. Pygidium with pronounced callus ring (flange) andanal funnel with up to 19 subequal filaments (numbervaries with size). No anal cone, except as artifactof contraction.

8. Staining pattern (Fig. 1)- palpode, marginal flange and carina dark, rest of

palpode light.- presetal portions of setigers 1-6 usually darker

than postsetal sections, distinction may be blurredon setigers 4 and 5; bands on setigers 2-3 darkest.

- setiger 7 with three colored areas ventrally, twodorsally.

- lateral and ventral stripes extend from setiger 9through tori of 14th, thereafter only area aroundtori darkly stained. Dorsal stripe from setiger 9through setiger 11 or 12.



Euclymeninae sp. B SCAMIT

Maldanidae SCAMIT Vol. 6 No. 5


1. Euclymeninae sp. A: Different staining pattern,

especially absence of dorsal stripe; darkest bandson setigers 6-7-8; unstained carina; no asetigerouspre-anal segments; anal filaments alternate long andshort with mid-ventral one longest.

See other species listed for Euclymeninae sp. A.

ADDITIONAL NOTES:

See Euclymeninae sp. A.

REMARKS: See Euclymeninae sp. A.

DISTRIBUTION:

Euclymeninae sp. B is infrequently encountered insouthern California; it is found in soft sedimentsbetween 50 and 200m.



Euclymeninae sp. B SCAMITMaldanidae SCAMIT Vol. 6, No. 5

Figure 1. Euclymeninae sp. B SCAMIT. Methyl green staining pattA, anterior end, lateral view; B, posterior end'; C, set6 to 9, ventral view; D, setigers 6 to 9, dorsal view;E,F, cephalic plaques; G, prostomium, ventral view.



2 1 6 BULLETIN OF MARINE SCIENCE. VOL. 4«. NO. i 1991

Figure 1. Maldane caUforniensisr A, Cephalic plaque and first setiger, lateral view, ventral collarunshaded and to left o f notopodia; B , Cephalic plaque, top view; C, Anal plaque and setigers 17 -19 ,dorso-lateral view, glandular areas unshaded on neuropodial tori and on preanal pads below last

setiger; D, Anal plaque, bottom view , dorsal side down: E, Anterior nine setigers, shading indicatesmethyl green staining pattern on lateral (drawing on left), dorsal (drawing at center), and ventral



Maldane sarsi Malmgren, 1865Maldanidae

Vol. 3, No. 12

SCAMIT Codes: LACO 47, PL 53 Date examined: March 3, 1985Voucher by: Karen Green (MEC

Literature:Arwidsson, 1907Fauvel, 1914Green, 1984

Diagnostic characters:

1. 19 setigerous segments, 2 asetous preanal glandular pads.

2. Cephalic plaque with smooth lateral and posterior margins.Posterior margin forms deep pocket. Prostomiai palpode semi-triangular, keel long and high, and nuchal organs short andgently curved.

3. Anal plaque divided into two reduced lobes; dorsal margin smooth,ventral margin smooth to crenulate.

4. Dorsal glandular band behind glandular tori of setiger 5«

Additional notes:The syntype of Maldane cristata and type material of M. sarsi wereexamined and are equivalent; records of M. glebifex from the easternPacific are questionable (Green, 1934; unpubiishec thesis).

Distribution:Alaska south to Antarctica, Japan, Arctic Ocean, 3affin Bay, DavisStrait, and Skagerrak. Depths from 3 to 5034 a.

drawings from Green,



(Ae+A. ycUis4es*oh£& d i s -p a r i d en t a t* (Moore , 1904)Maidanidae

Vol. 3, No. 12

SCAMIT Codes: LACO 43 , AHF 31 Date examined: March 3, 1985

Voucher by: Karen- Green (MEC)

Synonymy:Maid an e d i s -o ar id e n ta ta Moore, 1904

L i t e r a t u r e :Hartman, 1969Faucha ld , 1972Ima j ima and S h i r a k i , 1982

D i a g n o s t i c c h a r a c t e r s :1 . 19 s e t i g e r o u s s e g m e n t s , no a s e t o u s p r e a n a l g l a n d u l a r p a d s .

2 . C e p ha lic p l aq u e w i t h s e r r a t e l a t e r a l a nd p o s t e r i o r m a r g i n s .

F r c s t o m i a l p a lp o d e b r o a d l y r o u n d e d , k e e l o f m o d e r a te l e n g t han d l o w , an d n u c h a l o r g a n s s h o r t - m o d e r a t e l e n g t h v ; i thc u r v e d t i p s .

3 . A n a l p l a q u e d i v i d e d i n t o tv/o s m o o th l o b e s ; e n l a r g e d d o r s a ll o b e p e t a l o i d i n s h a p e , v e n t r a l m a r g i n fo rm s d ee p f u n n e l - l i k ep o c k e t .

it. F i r s t s e t i g e r w i th v e n t r a l c e l l a r - l i k e f o l d .

D i s t r i b u t i o n :W e s te rn C an ad a s o u t h t o G ulf o f C a l i f o r n i a ; J a p a n . Shelf, b a s i n ,

a nd c an yo n d e p t h s i n c o a r s e s a n d , s h e l l y m ix ed s e d i m e n t s , s i l t ysands, and mud.



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