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Limnology(lentic = lakes and ponds; lotic = streams and rivers)
Properties of Lakes
• Origins
• Water Balance
• Light
• Heat and Stratification
• Oxygen
• Nutrients: Nitrogen and Phosphorous
Lake Zones
Generalizations of Lake Physical
Properties
Lakes evolve by succession
The more shoreline – the more mature the
lake
Lakes change from deep to shallow
Lakes become more biologically productive
with succession
The Maturing of a Lake
Chemical Factors of Trophic
Lakes
Origins of lakes
1. Tectonic
2. Volcanic
3. Landslides
4. Glacial action
5. Solution of rock
6. River activity
7. Shoreline activity
8. Biological (beavers)
9. Human
Spirit Lake
Bonneville ReservoirGlacier Bay
The distribution of reservoirs
Distribution of reservoirs over 500 acres in the United States, by surface
area, January 1970. One dot equals 10,000 acres. Total area, 90 million
acres (Jenkins, R.M. 1970. AFS Spec. Publ. No. 7).
Limnology: water balance
Water inputs:
Precipitation, surface runoff from basin, groundwater, diversion
Water losses:
Drainage, seepage, evaporation and evapo-transpiration (plants), diversion
Mono Lake, CA
Soap Lake, WA
Global water supply
Water body Volume (km3 * 1000)
% of total Renewal time (years)
Oceans 1,370,000 97.61% 37,000
Polar ice,
glaciers
29,000 02.08% 16,000
Groundwater 4,000 00.29% 300
Freshwater
lakes
125 00.009% 1-100
Saline lakes 104 00.008% 10-1,000
Atmospheric
water vapor
14 00.0009% 9 days
Rivers 1.2 00.00009% 12-20 days
Adapted from Wetzel 1975
Limnology: light
http://www.ucar.edu/learn/images/spectrum.gif
Limnology: light
Absorbance of different colors
of light on a log scale in a clear
unproductive lake.
Note the rapid loss in the
quantity, and change in the
quality of light. Pattern vary
among lakes, depending on the
nature of material in the water
such as glacial flour, tannin,
phytoplankton, etc.
Clear, relatively unproductive lake
Limnology 1994
Depth
(m
)
Percent surface light0.1 1.0 10 100
15
10
5
0 0
5
10
Limnology: light
0.1 1.0 10 100
Absorbance of different colors
of light on a log scale by a clear
unproductive lake and a highly
productive, shallow reservoir.
Clear, relatively unproductive lakeShallow, productive reservoir
Limnology 1994
Depth
(m
)
Percent surface light0.1 1.0 10 100
15
10
5
0 0
5
10
Heating and cooling
Sources of heat:
1. Direct solar radiation (most important)
2. Groundwater and springs
3. Ground (minor)
Losses of heat:
1. Thermal radiation (primary)
2. Conduction
3. Evaporation
4. Outflow
Heat and the density of water
Temperature (C)
Density g
/cm
3
The maximum density
of water is at 3.94 C.
Ice at 0 C is 8.5%
lighter than liquid
water at 0 , so it floats.
Above 3.94 water
gets lighter as it
warms up, so the
surface water is also
warmer than deep
water in summer.
x
Distribution of heat
Hypolimnion
Metalimnion
Epilimnion
Temperature (C)
Depth
(m
)
Horizontal distribution is
primarily driven by wind
Vertical stratification is
seasonal, driven by the
relationship between
density and temperature
Mixing and Satratification
Limnology: heatD
epth
(m
)
Time (month)
Ice Ice
Dimictic Lake (turns over twice a year)
Limnology: heat
Time (month)
Depth
(m
)Monomictic lake (turns over once each year)
Limnology: heatThe distribution of heat within a
lake depends on water movement
1. Surface waves
2. Langmuir circulation waves
3. River Influents
Clear Lake, California
Dissolved Oxygen
Dissolved Oxygen
•Air contains 300 mg O2/L; the
rest is mostly nitrogen
•Water contains 14.6 mg O2/L
•Gasses diffuse slowly in water,
so distribution is governed by
circulation, not diffusion
Eutrophic Lakes: hypolimnion
depleted by decomposition
(biotic processes)
Oligotrophic Lakes:
Distribution governed by
physical processes
In warm water, the metabolic
demands of fishes tend to increase,
and the capacity of the water to hold
oxygen decreases.
Dissolved Oxygen
Winter Kill:
Occurs when a shallow lake is ice-covered and hence dark for a long period. The DO demand of plants can drive levels below the 2 mg/l required by most fish.
Summer Kill:
Occurs when dense concentrations of macrophytes die at the end of their growing season. Decomposition can drive DO levels down. If the lake is shallow and macrophytes are found over the whole surface, a “squeeze” between warm surface temperatures and low DO levels on the bottom in late summer can kill fish.
Depth (cm)
Oxyg
en
(m
g/l)
0 50 100 150 200 250 300 350
05
1
0
1
5
2
0
Relationship between dissolved
oxygen level and depth below the ice
in Black Lake, Alaska.
G. Ruggerone
8 mg/l is needed to
maintain healthy salmon
050
100
150
Oxygen (
% o
f satu
ration)
Pe
rce
nt
su
rviv
al
Oxygen (% saturation)
0 25 50 75 100
0 2
0
4
0 6
0
8
0 1
00
Survival of juvenile sockeye salmon in Black Lake
at different dissolved oxygen concentrations (Ruggerone 1999)
Development of temperature – dissolved oxygen “squeeze”
from summer-kill conditions in Lake Sammamish
Data: Hans Berge, Metro – King County
Limnology:
Nitrogen and Phosphorous
Nitrogen and Phosphorous are
essential elements for living
organisms, and one, the other,
or both in concert may limit
primary and secondary
production. In lakes, nitrogen
comes primarily from microbial
action or terrestrial runoff and
phosphorous primarily from local
geology. However, both can be
greatly affected by human
inputs.
Limnology:
Nitrogen and Phosphorous
Simplified Nitrogen cycle
Primary and secondary production may be
limited by Nitrogen or Phosphorus
Total phosphorous
Fis
h b
iom
ass (
kg/h
a)
Fis
h b
iom
ass (
kg/h
a)
Macro-benthos biomass
Natural and human sources for Nitrogen and
Phosphorous