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Evidence of EvolutionRecent Fossil Finds Darwin also struggled with what he called the imperfection of the geological record. Darwins study of fossils had convinced him and other scientists that life evolved.

But paleontologists in 1859 hadnt found enough fossils of intermediate forms of life to document the evolution of modern species from their ancestors. The Age of Earth and FossilsHow do fossils help to document the descent of modern species from ancient ancestors?

Many recently discovered fossils form series that trace the evolution of modern species from extinct ancestors.

Recent Fossil Finds Since Darwin, paleontologists have discovered hundreds of fossils that document intermediate stages in the evolution of many different groups of modern species. The Fossil RecordRemains of animals and plants found in sedimentary rockdeposits give us an indisputable record of past changes through vast periods of time.

The Fossil RecordThe more that we learn about the evolution of specific species lines, the more that these so-called gaps or "missing links in the chain of evolution" are filled with transitional fossil specimens.Fossils and Ancient Life Paleontologist are scientist who study fossilsNature of Fossils:Fossils are traces of organisms that have been dead for a long timeFossils are often found in sedimentary rockOver time, hard minerals can replace hard body parts or tissueThe fossil record provides evidence about the history of life on Earth

Recent Fossil Finds One recently discovered fossil series documents the evolution of whales from ancient land mammals. Several reconstructions based on fossil evidence are shown on the following slides. The exceptions to the reconstructions are the modern Mysticete and Odontocete.Recent Fossil Finds

Recent Fossil Finds

The limb structure of Ambulocetus (walking whale) suggests that these animals could both swim in shallow water and walk on land. Recent Fossil Finds

The hind limbs of Rodhocetus were short and probably not able to bear much weight. Paleontologists think that these animals spent most of their time in the water.Recent Fossil Finds

Basilosarus had a streamlined body and reduced hind limbs. These skeletal features suggest that Basilosarus spent its entire life swimming in the ocean.Recent Fossil Finds

Modern whales retain reduced pelvic bones and, in some cases, upper and lower limb bones. However, these structures no longer play a role in locomotion.Recent Fossil FindsOther recent fossil finds connect the dots between dinosaurs and birds, and between fish and four-legged land animals.

All historical records are incomplete, and the history of life is no exception. The evidence we do have, however, tells an unmistakable story of evolutionary change.Comparing Anatomy and EmbryologyBy Darwins time, scientists had noted that all vertebrate limbs had the same basic bone structure. Similarities in Embryology Early stages of different vertebrate embryos are very similar to each otherMany vertebrates have gill like structures at some point as an embryo

Homologous Structures Darwin proposed that animals with similar structures evolved from a common ancestor with a basic version of that structure.

Structures that are shared by related species and that have been inherited from a common ancestor are called homologous structures. Comparing Anatomy and EmbryologyFor example, the front limbs of amphibians, reptiles, birds, and mammals contain the same basic bones.

Homologous Structures Homologous bones, as shown by color-coding, support the differently-shaped front limbs of modern vertebrates.

Homologous Structures These limbs evolved, with modifications, from the front limbs of a common ancestor whose bones resembled those of an ancient fish.

Similarities and differences among homologous structures help determine how recently species shared a common ancestor.

For example, the front limbs of reptiles and birds are more similar to each other than either is to the front limb of an amphibian or mammal. This similarityamong many othersindicates that the common ancestor of reptiles and birds lived more recently than the common ancestor of reptiles, birds, and mammals. Analogous Structures The clue to common descent is common structure, not common function. A birds wing and a horses front limb have different functions but similar structures.

Body parts that share a common function, but not structure, are called analogous structures. The wing of a bee and the wing of a bird are analogous structures.Genetics and Molecular BiologyDarwin had no idea how heredity worked, and he was worried that this lack of knowledge might prove fatal to his theory.

As it happens, some of the strongest evidence supporting evolutionary theory comes from genetics. A long series of discoveries, from Mendel to Watson and Crick to genomics, helps explain how evolution works.

Also, we now understand how mutation and the reshuffling of genes during sexual reproduction produce the heritable variation on which natural selection operates.Lifes Common Genetic Code All living cells use information coded in DNA and RNA to carry information from one generation to the next and to direct protein synthesis. Lifes Common Genetic Code This genetic code is nearly identical in almost all organisms, including bacteria, yeasts, plants, fungi, and animals. Lifes Common Genetic Code This compares a small portion of the DNA for the same gene in three animalsa mouse, a whale, and a chicken.

Lifes Common Genetic Code This similarity in genetic code is powerful evidence that all organisms evolved from common ancestors.

Similarities in MacromoleculesThere are similarities in RNA and DNA from one organism to anotherMany species have the red-blood-cell protein hemoglobinAmino acid sequence differs by one amino acid between a human and a gorillaAmino acid sequence differs by 67 amino acids between a human and a frogMolecular biology can be used to confirm evolutionary histories suggested by fossils and anatomy

Homologous Molecules In Darwins day, biologists could only study similarities and differences in structures they could see. But physical body structures cant be used to compare mice with yeasts or bacteria.

Today, we know that homology is not limited to physical structures.

Homologous proteins share extensive structural and chemical similarities.

One homologous protein is cytochrome c, which functions in cellular respiration. Remarkably similar versions of cytochrome c are found in almost all living cells, from cells in bakers yeast to cells in humans.Homologous Molecules Genes can be homologous, too. One example is a set of genes that determine the identities of body parts.

Know as Hox genes, they help to determine the head to tail axis in embryonic development.

In vertebrates, sets of homologous Hox genes direct the growth of front and hind limbs.

Small changes in these genes can produce dramatic changes in the structures they control. Homologous Molecules Relatively minor changes in an organisms genome can produce major changes in an organisms structure and the structure of its descendants.

At least some homologous Hox genes are found in almost all multicellular animals, from fruit flies to humans.

For example, bacteria that live in a hot spring are very different from animals, yet many of their genes, and therefore the proteins coded by those genes, are similar to those of animals.

Such profound biochemical similarities are best explained by Darwins conclusion: Living organisms evolved through descent with modification from a common ancestor.Testing Natural SelectionWhat does recent research on the Galpagos finches show about natural selection?

The Grants have documented that natural selection takes place in wild finch populations frequently, and sometimes rapidly.

The Grants work shows that variation within a species increases the likelihood of the species adapting to and surviving environmental change.

Testing Natural SelectionOne way to gather evidence for evolutionary change is to observe natural selection in action.

One of the best examples of natural selection in action comes from observations of animals living in their natural environmentthe Galpagos finches.A Testable Hypothesis Darwin hypothesized that the Galpagos finches he observed had descended from a common ancestor.

He noted that several finch species have beaks of very different sizes and shapes. Each species uses its beak like a specialized tool to pick up and handle its food. Different types of foods are most easily handled with beaks of different sizes and shapes.

Darwin proposed that natural selection had shaped the beaks of different bird populations as they became adapted to eat different foods. No one thought there was a way to test this hypothesis until Peter and Rosemary Grant of Princeton University came along.A Testable Hypothesis

Evaluating Evolutionary Theory Today, evolutionary theorywhich includes natural selectionoffers insights that are vital to all branches of biology, from research on infectious disease to ecology.

Thats why evolution is often called the grand unifying theory of the life sciences.Evaluating Evolutionary Theory Like any scientific theory, evolutionary theory is constantly reviewed as new data are gathered.

Researchers still debate important questions, such as precisely how new species arise and why species become extinct.

There is also significant uncertainty about exactly how life began.

However, any questions that remain are about how evolution worksnot whether evolution occurs. To scientists, evolution is the key to understanding the natural world. How Do We Know That Evolution Has Occurred?1.The fossil record of change in earlier species 2. The chemical and anatomical similarities of related life forms 3. The geographic distribution of related species 4. The recorded genetic changes in living organisms over many generationsEvidenceEvolution is the scientific explanation that accounts for the diversity of life on EarthEvolution theory is based on a collection of scientific facts, observations, and hypothesesScientific theory is a well-supported testable explanation of phenomena that have occurred in nature

veevolce .movEvidence of Evolution Homologous body structuresSimilarities in embryologySimilarities in genetic make-upGeographic distribution of living speciesThe fossil record

Geographic Distribution of Living Species In response to the environment, different unrelated species have evolved to look similar to each other