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8/13/2019 BIO311Dlecture5
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Announcements
Discussion sections start this week. You willtake a short pre-concept test-bring a pencil (itwill not influence your grade in the course).
First Faculty Fellows Dining with Students lunch
tomorrow, Tuesday, Sept. 10, Noon, Jester CityLimits.
Reminder: Lecture attendance is expected andto your benefit. I will take attendanceperiodically, as I did last Friday, and there willbe graded lecture activities throughout thesemester.
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The Origin of Species
A new era of biology began in 1859 whenCharles Darwin published The Origin ofSpecies.
The Origin of Speciesfocused biologistsattention on the great diversity of organisms.
And in fact, it is this knowledge of diversity thatlead to the development of evolutionary theory
and our understanding of the processes ofevolution today.
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The Origin of Species
The book that shook the worldOne long argument for evolution
20 years in the making
http://darwin-online.org.uk/
http://darwin-online.org.uk/http://darwin-online.org.uk/http://darwin-online.org.uk/http://darwin-online.org.uk/8/13/2019 BIO311Dlecture5
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Darwin noted that current species are
descendants of ancestral species. Evolution can be defined by Darwins
phrase descent with modification.
Evolution can be viewed as both apattern and a process.
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Figure 22.2
1809
1798
1812
1795
1830
17901809 183136
1844
18591870
Lamarck publishes hishypothesis of evolution.
Malthus publishes
Essay on the Principleof Population.
Hutton proposeshis principle of
gradualism.
Charles Darwinis born.
Darwin travels aroundthe world on HMS
Beagle.
The Galpagos Islands
Darwin writes hisessay on descentwith modification.
On the Origin ofSpeciesis published.
While studying species inthe Malay Archipelago,Wallace (shown in 1848)sends Darwin his hypothesisof natural selection.
1858Cuvier publishes his extensivestudies of vertebrate fossils.
Lyell publishesPrinciples of Geology.
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Afred Russel Wallace (1823-1913)
1858Wallace sent letterto Darwin describing histheory of evolution
Naturalist & collector
Explored Amazon/SouthPacific
Independently developednotion of natural selection
and species change
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Chapter 1: Variation under domestication
Domestic species are variable
This variability is heritable
Small changes accumulate to make bigdifferences
Breeders have made changes > than differencesbetween species
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Chapter 2: Variation in nature
Things are the same way in nature
Population vs. typological approach
Variation is a natural state
Variation makes taxonomy difficult
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The struggle for existence & the tendency for
population to increase
Competition and predation EVERYWHERE,variation can be related to success
Darwin calculated that 1 pair of elephants (veryslow breeders) will leave 15 million descendentsin 500 years
Thus, most individuals do not survive(or the planet would be overrun with elephants)
Chapter 3: The struggle for existence
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Chapter 4: Natural selection
Can we doubtthat individuals havingany advantage, however slight, over others,would have the best chance surviving and of
procreating their kind? On the other hand,we may feel sure that any variation in theleast degree injurious would be rigidlydestroyed. This preservation of favorable
variations and the rejection of injuriousvariations, I call Natural Selection.
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Chapter 4: Natural selection
Discusses the potential strength of naturalselection
Discusses the possibility of sexual selection
Variation, inheritance, and the struggle forexistence inevitably lead to evolution
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Some doubt about the permanence of
species preceded Darwins ideas
Descent with modification by naturalselection explains the adaptations of
organisms and the unity and diversity oflife
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Darwins Focus on Adaptation
In reassessing his observations, Darwinperceived adaptation to the environmentand the origin of new species as closely
related processes. From studies made years after Darwins
voyage, biologists have concluded that
this is what happened to the Galpagosfinches.
Fi 22 6
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Figure 22.6
(a) Cactus-eater (b) Insect-eater
(c) Seed-eater
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In 1844, Darwin wrote an essay on naturalselection as the mechanism of descent with
modification,but did not introduce his theorypublicly.
Natural selection is a process in which individualswith favorable inherited traits are more likely to
survive and reproduce. In June 1858, Darwin received a manuscript from
Alfred Russell Wallace, who had developed atheory of natural selection similar to Darwins.
Darwin quickly finished The Origin of Speciesandpublished it the next year.
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The Origin of Species
Darwin explained three broadobservations:
The unity of life
The diversity of life The match between organisms and their
environment
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Descent with Modification
Darwin never used the word evolutionin thefirst edition of The Origin of Species.
The phrase descent with modificationsummarized Darwins perception of the unity
of life.
The phrase refers to the view that allorganisms are related through descent from
an ancestor that lived in the remote past.
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In the Darwinian view, the history oflife is like a tree with branchesrepresenting lifes diversity.
Darwins theory meshed well with thehierarchy of Linnaeus.
Figure 22 7
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Figure 22.7
Figure 22 8
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Figure 22.8Hyracoidea(Hyraxes)
Sirenia(Manateesand relatives)
Deinotherium
Mammut
Platybelodon
Stegodon
Mammuthus
Elephas maximus(Asia)
Loxodonta africana(Africa)
Loxodonta cyclotis(Africa)
Moeritherium
Barytherium
60
Millions of years ago
34 24 5.521040
Years ago
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Recommended books
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For next lecture
Read Chap. 23 and answer:
- What is a gene pool?
- Define a biological population.
- What is Hardy Weinberg equilibrium?