BIO311Dlecture5

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    Announcements

    Discussion sections start this week. You willtake a short pre-concept test-bring a pencil (itwill not influence your grade in the course).

    First Faculty Fellows Dining with Students lunch

    tomorrow, Tuesday, Sept. 10, Noon, Jester CityLimits.

    Reminder: Lecture attendance is expected andto your benefit. I will take attendanceperiodically, as I did last Friday, and there willbe graded lecture activities throughout thesemester.

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    The Origin of Species

    A new era of biology began in 1859 whenCharles Darwin published The Origin ofSpecies.

    The Origin of Speciesfocused biologistsattention on the great diversity of organisms.

    And in fact, it is this knowledge of diversity thatlead to the development of evolutionary theory

    and our understanding of the processes ofevolution today.

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    The Origin of Species

    The book that shook the worldOne long argument for evolution

    20 years in the making

    http://darwin-online.org.uk/

    http://darwin-online.org.uk/http://darwin-online.org.uk/http://darwin-online.org.uk/http://darwin-online.org.uk/
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    Darwin noted that current species are

    descendants of ancestral species. Evolution can be defined by Darwins

    phrase descent with modification.

    Evolution can be viewed as both apattern and a process.

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    Figure 22.2

    1809

    1798

    1812

    1795

    1830

    17901809 183136

    1844

    18591870

    Lamarck publishes hishypothesis of evolution.

    Malthus publishes

    Essay on the Principleof Population.

    Hutton proposeshis principle of

    gradualism.

    Charles Darwinis born.

    Darwin travels aroundthe world on HMS

    Beagle.

    The Galpagos Islands

    Darwin writes hisessay on descentwith modification.

    On the Origin ofSpeciesis published.

    While studying species inthe Malay Archipelago,Wallace (shown in 1848)sends Darwin his hypothesisof natural selection.

    1858Cuvier publishes his extensivestudies of vertebrate fossils.

    Lyell publishesPrinciples of Geology.

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    Afred Russel Wallace (1823-1913)

    1858Wallace sent letterto Darwin describing histheory of evolution

    Naturalist & collector

    Explored Amazon/SouthPacific

    Independently developednotion of natural selection

    and species change

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    Chapter 1: Variation under domestication

    Domestic species are variable

    This variability is heritable

    Small changes accumulate to make bigdifferences

    Breeders have made changes > than differencesbetween species

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    Chapter 2: Variation in nature

    Things are the same way in nature

    Population vs. typological approach

    Variation is a natural state

    Variation makes taxonomy difficult

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    The struggle for existence & the tendency for

    population to increase

    Competition and predation EVERYWHERE,variation can be related to success

    Darwin calculated that 1 pair of elephants (veryslow breeders) will leave 15 million descendentsin 500 years

    Thus, most individuals do not survive(or the planet would be overrun with elephants)

    Chapter 3: The struggle for existence

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    Chapter 4: Natural selection

    Can we doubtthat individuals havingany advantage, however slight, over others,would have the best chance surviving and of

    procreating their kind? On the other hand,we may feel sure that any variation in theleast degree injurious would be rigidlydestroyed. This preservation of favorable

    variations and the rejection of injuriousvariations, I call Natural Selection.

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    Chapter 4: Natural selection

    Discusses the potential strength of naturalselection

    Discusses the possibility of sexual selection

    Variation, inheritance, and the struggle forexistence inevitably lead to evolution

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    Some doubt about the permanence of

    species preceded Darwins ideas

    Descent with modification by naturalselection explains the adaptations of

    organisms and the unity and diversity oflife

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    Darwins Focus on Adaptation

    In reassessing his observations, Darwinperceived adaptation to the environmentand the origin of new species as closely

    related processes. From studies made years after Darwins

    voyage, biologists have concluded that

    this is what happened to the Galpagosfinches.

    Fi 22 6

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    Figure 22.6

    (a) Cactus-eater (b) Insect-eater

    (c) Seed-eater

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    In 1844, Darwin wrote an essay on naturalselection as the mechanism of descent with

    modification,but did not introduce his theorypublicly.

    Natural selection is a process in which individualswith favorable inherited traits are more likely to

    survive and reproduce. In June 1858, Darwin received a manuscript from

    Alfred Russell Wallace, who had developed atheory of natural selection similar to Darwins.

    Darwin quickly finished The Origin of Speciesandpublished it the next year.

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    The Origin of Species

    Darwin explained three broadobservations:

    The unity of life

    The diversity of life The match between organisms and their

    environment

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    Descent with Modification

    Darwin never used the word evolutionin thefirst edition of The Origin of Species.

    The phrase descent with modificationsummarized Darwins perception of the unity

    of life.

    The phrase refers to the view that allorganisms are related through descent from

    an ancestor that lived in the remote past.

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    In the Darwinian view, the history oflife is like a tree with branchesrepresenting lifes diversity.

    Darwins theory meshed well with thehierarchy of Linnaeus.

    Figure 22 7

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    Figure 22.7

    Figure 22 8

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    Figure 22.8Hyracoidea(Hyraxes)

    Sirenia(Manateesand relatives)

    Deinotherium

    Mammut

    Platybelodon

    Stegodon

    Mammuthus

    Elephas maximus(Asia)

    Loxodonta africana(Africa)

    Loxodonta cyclotis(Africa)

    Moeritherium

    Barytherium

    60

    Millions of years ago

    34 24 5.521040

    Years ago

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    Recommended books

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    For next lecture

    Read Chap. 23 and answer:

    - What is a gene pool?

    - Define a biological population.

    - What is Hardy Weinberg equilibrium?