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Benthic Invertebrates
and Demersal Fishes
at an Interim Dredge-Disposal Site off Wiiiapa BayWashington
by David R Miller
RobertL Ernmett and
RobertJ McConnell
Aprii 1988
BENTHIC INVERTEBRATES AND DEMERSAL FISHES AT AN
INTERIM DREDGE-DISPOSAL SITE OFF WILLAPA BAY WASHINGTON
by
David R Miller
Robert L Enunett
At and Robert J McConnell
Final Report of Research
funded by
Environmental Protection Agency Region 10
1200 Sixth Avenue
^ Seattle Washington 98101 Contract DW 13931463-01-0
and
Coastal Zone and Estuarlne Studies Division
Northwest and Alaska Fisheries Center
National Marine Fisheries Service
National Oceanic and Atmospheric Administration 2725 Montlake Boulevard East
Seattle Washington 98112
April 1988
CONTENTS
Page
INTRODUCTION 1
METHODS 3
Fish and Vaglle Epifauna 3
Benthic Invertebrates and Sediments 5
Data Analysis 5
Fish and Vaglle Epifauna 5
Benthic Invertebrates 3
Sediments ^
RESULTS 9
Fish and Vaglle Epifauna 9
Benthic Invertebrates 9
Sediments 13
DISCUSSION 13
Fish and Vaglle Epifauna 13
Benthic Invertebrates 13
Sediments 1^
SUMMARY 16
ACKNOWLEDGMENTS 18
LITERATURE CITED 19
APPENDIX 21
INTRODUCTION
The Environmental Protection Agency (EPA) and the U Slaquo Army Corps of
Engineers (COE) are responsible for designating and managing ocean dredged
material disposal sites under the Marine Protection Research and Sanctuaries
Act (MPRSA) of 1972 (P L- 92-532 called the Ocean Dumping Act) The MPRSA
was passed in recognition that the disposal of wastes into ocean waters could
potentially adversely affect the environment and MPRSA requires that such
disposal occur only at designated sites The EPA requires that site
designations be based on 1) environmental studies at each site and 2) historical
information on the impact of dredged material disposal on areas similar to such
sites in physical chemical and biological characteristics
Within the National Oceanic and Atmospheric Administration the National
Marine Fisheries Service (NMFS) has the primary Federal responsibility for the
conservation management and development of the Nations fishery resources
under the Fishery Conservation and Management Act P L 9^-265 The NMFS
Habitat Conservation Policy recognizes that healthy marine estuarine and
anadromous fish habitats are essential to maintaining the Nation s fisheries
Under this policy the NMFS has been cooperating with the EPA and COE in
evaluating interim offshore dredge disposal sites in Oregon and Washington
In July 1985 the National Marine Fisheries Service (NMFS) entered into a
cooperative agreement with the EPA and the Seattle District COE to conduct an
ecological survey at a proposed interim dredge disposal site off Wiliapa Bay
Washington between 30 July and 1 August 1985- Previous studies at the proposed
site (10-15 fathoms) by Simons (198A) and Shapiro and Associates Inc (1985)
indicated high benthic invertebrate densities similar to those found off
Tlllamook Bayj Oregon (gt70000 organismsm^) at 10-20 fathoms by Emmett et albull
(1987) As a result EPA requested additional benthlc Invertebrate sampling
prior to final site designation However the Wlllapa Bay Interim disposal site
was subsequently moved to deeper water (10-2A fathoms) because of the Intensive
commercial crab fishery at the original site and It was at this new site that
our sampling took place
Benthlc Invertebrate communities are commonly monitored during
environmental assessment surveys Benthlc Invertebrates are useful Indicators
of environmental conditions because 1) they are In contact with bottom sediments
(dredge spoils) 2) being relatively Immobile they may be either transported or
burled by dredge spoils and 3) they provide a critical link In the food chain
(Morton 1977)
Bottom trawl surveys sampling fishes and crabs at dredge sites can be
Important In determining what species use an area for migration spawning
rearing or feeding However dredging effects on fishes and crabs are
difficult to quantify because of the animals mobility contagious distribution
and ability to avoid disturbed habitats
Combined benthlc Invertebrate fish and crab surveys are especially
useful In Identifying both the diversity and structure of the benthlc community
and In distinguishing possible food-web linkages between Infauna and vaglle
eplfauna (fishes and crabs)
METHODS
Fish and Vagile Epifauna
Sampling was done with an 8-m semiballoon shrimp trawl having an overall
mesh size of 381 mm (stretch measure) a 127-mm mesh liner was inserted in the
cod end to ensure retention of smaller fish and invertebrates Fishing width of
the trawl was estimated to be about 5m A trawl tow of 10 min duration timed
from when the trawl reached bottom to the beginning of retrieval was made on
each of three depth contours mdash 20-fathom (37-m) 22-fathom (40-m) and 2kshy
fathom (44-m) (Fig 1) Position fixes and distance traveled were determined by
Loran-C (Appendix Table A)
Fish and Dungeness crab (Cancer magister) from each trawl catch were
identified and a subsample of 50 individuals of each species was measured (mm)
[total and standard lengths for fishes and carapace width (across the carapace
anterior to the 10th anterolateral spines) for crabs] and weighed (g) The
remainder of each species was counted and weighed as a group (Appendix Table B)
Nautical miles
1 1
I (
(
^ iTrawl 1
33 I1 i 46deg 44shyr 7-1p--j^ i i
N
T W
i L A p
B
A y
Dredge disposal VbullC 46deg 40shysite
Trawl I
l I 22 TravTraw
i I (
22fm 10fm I
24fm J ^ 20fm
1
V 46deg 36shy
LEGENDBOX CORE )2024 22
W-1-1W-3-1 W-2-1 VW-1-2
W-3-3 W-2-3 W-1-3
W-3-4 W-2-4 W-1-4
W-3-5 W-2-5 W-1-5
W-3-2 W-2-2
f 124deg 10
I z Figure 1mdashTrawl (fish and vagile epifauna) and box corer (benthic
invertebrates) sampling locations off Willapa Bay Washington
Benthlc Invertebrates and Sediments
Sampling was completed at 15 stations along three transects at the 20-raquo
22- and 2A-fathom contours (Fig 1) A 0096-m2 Gray-OHara box corer (Fig 2)
(Pequegnat et al 1981) was used to collect six samples at each station Three
of these were individually sieved through a QS-mm diameter mesh screen and the
residue containing the macro-invertebrates was placed in jars and preserved in a
buffered 5^ formaldehyde solution containing rose bengal (a protein stain) Of
the remaining three samples one was subsampled for sediment and two reserved
for possible future analysis Sediment grain size was determined by sieving and
organic content by burning for 1 h at 600deg C (Appendix Table C) Sediment
analysis was done by the COE sediment laboratory at Troutdale Oregon
At the NMFS field station in Hammond Oregon the benthic organisms were picked
from the preserved samples identified to the lowest possible taxonomic level
(usually species) and counted All specimens were placed in vials containing
705s ethyl alcohol and stored at Hammond
Data Analysis
Fish and Vagile Epifauna
Various computer programs on the Northwest and Alaska Fisheries Centers
(NWAFC) Burroughs (B7800)l^ computer were used to analyze the sampling data
Loran-C readings were used to estimate distances fished Fish and crab
Reference to trade names does not imply Indorsement by National Marine
Fisheries Service NOAA
Weight tray
Figure 2mdashBox corer (Gray-OHara modification of a standard box corer Pequegnat et al 1981) used for benthic invertebrate survey in Willapa Bay For deeper penetration in hard sediments weights can be attached to the shelves located on opposite sides
densities [numberhectare(ha)] were estimated using distance fishedj fishing
width of the trawl and catch data Descriptive summaries of each sample
effort include the following (1) a species list for each trawl (2) the number
and weight of fishes and crabs captured (total and by species) (3) numberha
(total and by species) (A) weight (gha) (total and by species) and (5) four
community structure indices [diversity index (H and SDV) species richness
(SR) and species evenness (J)]
Diversity was calculated for each trawl effort using the Shannon-Weaver
Diversity Index (Shannon and Weaver 1963)
S
H = - I Pilog2Pi ial
where H =gt Shannon-Weaver Diversity Index Pi = Xan (Xa is the number of
individuals of a particular species in a sample and n is the total number of
all individuals in the sample) and S raquo number of species
A second diversity index the Simpson Diversity Value (Simpson 19A9) was
also calculated for each trawl sample
S
SDV = 1 - H Pi2 i-1
Diversity values are sensitive to two components the number of species in
a sample (species richness) and the distribution of individuals among species
(evenness) (Lloyd and Ghelardi 196A) Species richness (SR) was estimated using
Margalefs formula (Margalef 1958)
SR = (S-1) ln(N)
where S = number of species and N = total number of individuals at the station
Species evenness was described using the Evenness Index (Pielou 1966)
J= H log2S
where J=laquo Evenness Indexgt H= Shannon-Weaver Diversity Index and S = number of
species-
Benthic Invertebrates
Benthic data were analyzed with the aid of a program on the NWAFC B7800
computer The three samples collected at each station were treated as
replicates enabling calculation of a mean numberm^ and a standard deviation
for each species- The program also calculated the number of species mean
number of invertebratessample and standard deviation and mean number
of invertebratesm^ and standard deviation The community structure indices
used in the trawling analyses were also calculated for the benthic
invertebrate data- Cluster analysis using the Bray-Curtis dissimilarity
index with a group averaging fusion strategy (Clifford and Stephenson 1975)
was used to identify station groupings that had similar species and densities-
A 0-5 dissimilarity value was considered a significant difference between
groups- The mean numberm^ (untransformed) for each species per station was
used in this analysis- Species which had mean densities lt10m^ were excluded
from the analysis to reduce the effect of rare species-
Sediments
Mean grain size (phi) percent silt-clay and percent organic matter were
calculated for each station (Appendix Table C) Within-area variability
tended to be low so no statistical analysis was performedshy
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
BENTHIC INVERTEBRATES AND DEMERSAL FISHES AT AN
INTERIM DREDGE-DISPOSAL SITE OFF WILLAPA BAY WASHINGTON
by
David R Miller
Robert L Enunett
At and Robert J McConnell
Final Report of Research
funded by
Environmental Protection Agency Region 10
1200 Sixth Avenue
^ Seattle Washington 98101 Contract DW 13931463-01-0
and
Coastal Zone and Estuarlne Studies Division
Northwest and Alaska Fisheries Center
National Marine Fisheries Service
National Oceanic and Atmospheric Administration 2725 Montlake Boulevard East
Seattle Washington 98112
April 1988
CONTENTS
Page
INTRODUCTION 1
METHODS 3
Fish and Vaglle Epifauna 3
Benthic Invertebrates and Sediments 5
Data Analysis 5
Fish and Vaglle Epifauna 5
Benthic Invertebrates 3
Sediments ^
RESULTS 9
Fish and Vaglle Epifauna 9
Benthic Invertebrates 9
Sediments 13
DISCUSSION 13
Fish and Vaglle Epifauna 13
Benthic Invertebrates 13
Sediments 1^
SUMMARY 16
ACKNOWLEDGMENTS 18
LITERATURE CITED 19
APPENDIX 21
INTRODUCTION
The Environmental Protection Agency (EPA) and the U Slaquo Army Corps of
Engineers (COE) are responsible for designating and managing ocean dredged
material disposal sites under the Marine Protection Research and Sanctuaries
Act (MPRSA) of 1972 (P L- 92-532 called the Ocean Dumping Act) The MPRSA
was passed in recognition that the disposal of wastes into ocean waters could
potentially adversely affect the environment and MPRSA requires that such
disposal occur only at designated sites The EPA requires that site
designations be based on 1) environmental studies at each site and 2) historical
information on the impact of dredged material disposal on areas similar to such
sites in physical chemical and biological characteristics
Within the National Oceanic and Atmospheric Administration the National
Marine Fisheries Service (NMFS) has the primary Federal responsibility for the
conservation management and development of the Nations fishery resources
under the Fishery Conservation and Management Act P L 9^-265 The NMFS
Habitat Conservation Policy recognizes that healthy marine estuarine and
anadromous fish habitats are essential to maintaining the Nation s fisheries
Under this policy the NMFS has been cooperating with the EPA and COE in
evaluating interim offshore dredge disposal sites in Oregon and Washington
In July 1985 the National Marine Fisheries Service (NMFS) entered into a
cooperative agreement with the EPA and the Seattle District COE to conduct an
ecological survey at a proposed interim dredge disposal site off Wiliapa Bay
Washington between 30 July and 1 August 1985- Previous studies at the proposed
site (10-15 fathoms) by Simons (198A) and Shapiro and Associates Inc (1985)
indicated high benthic invertebrate densities similar to those found off
Tlllamook Bayj Oregon (gt70000 organismsm^) at 10-20 fathoms by Emmett et albull
(1987) As a result EPA requested additional benthlc Invertebrate sampling
prior to final site designation However the Wlllapa Bay Interim disposal site
was subsequently moved to deeper water (10-2A fathoms) because of the Intensive
commercial crab fishery at the original site and It was at this new site that
our sampling took place
Benthlc Invertebrate communities are commonly monitored during
environmental assessment surveys Benthlc Invertebrates are useful Indicators
of environmental conditions because 1) they are In contact with bottom sediments
(dredge spoils) 2) being relatively Immobile they may be either transported or
burled by dredge spoils and 3) they provide a critical link In the food chain
(Morton 1977)
Bottom trawl surveys sampling fishes and crabs at dredge sites can be
Important In determining what species use an area for migration spawning
rearing or feeding However dredging effects on fishes and crabs are
difficult to quantify because of the animals mobility contagious distribution
and ability to avoid disturbed habitats
Combined benthlc Invertebrate fish and crab surveys are especially
useful In Identifying both the diversity and structure of the benthlc community
and In distinguishing possible food-web linkages between Infauna and vaglle
eplfauna (fishes and crabs)
METHODS
Fish and Vagile Epifauna
Sampling was done with an 8-m semiballoon shrimp trawl having an overall
mesh size of 381 mm (stretch measure) a 127-mm mesh liner was inserted in the
cod end to ensure retention of smaller fish and invertebrates Fishing width of
the trawl was estimated to be about 5m A trawl tow of 10 min duration timed
from when the trawl reached bottom to the beginning of retrieval was made on
each of three depth contours mdash 20-fathom (37-m) 22-fathom (40-m) and 2kshy
fathom (44-m) (Fig 1) Position fixes and distance traveled were determined by
Loran-C (Appendix Table A)
Fish and Dungeness crab (Cancer magister) from each trawl catch were
identified and a subsample of 50 individuals of each species was measured (mm)
[total and standard lengths for fishes and carapace width (across the carapace
anterior to the 10th anterolateral spines) for crabs] and weighed (g) The
remainder of each species was counted and weighed as a group (Appendix Table B)
Nautical miles
1 1
I (
(
^ iTrawl 1
33 I1 i 46deg 44shyr 7-1p--j^ i i
N
T W
i L A p
B
A y
Dredge disposal VbullC 46deg 40shysite
Trawl I
l I 22 TravTraw
i I (
22fm 10fm I
24fm J ^ 20fm
1
V 46deg 36shy
LEGENDBOX CORE )2024 22
W-1-1W-3-1 W-2-1 VW-1-2
W-3-3 W-2-3 W-1-3
W-3-4 W-2-4 W-1-4
W-3-5 W-2-5 W-1-5
W-3-2 W-2-2
f 124deg 10
I z Figure 1mdashTrawl (fish and vagile epifauna) and box corer (benthic
invertebrates) sampling locations off Willapa Bay Washington
Benthlc Invertebrates and Sediments
Sampling was completed at 15 stations along three transects at the 20-raquo
22- and 2A-fathom contours (Fig 1) A 0096-m2 Gray-OHara box corer (Fig 2)
(Pequegnat et al 1981) was used to collect six samples at each station Three
of these were individually sieved through a QS-mm diameter mesh screen and the
residue containing the macro-invertebrates was placed in jars and preserved in a
buffered 5^ formaldehyde solution containing rose bengal (a protein stain) Of
the remaining three samples one was subsampled for sediment and two reserved
for possible future analysis Sediment grain size was determined by sieving and
organic content by burning for 1 h at 600deg C (Appendix Table C) Sediment
analysis was done by the COE sediment laboratory at Troutdale Oregon
At the NMFS field station in Hammond Oregon the benthic organisms were picked
from the preserved samples identified to the lowest possible taxonomic level
(usually species) and counted All specimens were placed in vials containing
705s ethyl alcohol and stored at Hammond
Data Analysis
Fish and Vagile Epifauna
Various computer programs on the Northwest and Alaska Fisheries Centers
(NWAFC) Burroughs (B7800)l^ computer were used to analyze the sampling data
Loran-C readings were used to estimate distances fished Fish and crab
Reference to trade names does not imply Indorsement by National Marine
Fisheries Service NOAA
Weight tray
Figure 2mdashBox corer (Gray-OHara modification of a standard box corer Pequegnat et al 1981) used for benthic invertebrate survey in Willapa Bay For deeper penetration in hard sediments weights can be attached to the shelves located on opposite sides
densities [numberhectare(ha)] were estimated using distance fishedj fishing
width of the trawl and catch data Descriptive summaries of each sample
effort include the following (1) a species list for each trawl (2) the number
and weight of fishes and crabs captured (total and by species) (3) numberha
(total and by species) (A) weight (gha) (total and by species) and (5) four
community structure indices [diversity index (H and SDV) species richness
(SR) and species evenness (J)]
Diversity was calculated for each trawl effort using the Shannon-Weaver
Diversity Index (Shannon and Weaver 1963)
S
H = - I Pilog2Pi ial
where H =gt Shannon-Weaver Diversity Index Pi = Xan (Xa is the number of
individuals of a particular species in a sample and n is the total number of
all individuals in the sample) and S raquo number of species
A second diversity index the Simpson Diversity Value (Simpson 19A9) was
also calculated for each trawl sample
S
SDV = 1 - H Pi2 i-1
Diversity values are sensitive to two components the number of species in
a sample (species richness) and the distribution of individuals among species
(evenness) (Lloyd and Ghelardi 196A) Species richness (SR) was estimated using
Margalefs formula (Margalef 1958)
SR = (S-1) ln(N)
where S = number of species and N = total number of individuals at the station
Species evenness was described using the Evenness Index (Pielou 1966)
J= H log2S
where J=laquo Evenness Indexgt H= Shannon-Weaver Diversity Index and S = number of
species-
Benthic Invertebrates
Benthic data were analyzed with the aid of a program on the NWAFC B7800
computer The three samples collected at each station were treated as
replicates enabling calculation of a mean numberm^ and a standard deviation
for each species- The program also calculated the number of species mean
number of invertebratessample and standard deviation and mean number
of invertebratesm^ and standard deviation The community structure indices
used in the trawling analyses were also calculated for the benthic
invertebrate data- Cluster analysis using the Bray-Curtis dissimilarity
index with a group averaging fusion strategy (Clifford and Stephenson 1975)
was used to identify station groupings that had similar species and densities-
A 0-5 dissimilarity value was considered a significant difference between
groups- The mean numberm^ (untransformed) for each species per station was
used in this analysis- Species which had mean densities lt10m^ were excluded
from the analysis to reduce the effect of rare species-
Sediments
Mean grain size (phi) percent silt-clay and percent organic matter were
calculated for each station (Appendix Table C) Within-area variability
tended to be low so no statistical analysis was performedshy
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
CONTENTS
Page
INTRODUCTION 1
METHODS 3
Fish and Vaglle Epifauna 3
Benthic Invertebrates and Sediments 5
Data Analysis 5
Fish and Vaglle Epifauna 5
Benthic Invertebrates 3
Sediments ^
RESULTS 9
Fish and Vaglle Epifauna 9
Benthic Invertebrates 9
Sediments 13
DISCUSSION 13
Fish and Vaglle Epifauna 13
Benthic Invertebrates 13
Sediments 1^
SUMMARY 16
ACKNOWLEDGMENTS 18
LITERATURE CITED 19
APPENDIX 21
INTRODUCTION
The Environmental Protection Agency (EPA) and the U Slaquo Army Corps of
Engineers (COE) are responsible for designating and managing ocean dredged
material disposal sites under the Marine Protection Research and Sanctuaries
Act (MPRSA) of 1972 (P L- 92-532 called the Ocean Dumping Act) The MPRSA
was passed in recognition that the disposal of wastes into ocean waters could
potentially adversely affect the environment and MPRSA requires that such
disposal occur only at designated sites The EPA requires that site
designations be based on 1) environmental studies at each site and 2) historical
information on the impact of dredged material disposal on areas similar to such
sites in physical chemical and biological characteristics
Within the National Oceanic and Atmospheric Administration the National
Marine Fisheries Service (NMFS) has the primary Federal responsibility for the
conservation management and development of the Nations fishery resources
under the Fishery Conservation and Management Act P L 9^-265 The NMFS
Habitat Conservation Policy recognizes that healthy marine estuarine and
anadromous fish habitats are essential to maintaining the Nation s fisheries
Under this policy the NMFS has been cooperating with the EPA and COE in
evaluating interim offshore dredge disposal sites in Oregon and Washington
In July 1985 the National Marine Fisheries Service (NMFS) entered into a
cooperative agreement with the EPA and the Seattle District COE to conduct an
ecological survey at a proposed interim dredge disposal site off Wiliapa Bay
Washington between 30 July and 1 August 1985- Previous studies at the proposed
site (10-15 fathoms) by Simons (198A) and Shapiro and Associates Inc (1985)
indicated high benthic invertebrate densities similar to those found off
Tlllamook Bayj Oregon (gt70000 organismsm^) at 10-20 fathoms by Emmett et albull
(1987) As a result EPA requested additional benthlc Invertebrate sampling
prior to final site designation However the Wlllapa Bay Interim disposal site
was subsequently moved to deeper water (10-2A fathoms) because of the Intensive
commercial crab fishery at the original site and It was at this new site that
our sampling took place
Benthlc Invertebrate communities are commonly monitored during
environmental assessment surveys Benthlc Invertebrates are useful Indicators
of environmental conditions because 1) they are In contact with bottom sediments
(dredge spoils) 2) being relatively Immobile they may be either transported or
burled by dredge spoils and 3) they provide a critical link In the food chain
(Morton 1977)
Bottom trawl surveys sampling fishes and crabs at dredge sites can be
Important In determining what species use an area for migration spawning
rearing or feeding However dredging effects on fishes and crabs are
difficult to quantify because of the animals mobility contagious distribution
and ability to avoid disturbed habitats
Combined benthlc Invertebrate fish and crab surveys are especially
useful In Identifying both the diversity and structure of the benthlc community
and In distinguishing possible food-web linkages between Infauna and vaglle
eplfauna (fishes and crabs)
METHODS
Fish and Vagile Epifauna
Sampling was done with an 8-m semiballoon shrimp trawl having an overall
mesh size of 381 mm (stretch measure) a 127-mm mesh liner was inserted in the
cod end to ensure retention of smaller fish and invertebrates Fishing width of
the trawl was estimated to be about 5m A trawl tow of 10 min duration timed
from when the trawl reached bottom to the beginning of retrieval was made on
each of three depth contours mdash 20-fathom (37-m) 22-fathom (40-m) and 2kshy
fathom (44-m) (Fig 1) Position fixes and distance traveled were determined by
Loran-C (Appendix Table A)
Fish and Dungeness crab (Cancer magister) from each trawl catch were
identified and a subsample of 50 individuals of each species was measured (mm)
[total and standard lengths for fishes and carapace width (across the carapace
anterior to the 10th anterolateral spines) for crabs] and weighed (g) The
remainder of each species was counted and weighed as a group (Appendix Table B)
Nautical miles
1 1
I (
(
^ iTrawl 1
33 I1 i 46deg 44shyr 7-1p--j^ i i
N
T W
i L A p
B
A y
Dredge disposal VbullC 46deg 40shysite
Trawl I
l I 22 TravTraw
i I (
22fm 10fm I
24fm J ^ 20fm
1
V 46deg 36shy
LEGENDBOX CORE )2024 22
W-1-1W-3-1 W-2-1 VW-1-2
W-3-3 W-2-3 W-1-3
W-3-4 W-2-4 W-1-4
W-3-5 W-2-5 W-1-5
W-3-2 W-2-2
f 124deg 10
I z Figure 1mdashTrawl (fish and vagile epifauna) and box corer (benthic
invertebrates) sampling locations off Willapa Bay Washington
Benthlc Invertebrates and Sediments
Sampling was completed at 15 stations along three transects at the 20-raquo
22- and 2A-fathom contours (Fig 1) A 0096-m2 Gray-OHara box corer (Fig 2)
(Pequegnat et al 1981) was used to collect six samples at each station Three
of these were individually sieved through a QS-mm diameter mesh screen and the
residue containing the macro-invertebrates was placed in jars and preserved in a
buffered 5^ formaldehyde solution containing rose bengal (a protein stain) Of
the remaining three samples one was subsampled for sediment and two reserved
for possible future analysis Sediment grain size was determined by sieving and
organic content by burning for 1 h at 600deg C (Appendix Table C) Sediment
analysis was done by the COE sediment laboratory at Troutdale Oregon
At the NMFS field station in Hammond Oregon the benthic organisms were picked
from the preserved samples identified to the lowest possible taxonomic level
(usually species) and counted All specimens were placed in vials containing
705s ethyl alcohol and stored at Hammond
Data Analysis
Fish and Vagile Epifauna
Various computer programs on the Northwest and Alaska Fisheries Centers
(NWAFC) Burroughs (B7800)l^ computer were used to analyze the sampling data
Loran-C readings were used to estimate distances fished Fish and crab
Reference to trade names does not imply Indorsement by National Marine
Fisheries Service NOAA
Weight tray
Figure 2mdashBox corer (Gray-OHara modification of a standard box corer Pequegnat et al 1981) used for benthic invertebrate survey in Willapa Bay For deeper penetration in hard sediments weights can be attached to the shelves located on opposite sides
densities [numberhectare(ha)] were estimated using distance fishedj fishing
width of the trawl and catch data Descriptive summaries of each sample
effort include the following (1) a species list for each trawl (2) the number
and weight of fishes and crabs captured (total and by species) (3) numberha
(total and by species) (A) weight (gha) (total and by species) and (5) four
community structure indices [diversity index (H and SDV) species richness
(SR) and species evenness (J)]
Diversity was calculated for each trawl effort using the Shannon-Weaver
Diversity Index (Shannon and Weaver 1963)
S
H = - I Pilog2Pi ial
where H =gt Shannon-Weaver Diversity Index Pi = Xan (Xa is the number of
individuals of a particular species in a sample and n is the total number of
all individuals in the sample) and S raquo number of species
A second diversity index the Simpson Diversity Value (Simpson 19A9) was
also calculated for each trawl sample
S
SDV = 1 - H Pi2 i-1
Diversity values are sensitive to two components the number of species in
a sample (species richness) and the distribution of individuals among species
(evenness) (Lloyd and Ghelardi 196A) Species richness (SR) was estimated using
Margalefs formula (Margalef 1958)
SR = (S-1) ln(N)
where S = number of species and N = total number of individuals at the station
Species evenness was described using the Evenness Index (Pielou 1966)
J= H log2S
where J=laquo Evenness Indexgt H= Shannon-Weaver Diversity Index and S = number of
species-
Benthic Invertebrates
Benthic data were analyzed with the aid of a program on the NWAFC B7800
computer The three samples collected at each station were treated as
replicates enabling calculation of a mean numberm^ and a standard deviation
for each species- The program also calculated the number of species mean
number of invertebratessample and standard deviation and mean number
of invertebratesm^ and standard deviation The community structure indices
used in the trawling analyses were also calculated for the benthic
invertebrate data- Cluster analysis using the Bray-Curtis dissimilarity
index with a group averaging fusion strategy (Clifford and Stephenson 1975)
was used to identify station groupings that had similar species and densities-
A 0-5 dissimilarity value was considered a significant difference between
groups- The mean numberm^ (untransformed) for each species per station was
used in this analysis- Species which had mean densities lt10m^ were excluded
from the analysis to reduce the effect of rare species-
Sediments
Mean grain size (phi) percent silt-clay and percent organic matter were
calculated for each station (Appendix Table C) Within-area variability
tended to be low so no statistical analysis was performedshy
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
INTRODUCTION
The Environmental Protection Agency (EPA) and the U Slaquo Army Corps of
Engineers (COE) are responsible for designating and managing ocean dredged
material disposal sites under the Marine Protection Research and Sanctuaries
Act (MPRSA) of 1972 (P L- 92-532 called the Ocean Dumping Act) The MPRSA
was passed in recognition that the disposal of wastes into ocean waters could
potentially adversely affect the environment and MPRSA requires that such
disposal occur only at designated sites The EPA requires that site
designations be based on 1) environmental studies at each site and 2) historical
information on the impact of dredged material disposal on areas similar to such
sites in physical chemical and biological characteristics
Within the National Oceanic and Atmospheric Administration the National
Marine Fisheries Service (NMFS) has the primary Federal responsibility for the
conservation management and development of the Nations fishery resources
under the Fishery Conservation and Management Act P L 9^-265 The NMFS
Habitat Conservation Policy recognizes that healthy marine estuarine and
anadromous fish habitats are essential to maintaining the Nation s fisheries
Under this policy the NMFS has been cooperating with the EPA and COE in
evaluating interim offshore dredge disposal sites in Oregon and Washington
In July 1985 the National Marine Fisheries Service (NMFS) entered into a
cooperative agreement with the EPA and the Seattle District COE to conduct an
ecological survey at a proposed interim dredge disposal site off Wiliapa Bay
Washington between 30 July and 1 August 1985- Previous studies at the proposed
site (10-15 fathoms) by Simons (198A) and Shapiro and Associates Inc (1985)
indicated high benthic invertebrate densities similar to those found off
Tlllamook Bayj Oregon (gt70000 organismsm^) at 10-20 fathoms by Emmett et albull
(1987) As a result EPA requested additional benthlc Invertebrate sampling
prior to final site designation However the Wlllapa Bay Interim disposal site
was subsequently moved to deeper water (10-2A fathoms) because of the Intensive
commercial crab fishery at the original site and It was at this new site that
our sampling took place
Benthlc Invertebrate communities are commonly monitored during
environmental assessment surveys Benthlc Invertebrates are useful Indicators
of environmental conditions because 1) they are In contact with bottom sediments
(dredge spoils) 2) being relatively Immobile they may be either transported or
burled by dredge spoils and 3) they provide a critical link In the food chain
(Morton 1977)
Bottom trawl surveys sampling fishes and crabs at dredge sites can be
Important In determining what species use an area for migration spawning
rearing or feeding However dredging effects on fishes and crabs are
difficult to quantify because of the animals mobility contagious distribution
and ability to avoid disturbed habitats
Combined benthlc Invertebrate fish and crab surveys are especially
useful In Identifying both the diversity and structure of the benthlc community
and In distinguishing possible food-web linkages between Infauna and vaglle
eplfauna (fishes and crabs)
METHODS
Fish and Vagile Epifauna
Sampling was done with an 8-m semiballoon shrimp trawl having an overall
mesh size of 381 mm (stretch measure) a 127-mm mesh liner was inserted in the
cod end to ensure retention of smaller fish and invertebrates Fishing width of
the trawl was estimated to be about 5m A trawl tow of 10 min duration timed
from when the trawl reached bottom to the beginning of retrieval was made on
each of three depth contours mdash 20-fathom (37-m) 22-fathom (40-m) and 2kshy
fathom (44-m) (Fig 1) Position fixes and distance traveled were determined by
Loran-C (Appendix Table A)
Fish and Dungeness crab (Cancer magister) from each trawl catch were
identified and a subsample of 50 individuals of each species was measured (mm)
[total and standard lengths for fishes and carapace width (across the carapace
anterior to the 10th anterolateral spines) for crabs] and weighed (g) The
remainder of each species was counted and weighed as a group (Appendix Table B)
Nautical miles
1 1
I (
(
^ iTrawl 1
33 I1 i 46deg 44shyr 7-1p--j^ i i
N
T W
i L A p
B
A y
Dredge disposal VbullC 46deg 40shysite
Trawl I
l I 22 TravTraw
i I (
22fm 10fm I
24fm J ^ 20fm
1
V 46deg 36shy
LEGENDBOX CORE )2024 22
W-1-1W-3-1 W-2-1 VW-1-2
W-3-3 W-2-3 W-1-3
W-3-4 W-2-4 W-1-4
W-3-5 W-2-5 W-1-5
W-3-2 W-2-2
f 124deg 10
I z Figure 1mdashTrawl (fish and vagile epifauna) and box corer (benthic
invertebrates) sampling locations off Willapa Bay Washington
Benthlc Invertebrates and Sediments
Sampling was completed at 15 stations along three transects at the 20-raquo
22- and 2A-fathom contours (Fig 1) A 0096-m2 Gray-OHara box corer (Fig 2)
(Pequegnat et al 1981) was used to collect six samples at each station Three
of these were individually sieved through a QS-mm diameter mesh screen and the
residue containing the macro-invertebrates was placed in jars and preserved in a
buffered 5^ formaldehyde solution containing rose bengal (a protein stain) Of
the remaining three samples one was subsampled for sediment and two reserved
for possible future analysis Sediment grain size was determined by sieving and
organic content by burning for 1 h at 600deg C (Appendix Table C) Sediment
analysis was done by the COE sediment laboratory at Troutdale Oregon
At the NMFS field station in Hammond Oregon the benthic organisms were picked
from the preserved samples identified to the lowest possible taxonomic level
(usually species) and counted All specimens were placed in vials containing
705s ethyl alcohol and stored at Hammond
Data Analysis
Fish and Vagile Epifauna
Various computer programs on the Northwest and Alaska Fisheries Centers
(NWAFC) Burroughs (B7800)l^ computer were used to analyze the sampling data
Loran-C readings were used to estimate distances fished Fish and crab
Reference to trade names does not imply Indorsement by National Marine
Fisheries Service NOAA
Weight tray
Figure 2mdashBox corer (Gray-OHara modification of a standard box corer Pequegnat et al 1981) used for benthic invertebrate survey in Willapa Bay For deeper penetration in hard sediments weights can be attached to the shelves located on opposite sides
densities [numberhectare(ha)] were estimated using distance fishedj fishing
width of the trawl and catch data Descriptive summaries of each sample
effort include the following (1) a species list for each trawl (2) the number
and weight of fishes and crabs captured (total and by species) (3) numberha
(total and by species) (A) weight (gha) (total and by species) and (5) four
community structure indices [diversity index (H and SDV) species richness
(SR) and species evenness (J)]
Diversity was calculated for each trawl effort using the Shannon-Weaver
Diversity Index (Shannon and Weaver 1963)
S
H = - I Pilog2Pi ial
where H =gt Shannon-Weaver Diversity Index Pi = Xan (Xa is the number of
individuals of a particular species in a sample and n is the total number of
all individuals in the sample) and S raquo number of species
A second diversity index the Simpson Diversity Value (Simpson 19A9) was
also calculated for each trawl sample
S
SDV = 1 - H Pi2 i-1
Diversity values are sensitive to two components the number of species in
a sample (species richness) and the distribution of individuals among species
(evenness) (Lloyd and Ghelardi 196A) Species richness (SR) was estimated using
Margalefs formula (Margalef 1958)
SR = (S-1) ln(N)
where S = number of species and N = total number of individuals at the station
Species evenness was described using the Evenness Index (Pielou 1966)
J= H log2S
where J=laquo Evenness Indexgt H= Shannon-Weaver Diversity Index and S = number of
species-
Benthic Invertebrates
Benthic data were analyzed with the aid of a program on the NWAFC B7800
computer The three samples collected at each station were treated as
replicates enabling calculation of a mean numberm^ and a standard deviation
for each species- The program also calculated the number of species mean
number of invertebratessample and standard deviation and mean number
of invertebratesm^ and standard deviation The community structure indices
used in the trawling analyses were also calculated for the benthic
invertebrate data- Cluster analysis using the Bray-Curtis dissimilarity
index with a group averaging fusion strategy (Clifford and Stephenson 1975)
was used to identify station groupings that had similar species and densities-
A 0-5 dissimilarity value was considered a significant difference between
groups- The mean numberm^ (untransformed) for each species per station was
used in this analysis- Species which had mean densities lt10m^ were excluded
from the analysis to reduce the effect of rare species-
Sediments
Mean grain size (phi) percent silt-clay and percent organic matter were
calculated for each station (Appendix Table C) Within-area variability
tended to be low so no statistical analysis was performedshy
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
Tlllamook Bayj Oregon (gt70000 organismsm^) at 10-20 fathoms by Emmett et albull
(1987) As a result EPA requested additional benthlc Invertebrate sampling
prior to final site designation However the Wlllapa Bay Interim disposal site
was subsequently moved to deeper water (10-2A fathoms) because of the Intensive
commercial crab fishery at the original site and It was at this new site that
our sampling took place
Benthlc Invertebrate communities are commonly monitored during
environmental assessment surveys Benthlc Invertebrates are useful Indicators
of environmental conditions because 1) they are In contact with bottom sediments
(dredge spoils) 2) being relatively Immobile they may be either transported or
burled by dredge spoils and 3) they provide a critical link In the food chain
(Morton 1977)
Bottom trawl surveys sampling fishes and crabs at dredge sites can be
Important In determining what species use an area for migration spawning
rearing or feeding However dredging effects on fishes and crabs are
difficult to quantify because of the animals mobility contagious distribution
and ability to avoid disturbed habitats
Combined benthlc Invertebrate fish and crab surveys are especially
useful In Identifying both the diversity and structure of the benthlc community
and In distinguishing possible food-web linkages between Infauna and vaglle
eplfauna (fishes and crabs)
METHODS
Fish and Vagile Epifauna
Sampling was done with an 8-m semiballoon shrimp trawl having an overall
mesh size of 381 mm (stretch measure) a 127-mm mesh liner was inserted in the
cod end to ensure retention of smaller fish and invertebrates Fishing width of
the trawl was estimated to be about 5m A trawl tow of 10 min duration timed
from when the trawl reached bottom to the beginning of retrieval was made on
each of three depth contours mdash 20-fathom (37-m) 22-fathom (40-m) and 2kshy
fathom (44-m) (Fig 1) Position fixes and distance traveled were determined by
Loran-C (Appendix Table A)
Fish and Dungeness crab (Cancer magister) from each trawl catch were
identified and a subsample of 50 individuals of each species was measured (mm)
[total and standard lengths for fishes and carapace width (across the carapace
anterior to the 10th anterolateral spines) for crabs] and weighed (g) The
remainder of each species was counted and weighed as a group (Appendix Table B)
Nautical miles
1 1
I (
(
^ iTrawl 1
33 I1 i 46deg 44shyr 7-1p--j^ i i
N
T W
i L A p
B
A y
Dredge disposal VbullC 46deg 40shysite
Trawl I
l I 22 TravTraw
i I (
22fm 10fm I
24fm J ^ 20fm
1
V 46deg 36shy
LEGENDBOX CORE )2024 22
W-1-1W-3-1 W-2-1 VW-1-2
W-3-3 W-2-3 W-1-3
W-3-4 W-2-4 W-1-4
W-3-5 W-2-5 W-1-5
W-3-2 W-2-2
f 124deg 10
I z Figure 1mdashTrawl (fish and vagile epifauna) and box corer (benthic
invertebrates) sampling locations off Willapa Bay Washington
Benthlc Invertebrates and Sediments
Sampling was completed at 15 stations along three transects at the 20-raquo
22- and 2A-fathom contours (Fig 1) A 0096-m2 Gray-OHara box corer (Fig 2)
(Pequegnat et al 1981) was used to collect six samples at each station Three
of these were individually sieved through a QS-mm diameter mesh screen and the
residue containing the macro-invertebrates was placed in jars and preserved in a
buffered 5^ formaldehyde solution containing rose bengal (a protein stain) Of
the remaining three samples one was subsampled for sediment and two reserved
for possible future analysis Sediment grain size was determined by sieving and
organic content by burning for 1 h at 600deg C (Appendix Table C) Sediment
analysis was done by the COE sediment laboratory at Troutdale Oregon
At the NMFS field station in Hammond Oregon the benthic organisms were picked
from the preserved samples identified to the lowest possible taxonomic level
(usually species) and counted All specimens were placed in vials containing
705s ethyl alcohol and stored at Hammond
Data Analysis
Fish and Vagile Epifauna
Various computer programs on the Northwest and Alaska Fisheries Centers
(NWAFC) Burroughs (B7800)l^ computer were used to analyze the sampling data
Loran-C readings were used to estimate distances fished Fish and crab
Reference to trade names does not imply Indorsement by National Marine
Fisheries Service NOAA
Weight tray
Figure 2mdashBox corer (Gray-OHara modification of a standard box corer Pequegnat et al 1981) used for benthic invertebrate survey in Willapa Bay For deeper penetration in hard sediments weights can be attached to the shelves located on opposite sides
densities [numberhectare(ha)] were estimated using distance fishedj fishing
width of the trawl and catch data Descriptive summaries of each sample
effort include the following (1) a species list for each trawl (2) the number
and weight of fishes and crabs captured (total and by species) (3) numberha
(total and by species) (A) weight (gha) (total and by species) and (5) four
community structure indices [diversity index (H and SDV) species richness
(SR) and species evenness (J)]
Diversity was calculated for each trawl effort using the Shannon-Weaver
Diversity Index (Shannon and Weaver 1963)
S
H = - I Pilog2Pi ial
where H =gt Shannon-Weaver Diversity Index Pi = Xan (Xa is the number of
individuals of a particular species in a sample and n is the total number of
all individuals in the sample) and S raquo number of species
A second diversity index the Simpson Diversity Value (Simpson 19A9) was
also calculated for each trawl sample
S
SDV = 1 - H Pi2 i-1
Diversity values are sensitive to two components the number of species in
a sample (species richness) and the distribution of individuals among species
(evenness) (Lloyd and Ghelardi 196A) Species richness (SR) was estimated using
Margalefs formula (Margalef 1958)
SR = (S-1) ln(N)
where S = number of species and N = total number of individuals at the station
Species evenness was described using the Evenness Index (Pielou 1966)
J= H log2S
where J=laquo Evenness Indexgt H= Shannon-Weaver Diversity Index and S = number of
species-
Benthic Invertebrates
Benthic data were analyzed with the aid of a program on the NWAFC B7800
computer The three samples collected at each station were treated as
replicates enabling calculation of a mean numberm^ and a standard deviation
for each species- The program also calculated the number of species mean
number of invertebratessample and standard deviation and mean number
of invertebratesm^ and standard deviation The community structure indices
used in the trawling analyses were also calculated for the benthic
invertebrate data- Cluster analysis using the Bray-Curtis dissimilarity
index with a group averaging fusion strategy (Clifford and Stephenson 1975)
was used to identify station groupings that had similar species and densities-
A 0-5 dissimilarity value was considered a significant difference between
groups- The mean numberm^ (untransformed) for each species per station was
used in this analysis- Species which had mean densities lt10m^ were excluded
from the analysis to reduce the effect of rare species-
Sediments
Mean grain size (phi) percent silt-clay and percent organic matter were
calculated for each station (Appendix Table C) Within-area variability
tended to be low so no statistical analysis was performedshy
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
METHODS
Fish and Vagile Epifauna
Sampling was done with an 8-m semiballoon shrimp trawl having an overall
mesh size of 381 mm (stretch measure) a 127-mm mesh liner was inserted in the
cod end to ensure retention of smaller fish and invertebrates Fishing width of
the trawl was estimated to be about 5m A trawl tow of 10 min duration timed
from when the trawl reached bottom to the beginning of retrieval was made on
each of three depth contours mdash 20-fathom (37-m) 22-fathom (40-m) and 2kshy
fathom (44-m) (Fig 1) Position fixes and distance traveled were determined by
Loran-C (Appendix Table A)
Fish and Dungeness crab (Cancer magister) from each trawl catch were
identified and a subsample of 50 individuals of each species was measured (mm)
[total and standard lengths for fishes and carapace width (across the carapace
anterior to the 10th anterolateral spines) for crabs] and weighed (g) The
remainder of each species was counted and weighed as a group (Appendix Table B)
Nautical miles
1 1
I (
(
^ iTrawl 1
33 I1 i 46deg 44shyr 7-1p--j^ i i
N
T W
i L A p
B
A y
Dredge disposal VbullC 46deg 40shysite
Trawl I
l I 22 TravTraw
i I (
22fm 10fm I
24fm J ^ 20fm
1
V 46deg 36shy
LEGENDBOX CORE )2024 22
W-1-1W-3-1 W-2-1 VW-1-2
W-3-3 W-2-3 W-1-3
W-3-4 W-2-4 W-1-4
W-3-5 W-2-5 W-1-5
W-3-2 W-2-2
f 124deg 10
I z Figure 1mdashTrawl (fish and vagile epifauna) and box corer (benthic
invertebrates) sampling locations off Willapa Bay Washington
Benthlc Invertebrates and Sediments
Sampling was completed at 15 stations along three transects at the 20-raquo
22- and 2A-fathom contours (Fig 1) A 0096-m2 Gray-OHara box corer (Fig 2)
(Pequegnat et al 1981) was used to collect six samples at each station Three
of these were individually sieved through a QS-mm diameter mesh screen and the
residue containing the macro-invertebrates was placed in jars and preserved in a
buffered 5^ formaldehyde solution containing rose bengal (a protein stain) Of
the remaining three samples one was subsampled for sediment and two reserved
for possible future analysis Sediment grain size was determined by sieving and
organic content by burning for 1 h at 600deg C (Appendix Table C) Sediment
analysis was done by the COE sediment laboratory at Troutdale Oregon
At the NMFS field station in Hammond Oregon the benthic organisms were picked
from the preserved samples identified to the lowest possible taxonomic level
(usually species) and counted All specimens were placed in vials containing
705s ethyl alcohol and stored at Hammond
Data Analysis
Fish and Vagile Epifauna
Various computer programs on the Northwest and Alaska Fisheries Centers
(NWAFC) Burroughs (B7800)l^ computer were used to analyze the sampling data
Loran-C readings were used to estimate distances fished Fish and crab
Reference to trade names does not imply Indorsement by National Marine
Fisheries Service NOAA
Weight tray
Figure 2mdashBox corer (Gray-OHara modification of a standard box corer Pequegnat et al 1981) used for benthic invertebrate survey in Willapa Bay For deeper penetration in hard sediments weights can be attached to the shelves located on opposite sides
densities [numberhectare(ha)] were estimated using distance fishedj fishing
width of the trawl and catch data Descriptive summaries of each sample
effort include the following (1) a species list for each trawl (2) the number
and weight of fishes and crabs captured (total and by species) (3) numberha
(total and by species) (A) weight (gha) (total and by species) and (5) four
community structure indices [diversity index (H and SDV) species richness
(SR) and species evenness (J)]
Diversity was calculated for each trawl effort using the Shannon-Weaver
Diversity Index (Shannon and Weaver 1963)
S
H = - I Pilog2Pi ial
where H =gt Shannon-Weaver Diversity Index Pi = Xan (Xa is the number of
individuals of a particular species in a sample and n is the total number of
all individuals in the sample) and S raquo number of species
A second diversity index the Simpson Diversity Value (Simpson 19A9) was
also calculated for each trawl sample
S
SDV = 1 - H Pi2 i-1
Diversity values are sensitive to two components the number of species in
a sample (species richness) and the distribution of individuals among species
(evenness) (Lloyd and Ghelardi 196A) Species richness (SR) was estimated using
Margalefs formula (Margalef 1958)
SR = (S-1) ln(N)
where S = number of species and N = total number of individuals at the station
Species evenness was described using the Evenness Index (Pielou 1966)
J= H log2S
where J=laquo Evenness Indexgt H= Shannon-Weaver Diversity Index and S = number of
species-
Benthic Invertebrates
Benthic data were analyzed with the aid of a program on the NWAFC B7800
computer The three samples collected at each station were treated as
replicates enabling calculation of a mean numberm^ and a standard deviation
for each species- The program also calculated the number of species mean
number of invertebratessample and standard deviation and mean number
of invertebratesm^ and standard deviation The community structure indices
used in the trawling analyses were also calculated for the benthic
invertebrate data- Cluster analysis using the Bray-Curtis dissimilarity
index with a group averaging fusion strategy (Clifford and Stephenson 1975)
was used to identify station groupings that had similar species and densities-
A 0-5 dissimilarity value was considered a significant difference between
groups- The mean numberm^ (untransformed) for each species per station was
used in this analysis- Species which had mean densities lt10m^ were excluded
from the analysis to reduce the effect of rare species-
Sediments
Mean grain size (phi) percent silt-clay and percent organic matter were
calculated for each station (Appendix Table C) Within-area variability
tended to be low so no statistical analysis was performedshy
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
Nautical miles
1 1
I (
(
^ iTrawl 1
33 I1 i 46deg 44shyr 7-1p--j^ i i
N
T W
i L A p
B
A y
Dredge disposal VbullC 46deg 40shysite
Trawl I
l I 22 TravTraw
i I (
22fm 10fm I
24fm J ^ 20fm
1
V 46deg 36shy
LEGENDBOX CORE )2024 22
W-1-1W-3-1 W-2-1 VW-1-2
W-3-3 W-2-3 W-1-3
W-3-4 W-2-4 W-1-4
W-3-5 W-2-5 W-1-5
W-3-2 W-2-2
f 124deg 10
I z Figure 1mdashTrawl (fish and vagile epifauna) and box corer (benthic
invertebrates) sampling locations off Willapa Bay Washington
Benthlc Invertebrates and Sediments
Sampling was completed at 15 stations along three transects at the 20-raquo
22- and 2A-fathom contours (Fig 1) A 0096-m2 Gray-OHara box corer (Fig 2)
(Pequegnat et al 1981) was used to collect six samples at each station Three
of these were individually sieved through a QS-mm diameter mesh screen and the
residue containing the macro-invertebrates was placed in jars and preserved in a
buffered 5^ formaldehyde solution containing rose bengal (a protein stain) Of
the remaining three samples one was subsampled for sediment and two reserved
for possible future analysis Sediment grain size was determined by sieving and
organic content by burning for 1 h at 600deg C (Appendix Table C) Sediment
analysis was done by the COE sediment laboratory at Troutdale Oregon
At the NMFS field station in Hammond Oregon the benthic organisms were picked
from the preserved samples identified to the lowest possible taxonomic level
(usually species) and counted All specimens were placed in vials containing
705s ethyl alcohol and stored at Hammond
Data Analysis
Fish and Vagile Epifauna
Various computer programs on the Northwest and Alaska Fisheries Centers
(NWAFC) Burroughs (B7800)l^ computer were used to analyze the sampling data
Loran-C readings were used to estimate distances fished Fish and crab
Reference to trade names does not imply Indorsement by National Marine
Fisheries Service NOAA
Weight tray
Figure 2mdashBox corer (Gray-OHara modification of a standard box corer Pequegnat et al 1981) used for benthic invertebrate survey in Willapa Bay For deeper penetration in hard sediments weights can be attached to the shelves located on opposite sides
densities [numberhectare(ha)] were estimated using distance fishedj fishing
width of the trawl and catch data Descriptive summaries of each sample
effort include the following (1) a species list for each trawl (2) the number
and weight of fishes and crabs captured (total and by species) (3) numberha
(total and by species) (A) weight (gha) (total and by species) and (5) four
community structure indices [diversity index (H and SDV) species richness
(SR) and species evenness (J)]
Diversity was calculated for each trawl effort using the Shannon-Weaver
Diversity Index (Shannon and Weaver 1963)
S
H = - I Pilog2Pi ial
where H =gt Shannon-Weaver Diversity Index Pi = Xan (Xa is the number of
individuals of a particular species in a sample and n is the total number of
all individuals in the sample) and S raquo number of species
A second diversity index the Simpson Diversity Value (Simpson 19A9) was
also calculated for each trawl sample
S
SDV = 1 - H Pi2 i-1
Diversity values are sensitive to two components the number of species in
a sample (species richness) and the distribution of individuals among species
(evenness) (Lloyd and Ghelardi 196A) Species richness (SR) was estimated using
Margalefs formula (Margalef 1958)
SR = (S-1) ln(N)
where S = number of species and N = total number of individuals at the station
Species evenness was described using the Evenness Index (Pielou 1966)
J= H log2S
where J=laquo Evenness Indexgt H= Shannon-Weaver Diversity Index and S = number of
species-
Benthic Invertebrates
Benthic data were analyzed with the aid of a program on the NWAFC B7800
computer The three samples collected at each station were treated as
replicates enabling calculation of a mean numberm^ and a standard deviation
for each species- The program also calculated the number of species mean
number of invertebratessample and standard deviation and mean number
of invertebratesm^ and standard deviation The community structure indices
used in the trawling analyses were also calculated for the benthic
invertebrate data- Cluster analysis using the Bray-Curtis dissimilarity
index with a group averaging fusion strategy (Clifford and Stephenson 1975)
was used to identify station groupings that had similar species and densities-
A 0-5 dissimilarity value was considered a significant difference between
groups- The mean numberm^ (untransformed) for each species per station was
used in this analysis- Species which had mean densities lt10m^ were excluded
from the analysis to reduce the effect of rare species-
Sediments
Mean grain size (phi) percent silt-clay and percent organic matter were
calculated for each station (Appendix Table C) Within-area variability
tended to be low so no statistical analysis was performedshy
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
Benthlc Invertebrates and Sediments
Sampling was completed at 15 stations along three transects at the 20-raquo
22- and 2A-fathom contours (Fig 1) A 0096-m2 Gray-OHara box corer (Fig 2)
(Pequegnat et al 1981) was used to collect six samples at each station Three
of these were individually sieved through a QS-mm diameter mesh screen and the
residue containing the macro-invertebrates was placed in jars and preserved in a
buffered 5^ formaldehyde solution containing rose bengal (a protein stain) Of
the remaining three samples one was subsampled for sediment and two reserved
for possible future analysis Sediment grain size was determined by sieving and
organic content by burning for 1 h at 600deg C (Appendix Table C) Sediment
analysis was done by the COE sediment laboratory at Troutdale Oregon
At the NMFS field station in Hammond Oregon the benthic organisms were picked
from the preserved samples identified to the lowest possible taxonomic level
(usually species) and counted All specimens were placed in vials containing
705s ethyl alcohol and stored at Hammond
Data Analysis
Fish and Vagile Epifauna
Various computer programs on the Northwest and Alaska Fisheries Centers
(NWAFC) Burroughs (B7800)l^ computer were used to analyze the sampling data
Loran-C readings were used to estimate distances fished Fish and crab
Reference to trade names does not imply Indorsement by National Marine
Fisheries Service NOAA
Weight tray
Figure 2mdashBox corer (Gray-OHara modification of a standard box corer Pequegnat et al 1981) used for benthic invertebrate survey in Willapa Bay For deeper penetration in hard sediments weights can be attached to the shelves located on opposite sides
densities [numberhectare(ha)] were estimated using distance fishedj fishing
width of the trawl and catch data Descriptive summaries of each sample
effort include the following (1) a species list for each trawl (2) the number
and weight of fishes and crabs captured (total and by species) (3) numberha
(total and by species) (A) weight (gha) (total and by species) and (5) four
community structure indices [diversity index (H and SDV) species richness
(SR) and species evenness (J)]
Diversity was calculated for each trawl effort using the Shannon-Weaver
Diversity Index (Shannon and Weaver 1963)
S
H = - I Pilog2Pi ial
where H =gt Shannon-Weaver Diversity Index Pi = Xan (Xa is the number of
individuals of a particular species in a sample and n is the total number of
all individuals in the sample) and S raquo number of species
A second diversity index the Simpson Diversity Value (Simpson 19A9) was
also calculated for each trawl sample
S
SDV = 1 - H Pi2 i-1
Diversity values are sensitive to two components the number of species in
a sample (species richness) and the distribution of individuals among species
(evenness) (Lloyd and Ghelardi 196A) Species richness (SR) was estimated using
Margalefs formula (Margalef 1958)
SR = (S-1) ln(N)
where S = number of species and N = total number of individuals at the station
Species evenness was described using the Evenness Index (Pielou 1966)
J= H log2S
where J=laquo Evenness Indexgt H= Shannon-Weaver Diversity Index and S = number of
species-
Benthic Invertebrates
Benthic data were analyzed with the aid of a program on the NWAFC B7800
computer The three samples collected at each station were treated as
replicates enabling calculation of a mean numberm^ and a standard deviation
for each species- The program also calculated the number of species mean
number of invertebratessample and standard deviation and mean number
of invertebratesm^ and standard deviation The community structure indices
used in the trawling analyses were also calculated for the benthic
invertebrate data- Cluster analysis using the Bray-Curtis dissimilarity
index with a group averaging fusion strategy (Clifford and Stephenson 1975)
was used to identify station groupings that had similar species and densities-
A 0-5 dissimilarity value was considered a significant difference between
groups- The mean numberm^ (untransformed) for each species per station was
used in this analysis- Species which had mean densities lt10m^ were excluded
from the analysis to reduce the effect of rare species-
Sediments
Mean grain size (phi) percent silt-clay and percent organic matter were
calculated for each station (Appendix Table C) Within-area variability
tended to be low so no statistical analysis was performedshy
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
Weight tray
Figure 2mdashBox corer (Gray-OHara modification of a standard box corer Pequegnat et al 1981) used for benthic invertebrate survey in Willapa Bay For deeper penetration in hard sediments weights can be attached to the shelves located on opposite sides
densities [numberhectare(ha)] were estimated using distance fishedj fishing
width of the trawl and catch data Descriptive summaries of each sample
effort include the following (1) a species list for each trawl (2) the number
and weight of fishes and crabs captured (total and by species) (3) numberha
(total and by species) (A) weight (gha) (total and by species) and (5) four
community structure indices [diversity index (H and SDV) species richness
(SR) and species evenness (J)]
Diversity was calculated for each trawl effort using the Shannon-Weaver
Diversity Index (Shannon and Weaver 1963)
S
H = - I Pilog2Pi ial
where H =gt Shannon-Weaver Diversity Index Pi = Xan (Xa is the number of
individuals of a particular species in a sample and n is the total number of
all individuals in the sample) and S raquo number of species
A second diversity index the Simpson Diversity Value (Simpson 19A9) was
also calculated for each trawl sample
S
SDV = 1 - H Pi2 i-1
Diversity values are sensitive to two components the number of species in
a sample (species richness) and the distribution of individuals among species
(evenness) (Lloyd and Ghelardi 196A) Species richness (SR) was estimated using
Margalefs formula (Margalef 1958)
SR = (S-1) ln(N)
where S = number of species and N = total number of individuals at the station
Species evenness was described using the Evenness Index (Pielou 1966)
J= H log2S
where J=laquo Evenness Indexgt H= Shannon-Weaver Diversity Index and S = number of
species-
Benthic Invertebrates
Benthic data were analyzed with the aid of a program on the NWAFC B7800
computer The three samples collected at each station were treated as
replicates enabling calculation of a mean numberm^ and a standard deviation
for each species- The program also calculated the number of species mean
number of invertebratessample and standard deviation and mean number
of invertebratesm^ and standard deviation The community structure indices
used in the trawling analyses were also calculated for the benthic
invertebrate data- Cluster analysis using the Bray-Curtis dissimilarity
index with a group averaging fusion strategy (Clifford and Stephenson 1975)
was used to identify station groupings that had similar species and densities-
A 0-5 dissimilarity value was considered a significant difference between
groups- The mean numberm^ (untransformed) for each species per station was
used in this analysis- Species which had mean densities lt10m^ were excluded
from the analysis to reduce the effect of rare species-
Sediments
Mean grain size (phi) percent silt-clay and percent organic matter were
calculated for each station (Appendix Table C) Within-area variability
tended to be low so no statistical analysis was performedshy
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
densities [numberhectare(ha)] were estimated using distance fishedj fishing
width of the trawl and catch data Descriptive summaries of each sample
effort include the following (1) a species list for each trawl (2) the number
and weight of fishes and crabs captured (total and by species) (3) numberha
(total and by species) (A) weight (gha) (total and by species) and (5) four
community structure indices [diversity index (H and SDV) species richness
(SR) and species evenness (J)]
Diversity was calculated for each trawl effort using the Shannon-Weaver
Diversity Index (Shannon and Weaver 1963)
S
H = - I Pilog2Pi ial
where H =gt Shannon-Weaver Diversity Index Pi = Xan (Xa is the number of
individuals of a particular species in a sample and n is the total number of
all individuals in the sample) and S raquo number of species
A second diversity index the Simpson Diversity Value (Simpson 19A9) was
also calculated for each trawl sample
S
SDV = 1 - H Pi2 i-1
Diversity values are sensitive to two components the number of species in
a sample (species richness) and the distribution of individuals among species
(evenness) (Lloyd and Ghelardi 196A) Species richness (SR) was estimated using
Margalefs formula (Margalef 1958)
SR = (S-1) ln(N)
where S = number of species and N = total number of individuals at the station
Species evenness was described using the Evenness Index (Pielou 1966)
J= H log2S
where J=laquo Evenness Indexgt H= Shannon-Weaver Diversity Index and S = number of
species-
Benthic Invertebrates
Benthic data were analyzed with the aid of a program on the NWAFC B7800
computer The three samples collected at each station were treated as
replicates enabling calculation of a mean numberm^ and a standard deviation
for each species- The program also calculated the number of species mean
number of invertebratessample and standard deviation and mean number
of invertebratesm^ and standard deviation The community structure indices
used in the trawling analyses were also calculated for the benthic
invertebrate data- Cluster analysis using the Bray-Curtis dissimilarity
index with a group averaging fusion strategy (Clifford and Stephenson 1975)
was used to identify station groupings that had similar species and densities-
A 0-5 dissimilarity value was considered a significant difference between
groups- The mean numberm^ (untransformed) for each species per station was
used in this analysis- Species which had mean densities lt10m^ were excluded
from the analysis to reduce the effect of rare species-
Sediments
Mean grain size (phi) percent silt-clay and percent organic matter were
calculated for each station (Appendix Table C) Within-area variability
tended to be low so no statistical analysis was performedshy
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
Species evenness was described using the Evenness Index (Pielou 1966)
J= H log2S
where J=laquo Evenness Indexgt H= Shannon-Weaver Diversity Index and S = number of
species-
Benthic Invertebrates
Benthic data were analyzed with the aid of a program on the NWAFC B7800
computer The three samples collected at each station were treated as
replicates enabling calculation of a mean numberm^ and a standard deviation
for each species- The program also calculated the number of species mean
number of invertebratessample and standard deviation and mean number
of invertebratesm^ and standard deviation The community structure indices
used in the trawling analyses were also calculated for the benthic
invertebrate data- Cluster analysis using the Bray-Curtis dissimilarity
index with a group averaging fusion strategy (Clifford and Stephenson 1975)
was used to identify station groupings that had similar species and densities-
A 0-5 dissimilarity value was considered a significant difference between
groups- The mean numberm^ (untransformed) for each species per station was
used in this analysis- Species which had mean densities lt10m^ were excluded
from the analysis to reduce the effect of rare species-
Sediments
Mean grain size (phi) percent silt-clay and percent organic matter were
calculated for each station (Appendix Table C) Within-area variability
tended to be low so no statistical analysis was performedshy
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
RESULTS
Fish and Vagile Epifauna
The three trawl tows captured 10394 fish represented by 15 species
(Appendix Tables D and E) Pacific tomcod (Microgadus proximus) was the
dominant species captured at all three stations accounting for 95diams45regof the
total catch (Table 1) Densities (numberha) decreased dramatically as depth
increased with a 93956 decrease in densities between Stations W-1 and W-2 and
an 84656 decrease between W-2 and W-3 Conversely diversity (H and SDV)
increased as depth increased because of the reduced numbers of Pacific tomcod
and whitebait smelt (Allosmerus elongatus) at the deeper stations This
condition made the number of species in the sample nonproportional to the
number of individuals within the species
Benthic Invertebrates
A total of 229 taxa (Appendix Table F) with a mean density of 16113m2
were identified at the Willapa Bay site (Table 2) Highest invertebrate
densities were found along the 20-fathom contour (25885m^) densities
gradually decreased at the 22- (15440m2) 24-fathom transects (7014m2)
The highest invertebrate density was 35045m^ at Station W-1-2bull Mean taxa
numbers ranged from 80 to 85 and were similar for all transects Diversity
values (H and SDV) showed a mild inverse relationship to Invertebrate
densities with some higher diversity values associated with lower
invertebrate densities- The mildness of the relationship was due to the
domination of Owenia fusiformis (Table 3) a tube-dwelling polychaete worm at
all stations except W-3-4 (0_^ fusiformis 805m2 mdash Spiophanes bombyx
l847m2) and W-3-5 (0 fusiformis l622m2 mdash Spiophanes bombyx l545m2)
The highest density of 0- fusiformis was 20986m^ at station W-1-2
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
10
Table 1mdashCatch summaries (numberhectare and weighthectare) and diversity indices (by species) for trawl tows near Willapa Bay Washington 1 August 1985shy
Trawl W - 1 Trawl W - 2 Trawl W - 3
(depth 20 fathoms) (depth 22 fathoms) (depth 24 fathoms)
Species Nobullha gha Noha gha Noha gha
Big skate 13 84905
Longfin smelt 13 54 - - -
Whitebait smelt 867 7137 54 589 5 81
Plainfin
midshipman 1 41 - -
Pacific tomcod 29067 95838 1633 4418 200 535
Pacific staghorn sculpin 6 844 - - -
Warty poacher 6 60 7 88 -
Showy snailfish 13 210 - - -
Pacific sanddab - - 16 1838 16 2465
Speckled sanddab - - - - 16 124
Butter sole 130 6498 65 4454 22 1470
Slender sole - - - - 5 303
English sole 51 2156 36 1737 11 422
Sand sole 6 1873 7 1105 -
Dover sole 35 273 38 2112 11 297
Dungeness crab ^8 4610 mdash mdash mdash mdash
Totals 30258 204499 1856 16341 286 5697
Diversity Indices
H 030 082 169
SDV 008 022 050
SR 091 072 122
J 008 027 056
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
11
Table 2mdashGeneral description of the benthic invertebrate community sampled with a box corer off Willapa Bay Washington 30-31 July 1985 Grand totals were calculated by combining replicates from all stations
Station amp
depth (ft) No of
taxa
Mean no
m^ St dev
m^ H SDV SR J
W-1-1 121
W-1-2 123
W-1-3 123
W-l-A 120
W-1-5 120
70
98
75
85
71
25892
35045
23427
26174
18889
2601
11266
6883
15472
4890
299
241
328
289
281
076
036
081
071
067
774
1052
839
941
814
049
036
053
045
046
Mean total 80 25885
W-2-1 133
W-2-2 133
W-2-3 135
W-2-4 133
W-2-5 132
77
106
78
90
71
21993
24917
10670
11875
7747
6223
16111
1975
1039
1574
226
298
296
322
315
050
073
067
070
070
868
1183
959
1106
908
036
044
047
050
051
Mean total 85 15440
m
W-3-1 1U5
W-3-2 1A5
W-3-3 1A6
W-3-A 141
W-3-5 144
86
108
70
76
69
8816
11177
4910
4986
5181
3187
3769
1595
1617
1707
310
355
379
384
351
067
074
084
083
081
1097
1325
951
1032
931
048
053
062
061
057
Mean total 82 7014
Grand total 229 16113 11122 335 073 2045 043
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
12
Table 3 Summary of benthlc invertebrates collected near Wlllapa Baygt Washington 30-31 July 1985 Only dominant taxa from each category are shown
O
Taxon Total number Mean nom^
FOLTCHAETA
Owenia fusiformis 34A20 7968 Splophanes bombyx 3368 780 Spiophanes berkeleyorum 983 228 Onuphis sp indet 912 211 Onuphis iridescens 364 84 Chaetozone setosa 675 156 Leitoscoloplos pugettensis 599 139 Nephtys sp juv 459 106 Nephtys caecoides 264 61 Miscellaneous 3650 845
Total 45795 10600
CUMACEA
Diastylopsis dawsoni 7416 1717 Diastylopsis sp juv 4976 1152 Diastylopsis sp 687 159 Leucon subnasica 673 156 Miscellaneous 19 4
Total 13771 3188
AMPHIPODA
Photis sp juv 534 124
181 42Photis parvidons 107 25Photis brevipes 477 110Rhepoxynius sp
Rhepoxynius dabouis 271 63
Rhepoxynius abronius 202 47
Protomedeia sp juv 266 61 204 47Protomedeia prudens
Miscellaneous 973 225
Total 3215 744
OTHER
Bivalvia 4339 1004 Miscellaneous 2489 576
Total 6828 1581
Grand total 69609 16113
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
13
Cluster analysis of the data revealed two strong cluster groups (Fig- 3)shy
A high invertebrate density group (18000 - 35000m2) represented by the
20-fathom transect stations plus the two northern 22-fathom transect stations-
A moderate density group (5000 - 12000m2) was represented by the three
southern 22-fathom transect stations and all but one station of the 24-fathom
transect The latter station W-3-2 did not fit into any cluster group
Sediments
The Willapa Bay sediments were primarily fine sand mean grain size was
377 phi (Appendix Table C) Samples were moderately high in silt-clay (mean
value 23A) and low in organic matter (mean value lTs)
DISCUSSION
Fish and Vagile Epifauna
The 1985 Willapa Bay study was designed primarily to obtain additional
information on the benthie infaunal communities in the area of the proposed
interim disposal site Trawl surveys provided a species list and relative
abundances of larger organisms in the study area However to adequately
describe the fish and crab fauna replicate samples should be taken from each
depth contour to ensure statistical validity
Theoretically maximum diversity exists where there are many species
each represented by one Individual and minimum diversity exists when all
individuals belong to one species neither situation exists in a natural
community Although Trawl 1 had the highest number of species diversity was
low due to the unbalanced proportion of Pacific tomcod and whitebait smelt
(Table 1) Without more data It Is difficult to speculate why such great
differences occurred among stations- Since the greatest numbers in Trawl 1
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
Nautical miles
) 1
I N
I 46deg 44- ^
I l-~i-1^
1 i
Dredge disposal li site 46deg 40
22fm ( I
24fm 20fm
46deg 36
Density(numberm
5000-12000
18000-35000
124deg 10
Figure 3mdashLocations of benthic invertebrate cluster groups identified off Willapa Bay Washington
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
15
were due to Pacific tomcod which are epibenthic feeders it may be assumed
that they were near shore feeding on epibenthic prey
Benthic Invertebrates
The Willapa Bay benthic invertebrate community was high in both number of
taxa and density The dominant species was Owenia fusiformis The
distribution of this tube-dwelling worm was probably strongly affected by
sediment stability Oliver et al(1980) found tube-building polychaetes
abundant only in deeper water (gtA6 ft) where sediment disturbance due to wave
action is minimal Emmett et al(1987) in their September 198A survey off
Tillamook Bay found low densities of 0^^ fusiformis at 60 ft with highest
densities at 80 and 90 ft The Willapa Bay survey transects began at the 20shy
fathom depth contour and showed species and densities similar to the high
density samples off Tillamook Bay High densities of 0 fusiformis were also
found off Grays Harbor Washington (Pearson et al 1985) adjacent to and
north of Willapa Bay
Tillamook Bay Willapa Bay and Grays Harbor estuaries have extensive mud
flats with macroalgae and eelgrass Trawl tows at the Tillamook Bay and
Willapa Bay interim offshore disposal sites collected large amounts of
estuarine macroalgae Perhaps the export of these estuarine materials
enhances the nearshore benthic community The densities of benthic
invertebrates adjacent to these three estuaries are much higher than typically
found in the Oregon-Washington nearshore environment (Lie and Klsker 1970
Hancock et albull 198A Emmett et al 1987) They may represent unique marine
benthic communities linked to adjacent estuarine primary productionshy
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
16
Sediments
Sediments were typical of nearshore environments that are exposed to
waves and ocean swells (Fager 1968) Although sediments have a significant
effect on the distribution and abundance of benthic invertebrates (Gary 1965)
wave disturbance also plays an important role in determining the nearshore
benthic invertebrate community (Oliver et al 1980) Since invertebrate
densities at Willapa and Tillamook bays were much greater than those found at
other disposal sites along the Oregon coast and sediment structure was similar
(Emmett et al- 1987) sediment characteristics alone probably do not provide
enough information to predict either the benthic invertebrate community or
densities along the Oregon-Washington coast
SUMMARY
The Willapa Bay interim disposal site trawl-samples enabled a listing of
the larger benthic organisms and their relative abundances within the study
area However to adequately describe the fish and crab fauna replicate
samples should be taken from each depth contour to ensure statistical
validity
The benthic invertebrate community was high in both number of species and
densities of individuals The dominant species was the tube-dwelling worm
Owenia fusiformis Benthic invertebrate densities were much higher than
typically found in the Washington-Oregon nearshore environment
Sediments were typical of Washington-Oregon nearshore environments that
are exposed to waves and ocean swells Sediments were composed of clean fine
sands with moderate amounts of silt-clay and low amounts of organic matter
The density of benthic invertebrates off Willapa Bay resembled those off
the adjacent Grays Harbor Washington and off Tillamook Bay Oregon These
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5
17
three estuaries have extensive mud flats with macroalgae and eelgrasS- The
cause of the high invertebrate densities is unknown but the contribution of
primary production from the adjacent estuary is a probable factor The areal
extent of the high Invertebrate densities off Willapa Bay Grays Harbor and
Tillaraook Bay is not known and should be defined We recommend that these
areas not be used for the disposal of dredged materials until after more
intensive studies and then only if it can be shown that there are no severe
impacts to the estuarine and marine fisheries
This report does not constitute NMFSs formal comment under the Fish and
Wildlife Coordination Act or the National Environmental Policy Act
18 (laquoN
ACKNOWLEDGMENTS ^ ltbullraquo
This study was funded by the EPA Region 10) and the NMFS The Portland
District) COE) Troutdale) Oregon) sediment laboratory provided the sediment
analyses- Special thanks go to Travis Coley) NMFS) and to Howard Jones)
Marine Taxonomic Services) CorvalliS) Oregon
19
LITERATURE CITED
Gary A G Jr
1965 Preliminary studies on animal-sediment interrelationships off the central Oregon coast Mar Tech Soc Ocean Sci Ocean Engineering 1100-110
Cliffordraquo H T and W Stephenson 1975 An introduction to numerical classification Academic Press New
York 229 p
Emmett R L T C Coley G T McCabe and R J McConnell 1987 Demersal fishes and benthic invertebrates at four interim dredge disposal sites off the Oregon coast US Dept Commer Natl Oceanic Atmos Admin Natl Mar Fish Serv Northwest and Alaska Fish Cent Seattle WA 69 p plus Appendices (Report to US Army Corps of Engineers Contract DACW57-85-F-0210)
Fager E W
1968 A sand-bottom epifaunal community of invertebrates in shallow water Limnol Oceanogr 13448-A64
Hancock D R P 0 Nelson C K Sollitt and K J Williamson 1984 Coos Bay offshore disposal site investigation Interim Report Phase 1 February 1979-March 1980 61 p plus Appendices (Report to U S Army Corps of Engineers Portland District PO Box 2946 Portland OR 97208)
Lie U and D S Kisker
1970 Species composition and structure of benthic infauna communities off the coast of Washington J Fish Res Bd Canada 272273-2285
Lloyd M and R J Ghelardi
1964 A table for calculating the equitability component of species diversity J Anim Ecol 33217-225
Margalef R
1958 Information theory in ecology Gen Syst 336-71
Morton J W
1977 Ecological effects of dredging and dredge spoil disposal a literature review U-S Dept Int Fish Wildl Serv Tech Paper 94 33 p
Oliver J S p N Slattery L- W Hulberg and J- W Nybakken 1980 Relationships between wave disturbance and zonation of benthic
invertebrate communities along a subtidal high-energy beach in Monterey Bay California Fish Bull US 78437-454
20
Pearson W H D L- Woodruff P Wilkinson J S- Young and S L Kiessger
1985 Ocean surveys to select and characterize an ocean disposal site off Grays Harbor Washington 27 p plus Appendices (Draft Data Report to ^ US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 9812A)
Pequegnat W E L H Pequegnat B M James E A Kennedy R R Fay and A D Fredericks
my
1981 Procedural guide for designation surveys of ocean dredged material ^ disposal sites Tech Report EL-81-1 to Chief of Engineers U S Army Washington D C 20314 268 p plus Appendices
Pielou E L 1966 The measurement of diversity in different types of biological
collections J Theor Biol 13131-144
Shannon C E and W Weaver 1963 The mathematical theory of communication Univ Illinois Press
Urbana 117 p
Shapiro and Associates Inc 1985 Willapa Bay interim ocean dredge material disposal site designation
study 24 p plus Appendices (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124)
Simons D D 1984 Razor clam study Willapa Bay ocean disposal site Washington ^
Dept Fish 10 p plus Appendix (Report to US Army Corps of Engineers Seattle District PO Box C-3755 Seattle WA 98124 Contract DACW 67-84-1693)
Simpson E H 1949 Measurement of diversity Nature 163688 ^
21
APPENDIX
Data Tables
Appendix Table AmdashWlllapa Baygt Washington Interim ocean disposal site
sampling stations mdash Loran-C coordinates and depth 30-31 July and 1 August 1985
Station Coordinates Depth (ft)
20-fathom contour BENTHIC
W-1-1 119073 28044A 121
W-1-2 119115 280447 123
W-1-3 119147 280435 123
W-1-4 119200 280430 120
W-1-5 119255 280430 120
22-fathom contour BENTHIC
W-2-1 119100 280440 133
W-2-2 119150 280430 133
W-2-3 119175 280425 135
W-2-4 119220 280415 133
W-2-5 119265 280415 132
24-fathom contour BENTHIC
W-3-1 119113 280424 145
W-3-2 119160 280421 145
W-3-3 119203 280402 146
W-3-A 119250 280400 141
W-3-5 119290 280395 144
20-fathom contour TRAWL
W-1 119118 280447 - 119104 280446 123 - 125
22-fathom contour TRAWL
W-2 119140 280422 - 119113 280424 135 - 139
24-fathom contour TRAWL
W-3 119104 280414 - 119111 280415 147 - 146
A-1
Appendix Table B---Individual lengths and weights for fish and Dungeness crab collected with an 8-m trawl off Willapa Bay Washington 1 August 1985 Mass data are included where there were more than 50 individuals of each species
Trawl W-1
Gear 8-ra trawl
Date 1 August 1985 Depth 123 ft
Distance 2067 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Big skate 262 262 166
lAlO 0 26000
210 210 77
377 377 502
Longfin smelt 103 84 7
45 39 1
105 90 8
48 40 1
Whitebait smelt 125 106 10
110 93 10
111 94 7
94 79 4
121 107 11
122 105 9
108 90 6
115 95 8
121 102 10
119 100 8
114 96 7
122 104 9
123 102 10
120 103 8
111 94 7
120 101 9
123 104 11
118 99 8
115 99 8
130 110 11
136 116 13
137 118 13
130 111 12
105 87 6
123 103 9
110 93 6
B-1
Species
Whitebait smelt
Plainfin midshipman
Pacific tomcod
Total
length (mm)
119
121
95
135
117
121
111
113
llA
125
119
127
137
120
115
117
120
117
113
115
116
122
113
127
113
86
77
78
82
82
78
72
86
77
84
86
70
78
72
71
75
67
77
76
87
Standard
length Weight
(mm) (8)
100 11
102 9
80 4
116 13
99 8
104 9
95 6
96 8
98 8
105 10
100 8
106 9
116 14
101 9
98 7
98 8
101 9
100 8
96 7
96 7
99 8
105 9
95 7
109 11
100 13
75 5
67 4
66 4
73 5
71 5
66 4
62 3
75 5
66 4
75 5
76 5
60 3
68 4
61 4
63 3
65 3
58 3
67 4
65 4
78 5
B-2
Total Standard
length length Weight
Species (nun) (mm) (g)
Pacific tomcod 7A 64 4
87 76 5
67 58 3
73 65 4
71 63 3
63 54 2
82 70 5
70 60 3
76 66 3
75 66 4
73 64 3
82 71 5
75 65 3
72 62 4
92 81 6
80 71 4
77 67 4
76 66 4
6A 55 2
72 62 4
66 57 2
81 71 5
67 58 3
73 65 4
87 76 6
64 55 2
69 60 3
61 51 2
82 71 5
58 48 2
86 75 5
77 67 4
Pacific staghorn sculpin 235 199 175 tf tt bullbull
196 167 91
Warty poacher 115 99 9
bullbull bullbull 126 110 10
Showy snailfish 138 121 41
107 95 21 bullbull 50 45 2 bullt bull
56 49 2
Butter sole 42 34 1
40 32 1
195 158 71
224 182 124
m
124 100 18
131 104 20
B-3
bullbull
Species
Butter sole
English sole
Total
length (mm)
132
175
132
44
157
175
201
155
215
136
47
200
142
134
133
49
198
158
142
210
195
220
134
198
220
271
130
160
139
195
155
197
252
115
181
172
188
96
163
172
171
170
175
170
204
181
Standard
length Weight
(mm) (g)
106 20
141 47
108 20
36 1
128 35
142 50
169 69
124 34
183 99
110 23
39 1
162 68
115 24
114 22
105 21
39 1
157 74
126 35
114 24
171 88
159 66
179 98
107 21
159 74
175 93
219 223
105 21
133 35
112 24
159 74
126 33
158 68
204 157
93 12
142 57
146 42
157 56
79 8
138 35
145 44
145 42
141 42
147 41
140 42
171 73
155 47
B-4
Total Standard
length length Weight
Species (mm) (mm) (g)
English sole 157 133 33 tt 185 160 53 it ft
170 145 42 it tt
162 137 33 it tt
173 147 46
Sand sole 246 200 141 it tt
335 273 449
Dover sole 95 80 7 81 68 5 li It
76 63 4 bulli ti
115 96 12 tt tt
98 80 8 tt tt
75 63 3
Dover sole 101 84 9 tt tt
77 60 3 it it
119 101 15 it tt
92 76 7 it tt
115 98 13
Dungeness crab 133 362 Females it tt
13 1 it tt
118 280 Females tt tt
16 1 tt tt
16 1 ft tt
14 1 tt tt
15 1 tt tt
14 1 ti ft
135 369 Females it ft
14 1 tt It
14 1 tt tt
10 1 tt It
17 1 tt ft
15 1 tt tt
144 430 Females
Supplemental (not individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Whitebait smelt 223 1815 81
Pacific tomcod 9104 29989 33
B-5
Appendix Table BmdashContinued
Trawl W-2
Gear 8-m trawl
Date 1 August 1985 Depth 135 ft
Distance 2917 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 132 115 12
123 103 9
131 110 11 102 85 4
131 111 12
128 106 11
130 110 12 1A2 120 17
125 105 11
128 106 11
118 100 9
126 109 10
129 110 11
124 107 11
119 101 9
1A7 125 16
120 100 9
132 112 11
lAO 120 14
126 108 11
M
122 103 10
130 112 12
122 105 10
115 99 9
Pacific tomcod 60 50 2
66 57 2
65 54 2
85 73 5
79 68 4
62 56 2
64 53 2
81 68 4
70 59 3
75 66 3
63 52 2
68 57 2 72 63 3
B-6
Species
Pacific tomcod
Warty poacher bullbull tlaquo
bullbull bull
Pacific sanddab bullbull bull
bullbull bull
bullt bullbull
Total
length (mm)
76
71
75
80
77
73
82
66
76
75
63
75
71
69
70
73
70
80
73
65
73
67
6A
58
72
67
65
60
65
63
70
Ik
12
62
63
71
72
83
61
128
1A7
136
215
229
235
221
Standard
length Weight
(mm) (g)
65 4
60 3
67 3
67 4
64 4
65 3
70 5
57 2
66 4
67 3
52 2
66 4
62 3
58 3
61 3
64 3
59 3
70 4
64 3
54 2
63 3
57 2
55 2
50 2
62 3
55 2
57 2
51 2
56 2
53 2
61 3
63 3
60 3
53 2
53 2
60 3
62 3
74 4
52 2
110 11
127 16
117 12
181 101
193 124
197 133
184 115
B-7
Total Standard
length length Weight
Species (mm) (mm) (8)
Pacific sanddab 201 168 83
241 200 133
225 191 128bull
Butter sole 257 207 169
23141 115 245 200 162
bullbull 252 209 151 M bullbull
210 168 90
214 174 92 1 I
222 181 112
206 163 76
215 175 99
132 110 21
200 160 74
223 181 103
133 107 20
128 102 18
230 191 114 bullbull 127 103 17
118 95 14
143 116 24
131 107 19
124 99 16
255 208 156
145 117 26
183 148 59
150 121 29
206 165 78
207 167 84
220 179 97 M bullbull
117 95 14
138 112 23
English sole 181 154 48
167 141 36
176 148 43 169 142 34
182 153 43
171 145 40
200 170 68
181 153 43
205 174 66
190 159 52
172 147 37
173 146 40 61
212 185
199 167
77
B-8
Total Standard
length length Weight Species (mm) (mm) (8)
English sole 159 133 30 180 152 48
Sand sole 285 238 234 bullbull bull 226 183 99 bullbull bull
259 211 158
Dover sole 252 216 157 bullbull bullbull
146 122 24
99 82 7
105 87 10
148 125 25
250 210 127
112 94 10
105 88 9
275 234 206
282 238 223
108 90 10
104 97 12
97 82 6
70 57 2
171 144 38
105 89 10 M
203 173 63
Supplemental (not Individually measured and weighed)
Mass Mass Mean
count weight (g) weight (g)
Pacific tomcod 67A 1816 27
B-9
Appendix Table BmdashContinued
Trawl W-3
Gear 8-m trawl
Date 1 August 1985 Depth 146 ft
Distance 1214 ft
Total Standard
length length Weight
Species (mm) (mm) (8)
Whitebait smelt 146 126 15
Pacific tomcod 74 62 3
62 51 2
61 51 2
61 51 2
61 51 2
56 46 1
70 59 3
83 71 5
66 55 3
56 46 2
79 65 4
54 45 1
59 49 2
74 63 3
75 63 4
80 70 4
64 53 2
69 60 3
63 54 2
66 57 2
71 61 3
72 62 3
79 68 4
64 54 2
71 61 3
62 51 2
79 68 4
56 45 1
67 57 2
76 64 4
70 60 3
66 54 2
72 62 3
75 65 3
75 63 3
69 59 3
B-10
Species
Pacific tomcod
Pacific sanddab
Speckled sanddab t bulllaquo
bullbull ti
Butter sole
Slender sole English sole
M 1
Dover sole
1^
Total
length (mm)
57
258
221
223
65
68 131 235 2U 122 1A3 200 160 111 178 104
Standard
length Weight
(mm) (8)
47 2
222 183
188 129
191 14
56 2
57 2
114 19
188 136 173 96
98 15
115 25 161 56
137 31 151 47 148 46
86 9
B-11
Appendix Table C--Willapa Bay benthic sampling stations sediment structure analysis-
Date Mean grain Percent Percent
Station (1985) size (phi) silt-clay organic
matter
20-fathom contour
3-8 267 27
W-1-2 31 July 3-8 163 17
W-1-3 31 July 3-6 112 17
W-l-A 30 July 3-9 16
W-1-1 31 July
2A7
W-1-5 30 July 30 A9 12
22-fathom contour
W-2-1 31 July A3 A38 19
21W-2-2 31 July A-l 360
W-2-3 31 July Al 280 18
lAW-2-A 30 July Al 276
W-2-5 30 July A-O 16268
2A-fathom contour
W-3-1 31 July A2 353 21
W-3-2 31 July 16 00 15
W-3-3 31 July AO 238 lA
W-3-4 30 July AO 229 12
W-3-5 30 July AO 2A0 13
377 23A 168Mean
Standard devlatIon 065 1116 039
C-1
Appendix Table D--Fish species collected with an 8-m trawl off Willapa Bay
Scientific name
Phylum CHORDATA
Class Chondrichthyes Family Rajidae
Raja binoculata
Class Osteichthyes Family Osmeridae
Allosmerus elongatus
Spirinchus thaleichthys Family Batrachoididae
Porichthys notatus
Family Gadidae Microgadus proximus
Family Cottidae Leptocottus armatus
Family Agonidae
Ocella verrucosa
Family Cyclopteridae Liparis pulchellus
Family Bothidae Citharichthys sordidus
Citharichthys stigmaeus
Family Pleuronectidae Isopsetta isolepis
Lyopsetta exilis
Microstomus pacificus
Parophrys vetulus
Psettichthys melanostictus
Common name
Cartilaginous fishes Skates
Big skate
Bony fishes Smelts
Whitebait smelt
Longfin smelt Toad fishes
Plainfin midshipman Cods and hake
Pacific tomcod
Sculpins Pacific staghorn sculpin
Poachers
Warty poacher Snailfishes
Showy snailfish Lefteye flounders Pacific sandab
Speckled sandab Righteye flounders
Butter sole
Slender sole
Dover sole
English sole Sand sole
D-l
Appendix Table EmdashDescriptive summary of the benthic invertebrates at 15 sampling stations off Willapa Bay Washington 30-31 July 1985
3Station no W-1-1 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) rsr m^
Nemertinea 15 1000 520 375
Leitoscoloplos puaettensis 76 1000 2638 1084
Scoloplos acmeceps 54 1000 1875 1160
Paraonidae 1 333 34 60
Aricidea nr A suecica 8 1000 277 216
Prionospio cirrifera 18 1000 625 312
Spiophanes berkelevorum 128 1000 4444 1496
Spiophanes bombvx 12 1000 416 454
Magelona sacculata 3 666 104 104
Maaelona hartmanae 2 333 69 120
Magelona longicornisXhartmanae 2 333 69 120
Chaetozone setosa 47 1000 1631 592
Heteromastus filobranchus 7 666 243 262
Mediomastus ambiseta 12 666 416 633
Mediomastus sp indet 2 333 69 120
Notomastus lineatus 7 1000 243 159
Euclvmene zonalis 1 333 34 60
Eteone fauchaldi 3 666 104 104
Eteone sp iuvenile 1 333 34 60
Eteone californica 10 666 347 300
Polvnoidae sp indet 2 333 69 120
Pholoe minuta 33 1000 1145 375
Sthenalais tertiaglabra 9 1000 312 180
Thalenessa spinosa 5 666 173 216
Hesionidae 4 666 138 159
Nereis zonata 2 666 69 60
Glvcera spp iuvenile 19 666 659 677
Glvcinde armigera 46 1000 1597 216
Goniada maculata 3 666 104 104
Nephtys caecoides 20 1000 694 469
Nephtys ciliata 1 333 34 60
Nephtys sp iuvenile 40 1000 1388 433
Diopatra ornata 1 333 34 60
Onuphis iridescens 17 1000 590 469
Onuphis sp indet 23 1000 798 573
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3226 1000 112013 38860
Ampharete acutifrons 10 1000 347 262
Polycirrus sp complex 21 1000 729 00
Gastropoda 11 1000 381 240
Bivalvia 707 1000 24548 7569
E-1
Station no W-1-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 152 1000 5277 573
Diastvlis sp iuvenile 1273 1000 44201 14503
Diastvlopsis dawsoni 881 1000 30590 22676
Leucon subnasica 177 1000 6145 2761
Svnidotea bicuspida 6 1000 208 180
Svnidotea sp 15 1000 520 375
Pleurogonium rubicundum 6 666 208 275
Ampelisca macrocephala 10 1000 347 216
Eohaustorius spp 3 333 104 180
Lvsianassidae juvenile 1 333 34 60
Svnchelium sp juvenile 1 333 34 60
Photis parvidens 20 666 694 677
Photis sp juvenile 73 1000 2534 1910
Protomedeia prudens 34 1000 1180 365
Protomedeia spp juvenile 35 1000 1215 693
Ischvrocerus spp juvenile 8 333 277 481
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 29 1000 1006 573
Rhepoxvnius dabouis 16 1000 555 159
Rhepoxvnius abronius 7 1000 243 60
Dulichia monacantha 12 1000 416 104
Caprellidae 1 333 34 60
Nebalia puaettensis 4 666 138 120
Decapoda larvae 7 666 243 262
Ophiuroidea 46 1000 1597 661
Holothuroidea 7 1000 243 60
Ifumber of taxa= 70
Mean number per sample 248567 Standard deviation (SD) 24971
Mean numberm^ 2589236 SDm^ 260119
H= 299 SDV= 076 SR= 774 J= 049
E-2
3Station no W-1-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 199
Polychaeta 1 333 34 60
Orbiniidae 1 333 34 60
Leitoscoloplos pucrettensis 33 1000 1145 477
Orbinia (Phvlo) felix 1 333 34 60
Scoloplos acmeceps 12 666 416 375
Aricidea nr A suecica 2 333 69 120
Polvdora socialis 1 333 34 60
Minuspio cirrifera 2 333 69 120
Prionospio cirrifera 5 666 173 159
Spiophanes berkelevorum 39 1000 1354 1816
Spiophanes bombvx 69 1000 2395 1721
Maaelona sacculata 3 666 104 104
Haaelona hartmanae 6 666 208 180
Chaetozone setosa 84 1000 2916 721
Tharvx multifilis 1 333 34 60
Heteromastus filobranchus 8 666 277 318
Heteromastus spp 2 333 69 120
Mediomastus ambiseta 1 333 34 60
Mediomastus sp indet 3 666 104 104
Notomastus lineatus 9 1000 312 104
Phvllodoce hartmanae 3 333 104 180
Anaitides aroenlandica 2 333 69 120
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 1 333 34 60
Eteone spp 1 333 34 60
Eteone sp juvenile 5 666 173 159
Halosvdna brevisetosa 1 333 34 60
Pholoe minuta 15 666 520 477
Sthenalais tertiaglabra 17 666 590 627
Thalenessa spinosa 6 666 203 208
Hesionidae 1 333 34 60
Autolvtus sp 1 333 34 60
Syllis sp indet 1 bull333 34 60
Glycinde armicjera 26 1000 902 513
Glycinde sp juvenile 14 666 486 524
Goniada maculata 2 666 69 60
Nephtys caeca 2 666 69 60
Nephtys caecoides 3 333 104 180
Wephtys spp 2 333 69 120
Nephtys sp juvenile 21 666 729 650
Onuphis iridescens 30 1000 1041 650
Onuphis sp indet 25 666 368 945
Lumbrineris bicirrata 7 1000 243 60
E-3
station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
() m^ m^
Lumbrineris luti 2 666 69 60
Arabellidae 1 333 34 60
Mvriochele heeri 3 333 104 180
Owenia fusiformis 6044 1000 209861 52180
Ampharetidae 1 333 34 60
Ampharete acutifrons 6 1000 173 60
Polvcirrus sp complex 9 666 312 275
Amaeana occidentalis 2 333 69 120
Hirudinea 1 333 34 60
Gastropoda 4 666 138 120
Cvlichna attonsa 3 333 104 180
Bivalvia 412 666 14305 12704
Modiolus spp 12 333 416 721
Lucinoma annulata 11 333 381 661
Macoma elimata 2 333 69 120
Macoma expanse 28 333 972 1683
Tellina modesta 29 333 1006 1744
Tellina nuculoides 2 333 69 120
Siliqua sloati 5 333 173 300
Gari californica 2 333 69 120
Copepoda 2 333 69 120
Ostracoda sp A 12 1000 416 375
Ostracoda sp B 4 333 138 240
Diastvlis sp juvenile 712 333 24722 42820
Diastvlopsis spp 686 333 23819 41256
Diastvlopsis dawsoni 1244 1000 43194 13388
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 99 666 3437 3603
Anthuridae 2 666 69 60
Haliophasma aeminata 1 333 34 60
Svnidotea bicuspida 24 1000 833 541
Svnidotea sp 15 666 520 650
Pleuroqonium rubicundum 4 666 138 120
Ampelisca aqassizi 3 666 104 104
Arapelisca macrocephala 4 666 138 159
Wecomedon spp 2 333 69 120
Photis parvidens 16 666 555 636
Photis sp juvenile 45 666 1562 1617
Photis macinerneyi 16 333 555 962
Protomedeia prudens 10 666 347 300
Protoinedeia spp juvenile 31 666 1076 933
Ischyroceridae 3 333 104 130
Ischvrocerus spp juvenile 4 333 138 240
Ischyrocerus pegalops 13 333 451 731
Rhepoxynius spp 16 666 555 636
E-4
3Station no W-1-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m2
Rhepoxvnius dabouis 7 666 243 262
Rhepoxvnius abronius 22 1000 763 1053
Foxiphalus spp 1 333 34 60
Dulichia monacantha 10 666 347 365
Caprellidae 6 1000 208 104
Nebalia puaettensis 1 333 34 60
Ophiuroidea 13 666 451 394
Amphiodia spp 3 333 104 180
Number of taxa= 98
Mean number per sample 336433 Standard deviation (SD) 108152
Mean numberm 2 3504514 SDm21126588
H= 241 SDV= 061 SR=1052 J= 036
E-5
Station no W-1-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Anthozoa 1
Nemertinea 13
Leitoscoloplos ouaettensis 62
Orbinia (Phvlo) felix 1
Scoloplos acmeceps 31
Aricidea nr A suecica 6
Spiophanes berkelevorum 252
Spiophanes bombvx 32
Maaelona lonaicornisXhartmanae 1
Frequency of
occurrence
()
333
1000
1000
333
1000
666
1000
1000
333
1000
1000
333
666
666
333
333
333
333
666
333
1000
333
1000
1000
666
333
666
333
666
1000
666
1000
666
1000
1000
1000
333
1000
666
1000
1000
1000
666
666
666
Mean Standard
number deviation
itr mr
34 60
451 323
2152 1011
34 60
1076 601
208 180
8750 2436
1111 365
34 60
1493 592
486 216
208 360
104 104
173 216
34 60
34 60
104 180
34 60
69- 60
34 60
763 262
34 60
1180 433
972 573
69 60
173 300
104 104
34 60
69 60
1597 334
729 633
277 159
659 739
1701 767
1180 1601
763 492
69 120
34340 53917
69 60
659 421
243 159
23472 6096
173 159
694 636
69 60
Chaetozone setosa
Heteromastus filobranchus
Hediomastus ambiseta
Notomastus lineatus
Maldanidae sp indet
Euclvmene zonalis
Phvllodoce hartmanae
Phvllodoce multiseriata
Anaitides multipapillata
Eteone fauchaldi
Eteone sp juvenile
Eteone californica
Polynoidae Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Glvcera capitata
Glycera convoluta
Glycinde arraiaera
Glycinde sp juvenile
Goniada maculata
Nephtys caecoides
Nephtys sp juvenile Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Owenia fusiformis
Ampharete acutifrons
Polycirrus sp complex
Gastropoda Bivalvia
Copepoda
Ostracoda sp A
Ostracoda sp 3
43
14
6
3
5
1
1
3
1
2
1
22
1
34
28
2
5
3
1
2
46
21
8
19
49
34
22
2
2429
2
19
7
676
5
20
2
E-6
3Station no W-1-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Diastvlis sp juvenile 957 1000 33229 19551
Diastvlopsis dawsoni 1161 1000 40312 17324
Leucon subnasica 162 1000 5625 3725
Anthuridae 1 333 34 60
Svnidotea bicuspida 12 1000 416 275
Synidotea sp 18 1000 625 360
Pleuroaonium rubicundum 23 1000 798 394
Ampelisca macrocephala 4 666 138 120
Anonvs adoxusXcerinatus 18 1000 625 360
Lysianassidae juvenile 2 333 69 120
Photis parvidens 87 1000 3020 1502
Photis sp juvenile 103 1000 3576 2376
Protomedeia prudens 93 1000 3229 1637
Protomedeia spp juvenile 33 333 1145 1984
ProtomedeiaXCheirimedeia sp 24 333 833 1443
Cheirimedeia sp 5 333 173 300
Ischvrocerus sp 3 666 104 104
Ischvrocerus peaalops 9 333 312 541
Rhepoxvnius spp 6 333 208 360
Rhepoxynius dabouis 12 1000 416 00
Rhepoxvnius abronius 22 1000 763 365
Svmpleustes subalaber 1 333 34 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 2 666 69 60
Caprellidae 1 333 34 60
Nebalia pugettensis 2 333 69 120
Pinnixa sp megalopa 1 333 34 60
Ophiuroidea 31 1000 1076 534
Holothuroidea 11 1000 381 60
Pycnogonida 1 333 34 60
Number of taxa= 75
bullMean number per sample 224900 Standard deviation (SD) 66079
Mean numberm^ 2342708 SDm 688326
H= 328 SDV= 081 SR= 839 J= 053
E-7
Station no W-1-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Stande
number occurrence number deviat]
) m^ m^
Nemertinea 6 1000 208 104
Leitoscoloolos puaettensis 38 1000 1319 318
Orbinia (Phylo) felix 2 333 69 120
Scoloplos acmeceps 21 1000 729 580
Aricidea nr A suecica 7 1000 243 120
Prionospio cirrifera 7 1000 243 159
Spiophanes berkeleyorum 86 1000 2986 627
Spiophanes bombyx 102 1000 3541 1603
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 61 1000 2118 433
Heteromastus filobranchus 9 1000 312 104
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 333 69 120
Notomastus lineatus 15 1000 520 375
Maldanidae 2 333 69 120
Euclymene zonalis 3 666 104 104
Phyllodoce hartmanae 1 333 34 60
Phyllodoce longipes 2 333 69 120
Phyllodoce multiseriata 1 333 34 60
Anaitides multipapillata 2 333 69 120
Eteone fauchaldi 5 666 173 216
Eteone sp juvenile 6 333 208 360
Eteone californica 7 666 243 216
Tenonia kitsapensis 1 333 34 60
Pholoe minuta 11 1000 381 60
Sthenalais tertiaglabra 14 1000 486 300
Thalenessa spinosa 2 333 69 120
Hesionidae 2 333 69 120
Heteropodarke heteromorpha 1 333 34 60
Autolytus sp 3 333 104 180
Nereis zonata 1 333 34 60
Glycinde armiaera 21 1000 729 203
Glycinde sp juvenile 4 333 138 240
Goniada maculata 12 1000 416 375
Goniada sp juvenile 2 666 69 60
Nephtys caecoides 22 666 763 781
Nephtys sp juvenile 47 1000 1631 573
Onuphis iridescens 59 1000 2043 1734
Onuphis sp indet 71 1000 2465 2294
Lumbrineris sp indet 3 666 104 104
Owenia fusiformis 3815 1000 132465 101553
Ampharete acutifrons 15 1000 520 104
Polvcirrus sp complex 13 1000 451 216
Gastropoda 2 333 69 120
E-8
3Station no W-1-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() VA nr
Bivalvia 704 1000 24444 5001
Copepoda 111 1000 3854 751
Ostracoda sp A 18 1000 625 275
Ostracoda sp B 12 1000 416 180
Diastvlis sp juvenile 901 1000 31284 16570
Diastvlopsis dawsoni 794 1000 27569 24369
Oxvurostvlis pacifica 3 666 104 104
Leucon subnasica 73 1000 2534 492
Edotea sublittoralis 4 333 138 240
Svnidotea bicuspida 7 333 243 421
Svnidotea sp 19 1000 659 524
Pleuroaonium rubicundum 25 666 868 1011
Bathvcopea daltonae 1 333 34 60
Ampelisca macrocephala 8 666 277 262
Ampelisca sp 1 333 34 60
Atvlus tridens 1 333 34 60
Eohaustorius spp 1 333 34 60
Eohaustorius sawveri 1 333 34 60
Anonvx spp () 1 333 34 60
Anonvs adoxusXcerinatus 4 666 138 159
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 29 1000 1006 318
Photis SP juvenile 62 666 2152 2135
Protomedeia prudens 17 666 590 513
Protomedeia spp juvenile 15 666 520 477
Ischvrocerus spp juvenile 26 666 902 1387
Ischvrocerus pegalops 14 333 486 842
Rhepoxvnius spp 62 1000 2152 510
Rhepoxynius dabouis 17 1000 590 313
Rhepoxvnius abronius 18 1000 625 360
Sympleustes subglaber 4 333 138 240
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 1 333 34 60
Caprellidae 7 666 243 334
Nebalia pugettensis 5 666 173 216
Decapoda larvae 21 1000 729 180
Callianassa californiensis 1 333 34 60
Ophiuroidea 29 1000 1006 492
Holothuroidea 2 666 69 60
Pycnogonida 1 333 34 60
E-9
station no W-1-4 Date 30 Jul 85 Sample size (CONTINUED)
Number of taxa= 85
Mean number per sample 251267 Standard deviation (SD) 148529
Mean numberm^ 2617361 SDm^1547174
H= 289 SDV= 071 SR= 941 J= 045
bullV
E-10
3Station no W-1-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrencej number deviation
() rtr m^
Nemertinea 5 1000 173 85
Leitoscoloplos puaettensis 62 1000 2152 1069
Phylo (Orbinia) felix 1 333 34 60
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 2 666 69 60
Prionospio cirrifera 1 333 34 60
Spiophanes berkelevorum 57 1000 1979 993
Spiophanes bombvx 131 1000 4548 1935
Maaelona sacculata 5 1000 173 60
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 72 1000 2500 477
Heteromastus filobranchus 1 333 34 60
Mediomastus sp indet 1 333 34 60
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 4 666 138 120
Phyllodoce hartmanae 3 666 104 104
Phvllodoce lonaipes 1 333 34 60
Anaitides multipapillata 2 666 69 60
Eteone fauchaldi 3 666 104 104
Eteone californica 7 666 243 262
Pholoe minuta 5 1000 173 60
Sthenalais tertiaglabra 11 1000 381 481
Thalenessa spinosa 5 666 173 159
Nereis zonata 1 333 34 60
Nereis spp 1 333 34 60
Glvcera convoluta 1 333 34 60
Glycinde armigera 26 1000 902 513
Glycinde sp juvenile 1 333 34 60
Goniada maculata 9 1000 312 203
Nephtys caecoides 21 1000 729 275
Nephtys sp juvenile 37 1000 1284 962
Onuphis iridescens 8 1000 277 300
Onuphis sp indet 149 1000 5173 1793
Lumbrineris bicirrata 5 666 173 159
Owenia fusiformis 3007 1000 104409 7337
Ampharete acutifrons 10 1000 347 60
Pista brevibranchiata 1 333 34 60
Polycirrus sp complex 14 1000 436 513
Gastropoda 4 333 133 240
Cylichna attonsa 7 333 243 421
Bivalvia 537 1000 18645 4453
Copepoda 22 333 763 1323
Ostracoda sp A 12 1000 416 360
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 499 1000 17326 25978
E-11
Station no W-1-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Diastvlopsis dawsoni
Leucon subnasica
Edotea sublittoralis
Idotea sp
Svnidotea bicuspida
Pleuroaonium rubicundum
Gammaridea
Arapelisca macrocephala
Ampelisca spp juvenile
Anonvx spp ()
Isaeidae
Photis brevipes
Photis sp juvenile
Protomedeia prudens
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Svmpleustes subalaber
Dulichia monacantha
Caprellidae Nebalia puaettensis
Brachyura Ophiuroidea Holothuroidea
Pycnogonida
Number of taxa= 71
Mean number per sample 181333
Mean numberm^ 1888889 SDm
H= 281 SDV= 067 SR= 814
Standard deviation (SD) 46942
488976
J= 046
Total
number
240
43
5
1
30
18
5
15
3
5
2
51
79
1
56
16
9
49
1
9
1
3
1
18
1
1
Frequency of
occurrence
()
1000
1000
666
333
666
666
666
1000
333
333
333
666
666
333
1000
333
666
1000
333
1000
333
666
333
1000
333
333
Mean Standard
number deviation m^ m^
8333 6969
1493 2047
173 159
34 60
1041 1458
625 683
173 159
520 312
104 180
173 300
69 120
1770 1825
2743 3641
34 60
1944 1421
555 962
312 375
1701 1147
34 60
312 104
34 60
104 104
34 60
625 375
34 60
34 60
E-12
station no W-2-1
Taxa
Nemertinea
Leitoscoloplos puaettensis
Phylo (Orbinia) felix
Scoloplos acmeceps
Aricidea nr A suecica
Prionospio cirrifera
Spiophanes berkeleyorum
Spiophanes bombyx
Hagelona hartmanae
Chaetozone setosa
Tharyx secundus
Tharyx sp indet Heteromastus filobranchus
Mediomastus ambiseta
Mediomastus sp indet
Notomastus lineatus
Unid Naldanidae
Euclymene zonalis
Phyllodoce hartmanae
Phyllodoce multiseriata
Eteone californica
Aphrodite nealiaens
Tenonia kitsapensis
Pholoe minuta
Sthenalais tertiaalabra
Thalenessa spinosa
Hesionidae
Gyptis brevipalpa
Autolytus so
Glycera spp iuvenile tlaquos Goniadidae
Glycinde armiaera
Goniada maculata
Nephtys caecoides
Nephtys sp iuvenile
Onuphis iridescens
Onuphis sp indet
Lumbrineris bicirrata
Notocirrus californiensis
Ovfenia fusiformis
Ampharete acutifrons
Melinna elisabethae
Polycirrus sp complex
Gastropoda Bivalvia
3Date 31 Jul 85 Sample size
Total
number
16
90
9
59
11
14
35
53
12
29
5
2
5
15
5
3
6
1
5
7
6
1
1
41
4
6
1
1
1
1
1
25
5
17
24
4
42
5
1
4471
4
1
15
6
293
Frequency of Mean Standard
occurrence
)
1000
1000
666
1000
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
666
666
333
333
666
333
666
333
333
1000
666
1000
333
333
333
333
333
1000
1000
1000
1000
1000
1000
1000
333
1000
1000
333
1000
1000
1000
number deviation
m^ m^
555 370
3125 1816
312 275
2048 753
381 318
486 394
1215 365
1840 1310
416 208
1006 159
173 120
69 120
173 120
520 180
173 216
104 104
208 360
34 60
173 216
243 421
208 275
34 60
34 60
1423 159
138 159
203 180
34 60
34 60
34 60
34 60
34 60
863 300
173 60
590 334
833 275
138 60
1458 729
173 60
34 60
155243 50907
138 60
34 60
520 312
203 104
10173 1355
E-13
Station no W-2-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m2 m^
Copepoda 85 1000 2951 2104
Ostracoda sp A 71 1000 2465 1896
Ostracoda sp B 8 1000 277 159
Diastylis sp juvenile 244 1000 8472 12063
Diastvlopsis dawsoni 217 1000 7534 3799
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 26 1000 902 481
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 2 333 69 120
Svnidotea sp 7 1000 243 60
Pleuroaonium rubicundum 13 1000 451 365
Gammaridea 2 333 69 120
Ampelisca macrocephala 13 1000 451 513
Ampelisca spp juvenile 2 333 69 120
Eohaustorius sawyeri 1 333 34 60
Lysianassidae juvenile 4 333 138 240
Photis parvidens 18 666 625 683
Photis sp juvenile 39 1000 1354 1366
Protomedeia prudens 9 666 312 275
Protomedeia spp juvenile 20 1000 694 394
Rhepoxynius spp 78 1000 2708 104
Rhepoxynius dabouis 53 1000 1840 365
Rhepoxynius abronius 2 333 69 120
Sympleustes subalaber 1 333 34 60
Dulichia monacantha 8 666 277 394
Dulichia sp juvenile 12 666 416 633
Caprellidae 6 1000 208 104
Nebalia puqettensis 10 1000 347 159
Decapoda larvae 4 333 138 240
Pinnixa sp megalopa 2 333 69 120
Ophiuroidea 15 666 520 520
Holothuroidea 1 333 34 60
Number of taxa= 77
Mean number per sample 211133 Standard deviation (SD 59744
Mean numberm^ 2199306 SDm^ 622333
H= 226 SDV= 050 SR= 868 J= 036
E-14
3station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr xs
Anthozoa 2 333 69 120
Nemertinea 10 1000 347 120
Orbiniidae 1 333 34 60
Leitoscoloplos puaettensis 96 1000 3333 1469
Orbinia (Phvlo) felix 3 666 104 104
Scoloplos acmeceps 38 666 1319 1302
Aricidea nr A suecica 4 666 138 159
Minuspio cirrifera 5 333 173 300
Prionospio cirrifera 16 666 555 524
Spiophanes berkelevorum 238 1000 8263 7010
Spiophanes bombyx 36 1000 1250 104
Maaelona sacculata 2 333 69 120
Maaelona hartmanae 5 666 173 159
Chaetozone setosa 30 1000 1041 360
Heteromastus filobranchus 83 1000 2881 3668
Hediomastus ambiseta 34 666 1180 1069
Mediomastus sp indet 25 666 868 945
Notomastus lineatus 1 333 34 60
Euclvmene zonalis 5 666 173 159
Axiothella rubrocincta 2 333 69 120
Phyllodocidae 5 333 173 300
Phvllodoce hartmanae 7 1000 242 340
Phvllodoce sp iuveniles 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone spp 2 333 69 120
Eteone sp juvenile 2 333 69 120
Eteone californica 1 333 34 60
Polynoidae 2 333 69 120
Polynoidae sp indet 1 333 34 60
Halosydna brevisetosa 2 333 69 120
Pholoe minuta 28 666 972 939
Sthenalais tertiaglabra 3 666 104 104
Thalenessa spinosa 14 666 486 753
Hesionidae 1 333 34 60
Gvptis brevipalpa 1 333 34 60
Glycera capitata 5 333 173 300
Glycera spp juvenile 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 20 1000 694 753
51ycinde sp juvenile 7 666 243 262
Glycinde picta 12 333 416 721
Goniada maculata 2 333 69 120
Mephtys caecoides 5 666 173 159
Mephtys longosetosa 3 333 104 130
E--15
Station no W-2-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() vr m^
Nephtys spp 33 333 1145 1984
Nephtys sp juvenile 40 666 1388 1202
Onuphis iridescens 5 666 173 216
Onuphis eleaans 19 333 659 1142
Onuphis sp indet 19 666 659 627
Lumbrineris bicirrata 2 333 69 120
Lumbrineris cruzensis 1 333 34 60
Lumbrineris luti 2 666 69 60
Lumbrineris sp indet 2 333 69 120
Mvriochele heeri 83 333 2881 4991
Owenia collaris 38 333 1319 2285
Owenia fusiformis 2635 1000 91493 31624
Pectinaria spp 2 333 69 120
Pectinaria californiensis 2 333 69 120
Ampharete acutifrons 7 666 243 334
Polvcirrus sp complex 12 666 416 375
Amaeana occidentalis 2 333 69 120
Gastropoda 1 333 34 60
Bivalvia 116 666 4027 3489
Modiolus spp 22 333 763 1323
Lucinoma annulata 17 333 590 1022
Nacoma elimata 2 333 69 120
Macoma expanse 27 333 937 1623
Tellina modesta 10 333 347 601
Siliqua sloati 33 333 1145 1984
Copepoda 12 1000 416 275
Ostracoda 7 333 243 421
Ostracoda sp A 13 666 451 693
Ostracoda sp B 10 333 347 601
Diastylis sp juvenile 319 666 11076 16808
Diastvlopsis dawsoni 2586 1000 89791 121282
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 71 1000 2465 1379
Edotea sublittoralis 1 333 34 60
Synidotea bicuspida 4 666 138 159
Synidotea sp 5 666 173 159
Munnoqonium waldronense 12 333 416 721
Pleurogonium rubicundum 23 666 798 1207
Ampelisca aqassizi 1 333 34 60
Ampelisca macrocephala 6 333 203 360
Ampelisca spp juvenile 1 333 34 60
Lysianassidae 11 666 381 334
Lysianassidae juvenile 4 333 138 240
Photis parvidens 9 666 312 312
Photis sp juvenile 12 666 416 375
Photis spp 22 333 763 1323
E-16
3
(CONTINUED)
Station no W-2-2 Date 31 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Protomedeia orudens 14 666 486 753
Protomedeia spp iuvenile 16 666 555 709
Ischvrocerus spp 1 333 34 60
Cheriphotis sp 19 333 659 1142
Rhepoxvnius spp 35 666 1215 1198
Rhepoxvnius dabouis 12 666 416 375
Rhepoxvnius abronius 6 333 208 360
Parapleustes den 1 333 34 60
Dulichia monacantha 10 333 347 601
Dulichia sp juvenile 2 333 69 120
Caprellidae 11 666 381 421
Nebalia pugettensis 2 333 69 120
Ophiuroidea 13 666 451 421
Amphiodia spp 5 333 173 300
Holothuroidea 4 666 138 120
Number of taxa= 106
Mean number per sample 239200 Standard deviation (SD) 154661
Mean numberm^ 2491667 SDra 1611054
H= 298 SDV= 073 SR=1183 J= 044
E-17
Station no W-2-3 Date 31 Jul 85 Sample size 3
Taxa Total
number
Nemertinea 10
Leitoscoloplos nuaettensis 11
Orbinia (Phylo) felix 6
Scoloplos acmeceps 1
Aricidea nr A suecica 78
Laonice cirrata 1
Polvdora socialis 1
Spiophanes berkelevorum 23
Spiophanes bombyx 429
Maaelona hartmanae 9
Maaelona lonciicornishartmanae 1 Chaetozone setosa 33
Tharvx multifilis 1
Tharyx secundus 2
Decamastus aracilis 4
Nediomastus ambiseta 2
Mediomastus sp indet 5
Notomastus lineatus 1
Maldanidae sp indet 1
Maldanidae 1
Euclymene zonalis 14
Phyllodoce hartmanae 7
Phyllodoce multiseriata 13
Anaitides multipapillata
Eteone sp indet 1
Eteone californica
Paranaitides polvnoides 1
Pholoe minuta 3
Sthenalais tertiaalabra 3
Gyptis brevipalpa 3
Autolytus sp 1
Glycera convoluta 1
Glycinde armiaera 11
Glycinde sp juvenile 3
Goniada maculata
Nephtys caecoides 17
Nephtys sp juvenile 23
Onuphis iridescens 45
Onuphis sp indet 90
Lumbrineris bicirrata 1
Lumbrineris luti
Lumbrineris sp indet 1
Galathowenia nr G oculata 7
Owenia fusiformis 16691
Melinna sp juvenile 3
Frequency of
occurrence
()
1000
1000
1000
333
1000
333
333
1000
1000
666
333
1000
333
666
666
333
666
333
333
333
1000
666
1000
333
333
1000
333
666
666
666
333
333
1000
333
1000
1000
1000
666
1000
333
333
333
1000
1000
1000
Mean Standard
number deviation
vr m^
347 199
381 120
208 180
34 60
2708 1201
34 60
34 60
798 421
14895 3428
312 312
34 60
1145 416
34 60
69 60
138 159
69 120
173 216
34 60
34 60
34 60
486 120
243 216
451 365
69 120
34 60
208 104
34 60
104 104
104 104
104 104
34 60
34 60
381 159
104 180
138 60
590 334
798 262
1562 2270
3125 1825
34 60
69 120
34 60
243 240
58715 22314 277 60
E-18
3Station no W-2-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total
number
Polycirrus sp complex 10
Euchone nr E hancocki 3
Gastropoda 8
Bivalvia 110
Copepoda 10
Ostracoda sp A 51
Ostracoda sp B 3
Diastylis sp juvenile 21
Diastvlopsis dawsoni 79
Oxvurostvlis pacifica 1
Leucon subnasica 6
Isopoda 1
Pleuroaonium rubicundum 1
Ampelisca macrocephala 21
Ampelisca spp juvenile 6
Eohaustorius spp 1
Lysianassidae juvenile 1
Photis brevipes 4
Photis parvidens 10
Photis sp juvenile 16
Protomedeia prudens 6
Protomedeia spp juvenile 17
Ischyrocerus spp juvenile 1
Rhepoxvnius spp 34
Rhepoxvnius dabouis 20
Rhepoxvnius abronius 15
Sympleustes subglaber 5
Dulichia monacantha 3
Dulichia sp juvenile 4
Caprellidae 8
Pinnixa sp megalopa 1
Ophiuroidea 17
Holothuroidea 1
Number of taxa= 78
Mean number per sample 102433
Mean numberm^ 1067014 SDra
H= 296 SDV= 067 SR= 959
Frequency of Mean Standard
occurrence number deviation
() m^ m^
1000
1000
1000
1000
333
1000
666
666
666
333
666
333
333
1000
1000
333
333
666
333
1000
333
1000
333
1000
1000
1000
666
333
666
1000
333
1000
333
347 60
104 00
277 159
3819 781
347 601
1770 729
104 104
729 650
2743 3641
34 60
208 275
34 60
34 60
729 275
208 180
34 60
34 60
138 159
347 601
555 60
208 360
590 216
34 60
1180 835
694 394
520 375
173 216
104 180
138 159
277 159
34 60
590 216
34 60
Standard deviation (SD) 13963
197536
J= 047
E-19
Station no W-2-4 Date 30 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 5 1000 173 85
Echiura 1 333 34 60
Leitoscoloolos nuaettensis 34 1000 1180 809
Phvlo (Orbinia) felix 6 1000 208 208
Scoloplos acmeceps 12 1000 416 208
Aricidea nr A suecica 91 1000 3159 2304
Spionidae 2 333 69 120
Prionospio cirrifera 9 333 312 541
Spio filicornis 1 333 34 60
Spiophanes berkelevorum 40 1000 1388 573
Spiophanes bombvx 381 1000 13229 4899
Maaelona sacculata 1 333 34 60
Haaelona hartmanae 7 666 243 216
Chaetozone setosa 56 1000 1944 1043
Heteromastus filobranchus 3 666 104 104
Mediomastus ambiseta 11 1000 381 318
Mediomastus sp indet 9 1000 312 104
Notomastus lineatus 2 666 69 60
Maldanidae 3 666 104 104
Euclvmene zonalis 15 1000 520 104
Phvllodoce hartmanae 13 1000 451 159
Phvllodoce multiseriata 2 333 69 120
Phvllodoce sp juveniles 3 666 104 104
Eteone sp juvenile 1 333 34 60
Eteone californica 7 1000 243 159
Pholoe minuta 35 1000 1215 394
Sthenalais tertiaalabra 4 666 138 159
Thalenessa spinosa 4 666 13-3 159
Syllidae 1 333 34 60
Autolvtus sp 1 333 34 60
Glvcinde armiaera 11 1000 381 120
Glvcinde sp juvenile 1 333 34 60
Goniada maculata 8 666 277 240
Goniada sp juvenile 1 333 34 60
Nephtvs caecoides 17 666 590 534
Nephtvs sp juvenile 51 1000 1770 1082
Onuphis iridescens 29 1000 1006 1294
Onuphis sp indet 91 1000 3159 2535
Lumbrineris bicirrata 9 666 312 275
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Galathowenia nr G oculata 3 666 104 104
Owenia fusiformis 1306 1000 62708 9504
Pectinaria californiensis 2 666 69 60
E-20
3Station no W-2-4 Date 30 Jul 85 Sample size
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrencegt number deviation
() m^ m^
Ampharete acutifrons 4 666 138 159
Melinna sp juvenile 2 333 69 120
Polvcirrus sp complex 13 1000 451 318
Euchone nr E hancocki 1 333 34 60
Gastropoda 14 1000 486 262
Bivalvia 140 1000 4861 573
Scaphopoda 1 333 34 60
Copepoda 26 666 902 939
Ostracoda sp A 117 1000 4062 208
Ostracoda sp B 1 333 34 60
Diastylis sp juvenile 3 333 104 180
Diastvlopsis dawsoni 50 1000 1736 1147
OxYurostvlis pacifica 4 666 138 159
Leucon subnasica 5 666 173 159
Camplvapsis sp 1 333 34 60
Anthuridae 1 333 34 60
Edotea sublittoralis 8 666 277 318
Synidotea sp 1 333 34 60
Nunna sp 1 333 34 60
Pleurogonium rubicundum 2 666 69 60
Gnorimosphaeroma oregonensis 1 333 34 60
Ampelisca agassizi 7 333 243 421
Ampelisca macrocephala 33 1000 1145 520
Ampelisca spp juvenile 4 333 138 240
m Eohaustorius spp 2 333 69 120
Lysianassidae 3 333 104 180
Anonys adoxuscerinatus 1 333 34 60
Lysianassidae juvenile 4 666 138 159
Photis brevipes 10 1000 347 216
Photis sp juvenile 39 1000 1354 454
m Protomedeia prudens 4 333 138 240
Protomedeia spp juvenile 16 1000 555 240
Ischyrocerus pegalops 1 333 34 60
Rhepoxynius spp 37 1000 1284 781
Rhepoxynius dabouis 21 1000 729 208
Rhepoxynius abronius 6 666 208 208
Ax Sympleustes subglaber 7 1000 243 60
Dulichia monacantha 4 333 138 240
Dulichia sp juvenile 12 666 416 375
Caprellidae 10 666 347 433
Nebalia pugettensis 4 666 138 159
Callianassa californiensis 1 333 34 60
m Pinnixa sp megalopa 1 333 34 60
Porcellanidae 1 333 34 60
Ophiuroidea 4 1000 138 60
E-21
station no W-2-4 Date 30 Jul 85 Sample size 3 (CONTINUED)
Taxa Total Frequency of Mean Standard number occurrence number deviation
) m m
Holothuroidea 333 34 60
Number of taxa= 90
Mean number per sample 114000 Standard deviation (SD) 9978
Mean numberm^ 1187500 SDm^ 103942
H= 322 SDV= 070 SR=1106 J= 050
E-2
3Station no W-2-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 3 666 104 104
Leitoscoloolos puaettensis 29 1000 1006 433
Phvlo (Orbinia) felix 10 1000 347 216
Scoloplos acmeceps 9 1000 312 180
Aricidea nr A suecica 22 1000 763 421
Spiophanes berkelevorum 23 1000 798 216
Spiophanes bombvx 221 1000 7673 910
Maaelona sacculata 1 333 34 60
Magelona hartmanae 5 666 173 159
Chaetozone setosa 36 1000 1250 551
Hediomastus ambiseta 5 1000 173 60
Mediomastus sp indet 3 333 104 180
Mediomastus californiensis 1 333 34 60
Maldanidae 4 666 138 159
Euclvmene zonalis 7 1000 243 159
Phvllodoce hartmanae 7 1000 243 60
Phyllodoce longipes 1 333 34 60
Phvllodoce multiseriata 2 333 69 120
Eteone fauchaldi 3 1000 104 00
Eteone spp 2 333 69 120
Eteone californica 3 333 104 180
Polynoidae 1 333 34 60
Pholoe minuta 20 1000 694 240
Sthenalais tertiaglabra 3 333 104 180
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armigera 7 1000 243 120
Glycinde sp juvenile 1 333 34 60
Goniada maculata 10 1000 347 334
Nephtys caecoides 26 1000 902 120
Nephtys sp juvenile 25 1000 868 524
Onuphis iridescens 60 666 2083 2678
Onuphis sp indet 72 666 2500 3801
Lumbrineris bicirrata 4 666 138 159
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 1 333 34 60
Owenia fusiformis 1132 1000 41041 12484
Arapharete acutifrons 4 666 133 120
Melinna elisabethae 2 333 69 120
Terebellidae 1 333 34 60
Polycirrus sp complex 11 1000 381 216
Euchone nr E hancocki 2 666 69 60
Gastropoda 5 1000 173 60
Bivalvia 127 1000 4409 1413
Ostracoda sp A 75 1000 2604 2226
E-23
Station no W-2-5 Date 30 Jul 85 Sample size 3
(CONTINUED)
Taxa
Ostracoda sp B Diastvlopsis dawsoni
Hemilamprops sp
Leucon subnasica
Edotea sublittoralis
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Ampelisca aaassizi
Ampelisca macrocephala
Ampelisca spp iuvenile
Eohaustorius sawveri
Corophidae Isaeidae
Photis brevipes
Photis sp juvenile
Rhepoxvnius spp
Rhepoxvnius dabouis
Rhepoxvnius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Callianassa californiensis
Pinnixa sp megalopa
Porcellanidae
Ophiuroidea
Number of taxa= 71
Mean number per sample
Mean numberm^ 774653
H= 315 SDV= 070
Total Frequency of
number occurrence
()
2 666
22 1000
1 333
2 333
5 666
1 333
1 333
2 333
12 1000
13 1000
18 333
2 333
2 333
4 333
3 333
10 666
43 666
12 666
11 666
3 333
3 333
4 1000
1 333
4 1000
1 333
10 1000
Mean Standard
number deviation
m^ m^
69 60
763 573
34 60
69 120
173 159
34 60
34 60
69 120
416 541
451 240
625 1082
69 120
69 120
138 240
104 180
347 513
1493 1693
416 551
381 492
104 180
104 180
138 60
34 60
138 60
34 60
347 60
74367 Standard deviation (SD) 15106
SDm 157358
SR= 908 J= 051
E-24
Station no W-3-1 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m2
Nemertinea 6 1000 208 104
Leitoscoloplos puaettensis 12 666 416 375
Phvlo (Orbinia) felix 4 666 138 159
Scoloplos acmeceos 2 666 69 60
Aricidea nr A suecica 90 1000 3125 633
Prionospio cirrifera 3 333 104 180
Prionospio cirriferaMighti 1 333 34 60
Spiophanes berkelevorum 5 1000 173 60
Spiophanes bombvx 224 1000 7777 1027
Magelona sacculata 1 333 34 60
Magelona hartmanae 8 666 277 318
Magelona longicornisXhartmanae 1 333 34 60
Chaetozone setosa 32 1000 1111 334
Tharvx multifilis 1 333 34 60
Decamastus gracilis 8 666 277 318
Mediomastus ambiseta 6 333 208 360
Mediomastus sp indet 2 333 69 120
Mediomastus californiensis 1 333 34 60
Notomastus lineatus 5 666 173 159
Haldanidae 1 333 34 60
Euclvmene zonalis 2 666 69 60
Praxillella affinis 3 333 104 180
Phyllodoce mucosa 1 333 34 60
Phyllodoce multiseriata 4 666 138 159
Anaitides multipapillata 1 333 34 60
Eteone fauchaldi 1 333 34 60
Eteone sp indet 2 333 69 120
Eteone sp juvenile 1 333 34 60
Eteone californica 2 666 69 60
Eumida sanguinea 1 333 34 60
Aphroditidae 1 333 34 60
Pholoe minuta 2 666 69 60
Sthenalais tertiaglabra 2 333 69 120
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Autolytus sp- 3 666 104 104
Glycera convoluta 1 333 34 60
Glycinde armigera 10 1000 347 60
Glycinde sp juvenile 2 333 69 120
Goniada maculata 3 666 104 104
Nephtys caeca 2 333 69 120
Nephtys caecoides 11 1000 381 394
Nephtys sp juvenile 21 1000 729 375
Onuphidae sp juvenile 23 333 798 1383
E-25
station no W-3-1 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
) m^ vr
OnuDhis iridescens 14 666 486 469
Onuphis sp indet 88 666 3055 2678
Lumbrineris bicirrata 4 333 138 240
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 8 1000 277 216
Galathowenia nr G oculata 6 1000 208 180
Owenia fusiformis 1426 1000 49513 30017
Pectinaria californiensis 1 333 34 60
Ampharetidae 4 666 138 159
Ampharete acutifrons 1 333 34 60
Polycirrus sp complex 6 1000 208 104
Euchone nr E hancocki 6 666 208 208
Hirudinea 1 333 34 60
Gastropoda 21 1000 729 275
Bivalvia 84 1000 2916 1201
Copepoda 32 1000 1111 573
Ostracoda sp A 95 1000 3298 739
Diastvlopsis dawsoni 26 1000 902 669
Oxvurostvlis pacifica 1 333 34 60
Leucon subnasica 6 666 208 180
Edotea sublittoralis 3 666 104 104
Pleuroaonium rubicundum 5 1000 173 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 11 666 381 334
Ampelisca macrocephala 18 1000 625 00
Ampelisca spp juvenile 18 666 625 903
Eohaustorius spp 4 333 138 240
Eohaustorius sawyeri 4 666 133 159
Photis parvidens 5 666 173 159
Photis sp juvenile 8 666 277 262
Protomedeia prudens 5 1000 173 60
Protomedeia spp juvenile 7 666 243 216
Rhepoxvnius spp 25 1000 368 731
Rhepoxynius dabouis 32 1000 1111 669
Rhepoxvnius abronius 10 1000 347 60
Dulichia sp juvenile 3 666 104 104
Caprellidae 10 666 347 433
Nebalia puqettensis 2 333 69 120
Decapoda larvae 5 666 173 216
Pinnixa sp megalopa 5 666 173 159
Ophiuroidea 11 1000 381 300
Holothuroidea 2 666 69 60
station no W-3-1 Date 31 Jul 85 Sample size 3 (CONTINUED)
Number of taxa= 86
Mean number per sample 84633 Standard deviation (SD) 30596
Mean number m^ 881597 SD m^ 318713
H= 310 SDV= 067 SR=1097 J= 048
E-27
Station no W-3-2 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 10 1000 347 134
Leitoscoloplos puaettensis 20 1000 694 216
Phylo (Orbinia) felix 1 333 34 60
Aricidea suecica 64 333 2222 3849
Aricidea nr A suecica 39 666 1354 1469
Minuspio cirrifera 1 333 34 60
Prionospio cirrifera 9 333 312 541
Spiophanes berkelevorum 7 1000 243 60
Spiophanes bombyx 332 1000 11527 4221
Maqelona spp 3 333 104 180
Maaelona sacculata 2 333 69 120
Macrelona hartmanae 5 666 173 216
Chaetozone setosa 31 1000 1076 524
Tharyx multifilis 1 333 34 60
Decamastus aracilis 4 666 138 120
Heteromastus filobranchus 1 333 34 60
Heteromastus filiformis 10 333 347 601
Mediomastus ambiseta 1 333 34 60
Medioraastus sp indet 3 666 104 104
Notomastus lineatus 2 333 69 120 CO
Haldanidae 1 333 34 60
Euclymene zonalis 8 666 277 394
Axiothella rubrocincta 5 333 173 300
Phyllodoce hartmanae 4 666 138 120
Anaitides hartmanae 5 333 173 300
Phyllodoce lonaipes 3 333 104 180
Anaitides multipapillata 4 1000 138 60
Eteone spp 1 333 34 60
Eteone sp juvenile 1 333 34 60
Eteone californica 666 159
Pholoe minuta 7 666 243 334
Sthenalais tertiaalabra 1 333 34 60
Thalenessa spinosa 3 666 104 104
Pilargidae 1 333 34 60
Autolytus sp 333 69 120
Nereis spp 1 333 34 60
Glycera convoluta 1 333 34 60
Glycinde armigera 15 1000 520 130
Glycinde sp juvenile 1 333 34 60
Glycinde picta 1 333 34 60
Goniada maculata 5 666 173 216
Nephtys caecoides 5 666 173 159
Wephtys spp 15 333 520 902
Nephtys sp juvenile 39 666 1354 1263
Onuphis iridescens 1 333 34 60
E-28
Station no W-3-2 Date 31 Jul 85 Sample size 3
(CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Onuphis eleaans 82 666 2847 2468
Onuphis sp indet 63 666 2187 2933
Lumbrineris spp 3 333 104 180
Lumbrineris bicirrata 8 1000 277 120
Lumbrineris luti 1 333 34 60
Lumbrineris sp indet 3 666 104 104
Galathowenia nr G oculata 2 666 69 60
Mvriochele heeri 27 333 937 1623
Owenia collaris 49 333 1701 2946
Owenia fusiformis 1597 1000 55451 16672 Pectinaria spp 3 333 104 180
Ampharetidae 4 333 138 240
Ampharete acutifrons 4 666 138 159
Polycirrus spp 1 333 34 60
Polvcirrus sp complex 1 333 34 60
Amaeana occidentalis 2 333 69 120
Sabellidae 4 333 138 240
Gastropoda 4 666 138 120
Bivalvia 64 666 2222 2097
Yoldia scissurata 1 333 34 60
Modiolus spp 7 333 243 421
Lucinoma annulata 14 333 486 842
Macoma elimata 1 333 34 60
Hacoma expansa 16 333 555 962
Tellina modesta 1 333 34 60
Siliaua sloati 5 333 173 300
Scaphopoda 1 333 34 60
Dentalium spp 4 333 138 240
Copepoda 7 333 243 421
Ostracoda sp A 43 1000 1493 2316
Ostracoda sp B 98 1000 3402 3505
Nebalia sp 1 333 34 60
Campvlaspis 1 333 34 60
Diastylis sp iuvenile 47 666 1631 2648
Diastylopsis dawsoni 112 1000 3888 3603
Leucon subnasica 2 666 69 60
Isopoda 2 333 69 120
Synidotea sp 1 333 34 60
Pleurogonium rubicundum 2 333 69 120
Ampelisca a3assjzi 12 333 416 721
Ampelisca macrocephala 15 666 520 520
Ampelisca spp juvenile 3 333 104 130
Annisogammarus pugettensis 14 333 486 342
Eohaustorius spp 3 666 104 104
Lysianassidae 1 333 34 60
E-29
station no W-3-2 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation () m^ m
Lysianassidae juvenile 2 333 69 120
Photis parvidens 4 333 138 240
Photis sp juvenile 11 666 381 421
Photis macinernevi 33 333 1145 1984
Protomedeia prudens 6 666 208 180
Protomedeia spp juvenile 5 333 173 300
Cheriphotis sp 5 333 173 300
Rhepoxynius spp 37 666 1284 1113
Rhepoxvnius dabouis 14 666 486 433
Rhepoxynius abronius 7 1000 243 120
Rhepoxynius lucubrans 50 333 1736 3007
Parapleustes den 2 333 69 120
Caprellidae 3 333 104 180
Paguridae 1 333 34 60
Pinnixa sp megalopa 4 666 138 120
Pinnixa occidentalis 4 333 138 240
Ophiuroidea 8 666 277 240
Amphiodia spp 7 333 243 421
Number of taxa= 108
Nean number per sample 107300 Standard deviation (SD) 36181
Mean numberm^ 1117708 SDm2 376889
H= 355 SDV= 074 SR=1325 J= 053
E-30
Station no W-3-3 Date 31 Jul 85 Sample size 3
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Nemertinea 3 1000 104 00
Echiura 1 333 34 60
Leitoscoloplos pucrettensis 12 1000 416 104
Orbinia (Phylo) felix 7 666 243 262
Scoloplos acmeceps 1 333 34 60
Aricidea nr A suecica 61 1000 2118 492
Paraonella platvbranchia 1 333 34 60
Boccardia pugettensis 1 333 34 60
Laonice cirrata 1 333 34 60
Prionospio cirrifera 7 666 243 262
Spiophanes berkelevorum 27 1000 937 375
Spiophanes bombvx 369 1000 12812 4864
Magelona sacculata 1 333 34 60
Magelona hartmanae 2 333 69 120
Chaetozone setosa 30 1000 1041 541
Mediomastus ambiseta 1 333 34 60
Mediomastus californiensis 2 666 69 60
Notomastus lineatus 2 666 69 60
Euclvmene zonalis 19 1000 659 159
Scalibregma inflatum 2 333 69 120
Phvllodoce hartmanae 1 333 34 60
Phvllodoce mucosa 1 333 34 60
Eteone californica 6 666 208 275
Aphrodite iaponica 1 333 34 60
Pholoe minuta 2 333 69 120
Thalenessa spinosa 2 333 69 120
Gyptis brevipalpa 1 333 34 60
Nereis zonata 2 333 69 120
Glycinde armigera 10 1000 347 262
Goniada maculata 5 1000 173 60
Nephtys caecoides 31 1000 1076 365
Nephtys sp iuvenile 20 1000 694 313
Onuphis iridescens 2 666 69 60
Onuphis sp indet 65 1000 2256 1421
Lumbrineris bicirrata 3 1000 277 216
Galathowenia nr G oculata 5 666 173 159
Owenia fusiformis 392 1000 13611 10585
Ampharete acutifrons 3 666 104 104
Helinna elisabethae 1 333 34 60
Helinna sp juvenile 2 666 69 60
Terebellidae sp iuvenile 1 333 34 60
Pplycirrus sp complex 12 1000 416 180
Euchone nr E hancocki 3 666 104 104
Gastropoda 7 1000 243 120
Bivalvia 45 1000 1562 541
E-31
station no W-3-3 Date 31 Jul 85 Sample size (CONTINUED)
Taxa Total Frequency of Mean Stand
number occurrence number deviat
() m^ m^
Scaphopoda 3 666 104 104
Copepoda 25 333 868 1503
Ostracoda sp A 76 1000 2638 492
Leucon subnasica 1 333 34 60
Anthuridae 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Ampelisca aaassizi 6 1000 208 104
Ampelisca macrocephala 16 1000 555 334
Lysianassidae 3 666 104 104
Svnchelidium shoemakeri 1 333 34 60
Photis brevipes 3 666 104 104
Photis sp iuvenile 18 1000 625 208
Protomedeia spp juvenile 2 333 69 120
Ischvrocerus peaalops 1 333 34 60
Rhepoxynius spp 19 1000 659 159
Rhepoxvnius dabouis 24 1000 833 312
Rhepoxynius abronius 10 1000 347 240
Pleustidae 2 666 69 60
Dulichia monacantha 4 666 138 120
Dulichia sp juvenile 2 333 69 120
Pinnixa sp meaalopa 7 666 243 334
Brachyura 1 333 34 60
Ophiuroidea 8 1000 277 159
Holothuroidea 1 333 34 60
Other 1 333 34 60
Number of taxa= 70
Mean number per sample 47133 Standard deviation (SD) 15314
Mean numberm^ 490972 SDm^ 159516
H= 379 SDV= 084 SR= 951 J= 062
E-32
3Station no W-3-4 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence1 number deviation
() m^ m^
Nemertinea 11 1000 381 120
Leitoscolonlos nuaettensis 14 1000 486 492
Phylo (Orbinia) felix 5 1000 173 120
Aricidea nr A suecica 40 1000 1388 469
Laonice cirrata 1 333 34 60
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 3 666 104 104
Scolelepis spp 1 333 34 60
Spio filicornis 2 333 69 120
Spiophanes berkelevorum 12 1000 416 00
Spiophanes bombyx 532 1000 18472 6056
Maaelona hartmanae 4 1000 138 60
Chaetozone setosa 45 1000 1562 580
Tharyx secundus 1 333 34 60
Decamastus aracilis 3 333 104 180
Mediomastus ambiseta 1 333 34 60
Notomastus lineatus 7 1000 243 240
Euclymene zonalis 24 1000 833 275
Travisia sp indet 1 333 34 60
Phyllodoce hartmanae 2 666 69 60
Phyllodoce lonaipes 2 666 69 60
Eteone sp juvenile 1 333 34 60
Eteone californica 4 666 138 120
Pholoe minuta 2 666 69 60
Sthenalais tertiaalabra 3 666 104 104
Thalenessa spinosa 2 666 69 60
Gyptis brevipalpa 2 333 69 120
Glycinde armigera 12 1000 416 208
Goniada maculata 7 1000 243 120
Nephtys caecoides 35 1000 1215 767
Nephtys sp juvenile 10 1000 347 60
Onuphis iridescens 15 1000 520 454
Onuphis sp indet 58 1000 2013 1829
Lumbrineris bicirrata 3 666 104 104
Lumbrineris cruzensis 3 666 104 104
Notocirrus californiensis 1 333 34 60
Galathowenia nr G oculata 7 1000 243 159
Ovfenia fusiformis 232 1000 3055 4030
Ampharete acutifrons 1 333 34 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
^lycirr^^s sp complex 11 1000 381 159
Chone sp indet 1 333 34 60
Gastropoda 1 333 34 60
Bivalvia 49 1000 1701 1000
E-33
station no W-3-4 Date 30 Jul 86 Sample size (CONTINUED)
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Copepoda 39 1000 1354 1804
Ostracoda sp A 69 1000 2395 1623
Ostracoda sp B 1 333 34 60
Diastvlopsis dawsoni 2 666 69 60
Hemilamprops sp 2 333 69 120
Camplvapsis sp 1 333 34 60
Edotea sublittoralis 1 333 34 60
Pleuroaonium rubicundum 1 333 34 60
Bathvcopea daltonae 1 333 34 60
Ampelisca aaassizi 6 333 208 360
Ampelisca macrocephala 16 1000 555 216
Ampelisca spp juvenile 22 666 763 781
Eohaustorius spp 10 1000 347 262
Eohaustorius sawyeri 2 333 69 120
Lysianassidae 1 333 34 60
Anonyx spp () 2 333 69 120
Corophium sp juvenile 1 333 34 60
Photis brevipes 7 666 243 334
Photis sp juvenile 7 666 243 334
Protomedeia prudens 333 173 300
Protomedeia spp juvenile 3 666 104 104
Rhepoxynius spp 21 1000 729 312
Rhepoxynius dabouis 16 1000 555 433
Rhepoxynius abronius 7 1000 243 60
Dulichia sp juvenile 1 333 34 60
Caprellidae 1 333 34 60
Nebalia puaettensis 2 666 69 60
Decapoda larvae 2 666 69 60
Pinnixa sp megalopa 3 666 104 104
Ophiuroidea 8 1000 277 159
Holothuroidea 2 666 69 60
Number of taxa= 76
Mean number per sample 47867 Standard deviation (SD) 15519
Mean numberm^ 498611 SDm^ 161658
H= 384 SDy= 083 SR=1032 J= 061
E-34
3Station no W-3-5 Date 30 Jul 85 Sample size
Taxa Total Frequency of Mean Standard
number occurrence number deviation
() m^ m^
Leitoscoloplos puaettensis 10 1000 347 60
Phylo (Orbinia) felix 14 1000 486 159
Aricidea nr A suecica 65 1000 2256 1207
Polvdora caullervi 1 333 34 60
Prionospio cirrifera 2 666 69 60
Spio filicornis 5 666 173 216
Spiophanes berkeleyorum 11 666 381 421
Spiophanes bombyx 445 1000 15451 1027
Maaelona sacculata 2 666 69 60
Maaelona hartmanae 12 1000 416 275
Chaetozone setosa 46 1000 1597 492
Tharyx multifilis 1 333 34 60
Decamastus crracilis 1 333 34 60
Notomastus lineatus 2 666 69 60
Maldanidae sp indet 2 333 69 120
Maldanidae 1 333 34 60
Euclymene zonalis 25 1000 868 365
Phyllodoce hartmanae 5 1000 173 60
Phyllodoce sp juveniles 1 333 34 60
Eteone californica 3 666 104 104
Pholoe minuta 3 1000 104 00
Sthenalais tertiaplabra 2 666 69 60
Thalenessa spinosa 1 333 34 60
Gyptis brevipalpa 1 333 34 60
Glycinde armiaera 6 1000 208 180
Goniada maculata 3 333 104 180
Nephtys caecoides 15 1000 520 275
Nephtys sp juvenile 12 1000 416 180
Onuphis iridescens 41 666 1423 1359
Onuphis sp indet 34 1000 1180 513
Lumbrineris bicirrata 2 333 69 120
Lumbrineris latreilli 1 333 34 60
Lumbrineris sp indet 2 333 69 120
Galathowenia nr G oculata 8 666 277 240
Owenia fusiformis 467 1000 16215 10554
Ampharete acutifrons 4 1000 138 60
Melinna elisabethae 1 333 34 60
Melinna sp juvenile 1 333 34 60
Terebellidae 1 333 34 50
Polycirrus sp complex 8 1000 277 60
Gastropoda 17 666 590 534
Bivalvia 28 666 972 898
Copepoda 1 333 34 60
Ostracoda sp A 56 1000 1944 1744
Ostracoda sp B 2 666 69 60
E-35
3Station no W-3-5 Date 30 Jul 85 Sample size (CONTINUED)
Taxa
Diastvlopsis spp
Diastvlopsis dawsoni
Svnidotea sp
Munna sp
Pleuroaonium rubicundum
Bathycopea daltonae
Gammaridea
Ampelisca agassizi
Ampelisca macrocephala
Eohaustorius spp
Eohaustorius sawveri
Lysianassidae Anonvx spp ()
Synchelidium shoemakeri
Photis parvidens
Photis sp juvenile
Protomedeia spp juvenile
Rhepoxvnius spp
Rhepoxynius dabouis
Rhepoxynius abronius
Sympleustes subglaber
Caprellidae Nebalia pugettensis
Ophiuroidea
Number of taxa= 69
Mean number per sample
2 Mean numberm 518056
H= 351 SDV= 081
Total
number
1
2
2
1
1
1
1
3
14
4
1
1
1
1
12
12
10
19
6
10
6
5
1
6
49733
SDra
SR= 931
Frequency of Mean Standard
occurrence number deviation
() m^ m^
333 34 60
333 69 120
333 69 120
333 34 60
333 34 60
333 34 60
333 34 60
333 104 180
1000 486 120
666 138 120
333 34 60
333 34 60
333 34 60
333 34 60
1000 416 375
666 416 633
1000 347 120
1000 659 262
1000 208 00
1000 347 60
666 208 275
666 173 159
333 34 60
1000 208 104
Standard deviation (SD) 16333
170656
J= 057
E-36
Appendix Table F Benthic Invertebrate taxa collected off Willapa Bay Washington 30-32 July 1985
Cnidaria
Anthozoa
Nemertinea
Annelida
Polychaeta ^ Orbiniidae
Leitoscoloplos pugettensis (Johnson 1910)
Phylo felix Kingberg 1866 Scoloplos acmeceps Chamberlin 1919
Paraonidae
Aricidea nr suecica Aricidea suecica (Elaison 1920)
Paraonella platybranchia (Hartman 1961)
Spionidae
Boccardia pugettensis Blake 1979
Laonice cirrata (Sars 1851)
Minuspio cirrifera (Wiren 1883)
Polydora caulleryi Mesnil 1897
Polydora socialis (Schmarda 1861)
Prionospio cirriferalighti
Scolelepis spp Spio filicornis Muller 1776
Spiophanes berkeleyorum Pettibone 1962
Spiophanes bombyx (Claparede 1870) Megelonidae
Megelona hartmanae Jones 1978 ()
Megelona longicornishartmanae
Megelona sacculata Hartman 1961
Megelona spp ^ Cirratulidae
Chaetozone setosa Malmgren 1867 Cirratulidae sp A Tharyx multifilis Moore 1909
Tharyx secundus Banse and Hobson 1968
Tharyx sp indeterminate ^ Capitellidae
Decamastus gracilis Hartman 1963
Heteromastus filobranchus Berkeley and Berkeley 1932 Heteromastus filiformis (Claparede 1864) Heteromastus spp
Mediomastus ambiseta (Hartman 1947)
Med^-Otnastus californiensis Hartman 1944
Mediomastus sp Indeterminate
Notomastus lineatus Claparede 1870 Maldanidae
Axiothella rubrocincta (Johnson 1901)
Euclymene zonalis (Verrill 1874)
Maldanidae (unidentified)
Praxillella affinis (Sars 1872)
F-1
Ophelildae Travisla sp Indeterminate
Scalibregmidae Scalibregma Inflatum Rathke 18A3
Phyllodocidae
Anaitides groenlandica (Oersted 18A3)
Anaitides hartmanae (Blake and Walton 1977)
Eteone californica Hartman 1936
Eteone fauchaldi Kravitz and Jones 1979
Eteone longa (Fabricius 1780)
Eteone spp
Eulalia (Eumida) sanguinea Orsted 18A3
Paranaitides polynoides (Moore 1909)
Phyllodocidae (unidentified) Phyllodoce (Anaitides) longipes (Kinberg 1866)
Phyllodoce mucosa Orsted 1843
Phyllodoce multiseriata Rioja 1941 Aphroditidae
Aphrodite japonica Marenzeller 1879
Aphrodite negligens Moore 1905
Polynoidae
Halosydna brevisetosa Kinberg 1855 Halosydna spp Lepidonotus spp Polynoidae spp (juveniles) Tenoia kitsapensis Nichols 1969
Sigalionidae
Pholoe minuta (Fabricius 1780)
Sthenalais tertiaglabra Moore 1910 Thalenessa spinosa (Hartman 1939)
Hesionidae
Heteropodarke heteromorpha Hartman-Schroeder 1962
Podarkeopsis (Gyptis) brevipalpa (Hartman-Schroeder 1959) Pilargidae Syllidae
Autolytus sp Nereidae
Nereis zonata Malmgren 1867
Nereis spp
Glyceridae Glycera capitata Oersted 1843
Glycera convoluta Keferstein 1862 ltv
Glycera tenuis Hartman 1944
Glycera spp (juveniles) Goniadidae
Glycinde armigera Moore 1911
Glycinde picta Berkeley 1927 Glycinde sp-
Goniada maculata Oersted 1843
Goniada sp
Nephtyidae Nephtys caeca (Fabricius 1780)
Nephtys caecoides Hartman 1938
Nephtys californiensis Hartman 1938 (5raquov
Nephtys ciliata (0 F Muller 1776)
F-2
(Ms
^
^
^
^
^
^
Nephtys longosetosa Oersted 1843 Nephtys spp
Onuphldae Dlopatra ornata Moore 1911 Onuphls elegans (Johnson 1901) Onuphls (Nothria)Irldescens (Johnson 1901) Onuphls sp
Lumbrlnerldae
LumbrInerIs blclrrata Treadwell 1929
LumbrInerls callfornlensls Hartman 1944
LumbrInerIs cruzensls Hartman 1944
LumbrInerls latrellll Audouln and Mllne-Edwards 1843 Lumbrlnerls lut1 Berkeley and Berkeley 1945 Lumbrlnerls spp
Arabellldae
Notoclrrus callfornlensls Hartman 1968
Owenlldae
Galathowenla (Myrlochele) oculata Zaks 1923 Myrlochele heerl Malmgren 1867 Owenla collarls Hartman 1955
Owenla fuslformls delle Chlaje 1841 PectInarlldae
Fectlnarla callfornlensls Hartman 1941
Fectlnarla sp
AmpharetIdae
Ampharete acutIfrons (Grube 1860) Ampharete spp
Mlllnna ellsabethae Mclntosh 1922
Mlllnna sp (juvenile) Terebellldae
Amaeane occldentalls Hartman 1944
Flsta brevlbranchlata Moore 1923
Folyclrrus spp
Folyclrrus sp complex Sabelllldae
Chone sp Euchone nr Ej^ hancockl Banse 1970
Archlannellda
Hlrudlnea
Mollusca
Gastropoda Cephalaspldae
ScaphandrIdae Cyllchna attonsa (Carpenter 1864)
Felecypoda
Nuculanldae
Yoldla sclssurata (Dall 1887)
Mytllldae Modlolus spp
LucInldae
Luclnoma annulate (Reeve 1850)
Telllnldae
Macoma elImata Dunnlll and Coan 1968
Macoma expanse Carpenter 1864 TellIna modesta (Carpenter 1864)
F-3
Telllna nuculoldes (Reeve 1854)
Psammobildae
Garl callfornlca (Conrad 1837)
Solenldae
Sillqua sloat1 Hertleln 1961
Scaphopoda Dentalium sppbull
Arthropoda Pycncgonldea
Crustacea
Leptostraca
Nebalildae
Neballa pugettensls (Clark 1932)
Nebalia spbull
Ostracoda
Ostracoda sp A Ostracoda sp B
Copepoda Cuntacea
Dlastylidae Dlastylls sp Dlastylopsls dawsonl Smith 1880 Diastylopsis spp
Oxyurostylls paciflea Zimmer 1936 Lampropidae
Hemilamprops callfornlca Zlmraer 1936
Leuconldae
Leucon subnaslca Given 1961
Nannastacldae
Camplyapsls spbull
Isopoda Anthurldae
Hallophasma gemlnata Menzles and Barnard 1959 Idoteldae
Edotea subllttoralls Menzles and Barnard 1959
Idotea sp
Synldotea blcusplda Owen 1939
Synldotea sp
Munnldae
Munna sp Munnogonlum tlllerae (Menzles and Barnard 1959)
Pleurogonlldae Pleurogonlum rublcundum G-O- Sars 1863
SpaeromatIdae
Bathycopea daltonae Menzles and Barnard 1959
laeroma oregonensIs (Dana 1854-1855) Amphlpoda
Ampellscldae Ampellsca agasslzl (Judd 1896)
AmpelIsca macrocephala Llljeborg 1851 AmpelIsca spp- (juveniles) AmpelIsca sp
^
^
^
F-4
_
^
^
^
^
^
Atylidae Atylus trldens (Alderman 1936)
Corophlldae Corophium spp
Gammaridae
Anisogammarus pugettensls (Dana 1853)
Haustoriidae
Eohaustorius sawyerl Bosworth 1973
Eohaustorius spp Isaeidae
Cheriphotls spbull Cheirimedeia sp Photis brevlpes Shoemaker 19A2 Photis macinerneyi Conlan 1983 Photis parvidons Conlan 1983
Photis spp (juveniles) Protomedeia prudens Barnard 1966 Protomedeia spp (juveniles)
Ischyroceridae Ischyrocerus pegalops Barnard 1962 Ischyrocerus sp Ischyrocerus spp (juveniles)
Lysianassidae
Oedicerotidae
Synchelidium shoemakeri Mills 1962
Synchelidium sp (juvenile) Phoxocephalidae
Foxiphalus spp (juveniles) Rhepoxynius abronlus (Barnard 1960) Rhepoxynius dabouis (Barnard 1960) Rhepoxynius lucubrans (Barnard 1960) Rhepoxynius spp
Pleustidae
Parapleustes den Barnard 1969
Sumpleustes subglaber Barnard and Given 1960 Podoceridae
Dulichia monacantha Metzger 1875 Dulichia spp
Caprellidae Decapoda
Decapoda larvae Natantia
Reptantia
Callianassidae
Callianassa californiensis Dana 185A
Paguridae
Pinnotheridae
Pinnixa occidentalis Rathbun 1893
Pinnixa spp raegalops
Porcellanidae
Echinodermata
Ophiuroidea Amphiodia sppbull
Holothuroidea
F-5