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THESIS 2016
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1
21
..
2-3
2
2-3
1 2
3-4
1
6-7
3
2
elytra (
elytron)
?
4
2
5
6
7
: Chalcosoma caucasus
: Giant rhinoceros beetle
120 .
3
8
4
9
10
5
11
12
6
..
13
14
7
17
18
Dynastes hercules
Scarabaeidae
190
65-70
16
(Hercules beetle)
19
150
60-65
(Neptune beetle)
20
100-120
60-65
(Elephant beetle)
70-100
87-100 70-75
(goliath beetle)
21
130
(Causcasus three-horned beetle)
22
8
23
24
(compound
eyes) 1
1-2
11
(scape)
(pedicel)
3-11 (flagellum)
(mandible)
(stag)
stag beetle
(labrum)
(labium)
(head)
25
3
(prothorax)
(pronotum)
(meso-
thorax)
1
2
7 8
(tergite)
(sternite)
,
, , , 5
(thorax)
(abdomen)
1
2
3
26
(Complete metamorphosis)
(prolegs)
Eruciform
4
Sca rabae i f o rm (Wh i te grub- l i ke )
C
3
27
Vermiform (maggot-like)
(Cerambycidae)
(round-headed bor-
er) (Buprestidae)
(flat-headed borer)
Eruciform (Scarabaeidae)
(Lucanidae)
(Passalidae) Campodeiform (diplu-
ran-like)
(Diplura) (Thysanura)
3
(Meloidae)
(Coccinellidae)
Elateriform (wireworm-like)
(Elateridae)
(Tenebrionidae)
28
4
1. (egg)
2. (larva)
3. (pupa)
(Exatate pupa)
4. (adult)
29
(predators)
Coccinellidae
Monochilus sexmaculatus
3
30
9
31
32
(Flighting beetle)
33
12
3
1
3
3
300
34
""
35
100 -120
15 20
C
(Scarabaeiform type)
30
6
8
30 35
2 5
36
10
37
38
11
39
40
41
12
42
2
(Dynastinae,
Scarabaeidae)
(Lucanidae)
48-70
1
2
2-3
(Eupatorus gracilicornis)
43
2.5-4.0
10.5
7.5
1
38.5-79.5
42.2-54.0
()(Japanese rhinoceros beetle)
(Hexarthrius parryi)
(Giraffe stag beetle)
44
1
15-20
1.
2-3
2.
10-20
3.
4.
45
1.
1:1 1:2
4-5
2.
5
15-20
()
3.
45-60
5-6
4.
5.
46
47
48