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An invertebrate survey of Penn Common
(concentrating on the wet habitats, especially the springs and flushes)
to cover those species and groups that help evaluate its biological
importance and help to inform its habitat management under
the Higher Level Scheme agreement.
Peter Boardman, Ian Cheeseborough, Nigel Jones, Richard Wright
November 2011
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 2
INDEX
i. Summary …………………………………………………………………………….. 3
1. Rationale …………………………………………………………………………………………. 4
2. Methodology…………………………………………………………………………………… 5
3. Results – overview………………………………………………………………………….. 5
4. Results – analysis of invertebrate communities………………………………. 9
5. Results – summary of ISIS analysis…………………………………………………… 10
6. Management Suggestions……………………………………………………………….. 11
7. Discussion……………………………………………………………………………………….. 13
ii. References………………………………………………………………………………. 15
iii. Acknowledgements…………………………………………………………………. 15
Appendix 1 – Full survey data for Penn Common golf course ……………………. 16
Appendix 2 – ISIS outputs for Penn Common …………………………………………….. 31
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
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Summary
An invertebrate survey of Penn Common, near Wolverhampton, recorded 457 species of invertebrates
during 2011.
Of these, the vast majority were common species with only 7 species recorded of national or regional
importance. These were the flies; Spania nigra, Syntormon macula, Sphaerophoria taeniata, and Herina
frondescentiae. The weevil Trachyphloeus asperatus, the spider-hunting wasp Anoplius caviventris, and the
solitary wasp Tiphia minuta.
24 species were noted as new to Staffordshire (V.C. 39).
Data was put through ISIS software to identify species assemblages for the site. As a whole site, significant
species assemblage types (SATs) were identified for scrub edge, rich flower resource, and bark & sapwood
decay assemblages. The most significant broad assemblage type (BATs) noted was that for flowing water
suggesting the wetland habitats are important at the site. Heathland communities were not significant
across the whole site data.
When data was split down into habitat areas the species assemblage types produced significant results for
scrub-heath and moorland species assemblage amongst the dry acid grassland habitats, and the bark and
sapwood decay assemblages amongst the aculeate habitats.
The calcareous flush was found to be of little significance currently.
Penn Common was found to be regionally important for its aculeate hymenoptera assemblage, though much
of this is associated with dead-wood rather than heathland species.
Management suggestions were made to increase the suitability of habitat for heathland hymenoptera,
amongst other habitats across the site.
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
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1. Rationale
1.1 An invertebrate survey was carried out on Penn Common (SO898945), near Wolverhampton, to identify the
most important invertebrate communities at the site and examine how they compare to other similar sites
in the area (Staffordshire and West Midlands primarily but also east Shropshire and north Worcestershire).
1.2 Outputs were to examine how the invertebrate communities can be conserved and, where possible,
enhanced (under management through the Higher Level Stewardship (HLS) agreement).
1.3 Penn Common has historically been recognised as an important site in terms of nature conservation with
historical botanical records suggesting important habitats (particularly base rich flushes and heathland) for
invertebrates at the site. It was noted that grazing had recently ceased but more up to date information
regarding habitat quality and suitability for invertebrate assemblages was lacking, hence the need to assess
the site through survey work.
1.4 The site hosts Penn Common Golf Course, where golf has been played since 1893 (see map 1), and is
intersected roughly down the centre on a north / south plane by a minor road. It sits to the south of Penn, a
suburb of the major conurbation of Wolverhampton, and to the west of Sedgley, and is located in
Staffordshire (V.C. 39).
Map 1 – Penn Common Golf Course (www.maps.google.co.uk.)
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
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2. Methodology
2.1 The survey methodology followed that laid out in ‘Surveying terrestrial and freshwater invertebrates for
conservation evaluation’ (Drake et al, 2007) and results were analysed using the ISIS software, developed by
Natural England, to assess invertebrate communities at the site.
2.2 The surveyors used appropriate methodologies for their particular invertebrate interests; Pete Boardman &
Nigel Jones (diptera – timed sweeping over flushes, damp and marginal habitats, plus netting individual
species of interest, water traps, limited Malaise trapping), Ian Cheeseborough (aculeate hymenoptera –
netting individual species and observation of nesting aggregations and dead wood habitats, water traps,
limited Malaise trapping), Richard Wright (coleoptera – pitfall trapping and sweeping / beating heather and
other vegetation, hand searching).
2.3 Where possible other invertebrate and insect orders were recorded when encountered. These included the
Lepidoptera which were being surveyed by another contractor so little attention was given to the group
during our site visits.
2.4 Visits were timed to assess the peak phenology of the groups studied.
3. Results - overview
3.1 Survey visits were undertaken in April, May, June, July, September and October by the surveyors, after an
initial visit in early April accompanied by the course manager, Paul Mills from Penn Common Golf Course.
3.2 Survey work during 2011 was in some ways hampered by the very dry conditions that occurred through the
year. A hot, dry spring was followed by a generally dull, dry summer and the dry weather continued on into
autumn. Several of the wet or damp areas where survey work was focused were subject to desiccation and
at times insects were hard to come across. Nevertheless a total of 457 species were recorded at the site.
3.3 The table below details the breakdown of invertebrate orders recorded;
Order No of Species
Diptera (true flies) 220 Coleoptera (beetles) 82 Hymenoptera (bees, wasps and ants) 73 Aranae & Opiliones (spiders and harvestman) 33 Hemiptera (true bugs) 21 Lepidoptera (butterflies and moths) 15 Diplopoda (millipedes) 4 Odonata (damselflies and dragonflies) 4 Isopoda (woodlice) 3 Orthoptera (grasshoppers and crickets) 2 Total - 457
Table 1 – summary of invertebrate orders recorded at Penn Common during the 2011 survey
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 6
3.4 Of the 457 species recorded few were noted to be of conservation importance except for the following 7
species;
3.5 Diptera (Spaniidae) Spania nigra Meigen, 1830 Liverwort Snipefly. This species is listed as Notable but being
small is often overlooked. Larvae leaf mine species of liverworts and Pellia neesia has been noted to host the
fly (Stubbs & Drake, 2001). It has a widespread distribution within the United Kingdom
(www.data.nbn.org.uk) and appears here at its first Staffordshire site (Jukes, 2005). It has been recorded in
Shropshire from dingle woodland near Ironbridge Gorge and at three sites around the Long Mynd in south
Shropshire (Shropshire Ecological Data Network pers. comm.)
3.6 Diptera (Dolichopodidae) Syntormon macula Parent, 1927. This uncommon long-legged fly is often
associated with dingle woodlands or damp places and has a mostly south-western distribution. It was
originally designated a Red Data Book species but is now recognised as more commonly occurring, so was
given the lower risk (near threatened) status to reflect this (Falk & Crossley, 2005). It appears here at its first
Staffordshire site (Jukes, 2005). It is known from three dingle woodlands in Shropshire (Boardman, 2005 &
Shropshire Ecological Data Network pers. comm.).
3.7 Diptera (Syrphidae) Sphaerophoria taeniata (Meigen, 1822). This hoverfly has a mostly southern distribution
with Kent across to Somerset seeing the majority of records. It is uncommon outside of this range
(www.hoverfly.org.uk), and it appears to be heading north and is recorded here at its first Staffordshire
location (Jukes, 2005).
3.8 Diptera (Ulidiidae) Herina frondescentiae (Linnaeus, 1758). This non-tephritid picture-wing fly has a
widespread distribution nationally but can be considered local in the Midlands. It is associated with damp
grasslands and appears new to Staffordshire at this site (Jukes, 2005).
Herina frondescentiae (photo by Nigel Jones)
3.9 Coleoptera (Curculionidae) Trachyphloeus asperatus Boheman, 1843. This weevil is associated with lowland
calcareous grassland and has a widespread distribution in England, with mainly a maritime distribution in
Wales. It is classed as Notable A, and is the first Staffordshire record since 1907 (Slawson & Brown, 2009).
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 7
3.10 Hymenoptera (Pompilidae) Anoplius caviventris (Aurivillius, 1907). This spider-hunting wasp has a mostly
southern distribution (www.data.nbn.org) and is classed as Nationally Notable B by Falk (1991) but ‘very
rare’ by Michael Archer in his more recent review. It appears here as new to Staffordshire (Webb, Bloxham
and Slawson, 2005). It is known from one site in Worcestershire (Trevis, 2004) and a single site in Shropshire
(Ian Cheeseborough pers. comm. & Shropshire Ecological Data Network pers. comm.).
3.11 Hymenoptera (Tiphiidae) Tiphia minuta Vander Linden, 1827. This solitary wasp is a parasite of dung beetles
(Scarabaeidae) and is considered to be regionally scarce (Jukes, 2004).
3.12 As well as these important species several others were noted as new to Staffordshire when checked against
current checklists;
3.13 Diptera;
craneflies (Limoniidae); Dicranomyia affinis, Eloeophila submarmorata, Erioptera squalida.
long-legged flies (Dolichopodidae); Hercostomus parvilamellatus, Teuchophorus calcaratus, T.
monacanthus, T. nigricosta, T. simplex,
snail-killing flies (Sciomyzidae); Pherbellia schoenherri, Tetranocera fuscinervis.
flat-footed flies (Platypezidae); Agathomyia antennata
druid flies (Clusiidae); Clusiodes gentilis
stalk flies (Psilidae); Chamaepsila rosae
shore flies (Ephydridae); Ditrichophora calceata
big-headed flies (Pipunculidae); Jassidophaga villosa
lauxaniid flies (Lauxaniidae); Minettia fasciata
parasitic flies (Tachinidae); Lydella stabulans, Vibrissina debilitata
dung flies (Scathophagidae); Nanna flavipes
anthomyid flies (Anthomyiidae); Anthomyia procellaris
muscid flies (Muscidae); Coenosia infantula
3.14 Hemiptera;
leatherbugs (Coreidae); Coreus marginatus (Dock Bug)
3.15 Coleoptera;
ground beetle (Carabidae); Agonum emarginatum
3.16 Hymenoptera;
solitary wasp (Sphecidae); Psenulus concolor
3.17 As well as assessing the site as a whole, it was broken down to the main areas of interest as detailed in the
tender document, to begin to assess the invertebrate communities present at the site.
3.18 They were as follows (see map 2);
1 – damp grassland, pond etc at SO89709483
2 – damp, rushy grassland with scrub at SO89629479
3 – species-rich dry grassland at SO895294444
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
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4 – acid grassland / heather at SO89459440
5 – wet woodland / flush with calcareous influence at SO89599442
6 – acid grassland / heather at SO89799476
7 – damp woodland at SO90309478
8 – marsh / wet woodland and wet grassland at SO90379448
8a* – open scrub / woodland with bracken at SO903944
9 – woodland flush / pond / open ditch at SO90259456
9a* –cleared patch with dead wood at SO90269452
10 – flower-rich clearing at SO90269452
11 – damp woodland at SO89639441
12 – eastern footpath / sheltered hedgerow at SO90349456
3.19 Other records away from these areas were noted as ‘general site records’ and given a site centroid grid
reference of SO898945.
3.20 Areas 8a and 9a were separated from areas 8 and 9 respectively to take into account good aculeate
hymenoptera habitat including dead wood, which would otherwise potentially biased species assemblage
analysis in the larger parent areas.
3.21 All records are detailed in Appendix of this report within Table 2.
Map two – compartments survey during Penn Common survey 2011
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
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4. Results - Analysis of Communities
4.1 Data was put into the ISIS spreadsheet to identify Species Assembly Types (SATs) and Broad Assemblage
Types (BATs) that may exist at Penn Common. It should be noted that approximately 30 species could not be
used in the ISIS analysis as they registered an “error”. These species are ones excluded from ISIS as they are
ones where sufficient information is known about them, are vagrants, or their habitats do not fit into the
classification.
4.2 All SATs and BATs tables used in this analysis are produced in the Appendix of this report.
4.3 Whole site data - ISIS recognised 11 SATs of which three (scrub edge, rich flower resource and bark &
sapwood decay) suggested strong species assemblies (favourable condition in CSM). Less strong SATs were
those from the dung, open short sward, shingle banks, seepage, reed-fen and pools, scrub-heath and
moorland, bare sand and chalk, and heartwood decay assemblies.
4.4 This suggests that the site as a whole has more value for its scrub / woodland edge, associated dead wood,
and floral resources than for its heathland or wetland features. This may represent a decline in the state of
the habitat but this can only be suggested, rather than proven, without previous evidence of invertebrate
survey work at the site and is based upon the perception that features such as the calcareous flush and
heathland were perhaps once in better condition due to the historical botanical records that exist.
4.5 Whole site data - 8 BATs were identified, of which the flowing water broad assembly was noted as
favourable (in terms of CSM) and achieved the highest rarity score (154). The grassland and scrub matrix
achieved the highest BAT species richness score of 154. These figures suggest that within the site as a whole
the grassland and scrub matrix and the flushes are key features. Other BATs identified were; unshaded early
successional mosaic, permanent wet mire, wood decay, shaded field and ground layer, mineral marsh and
open water, arboreal canopy.
4.6 Calcareous flush at SO89599442 (area 5) - ISIS only recognised a single SAT from this data, a single species
of the shingle bank assembly.
4.7 Calcareous flush at SO89599442 (area 5) - 5 BATs were identified; grassland and scrub matrix, flowing
water, mineral marsh and open water, shaded field and ground layer, and permanent wet mire. No features
of either SATs or BATs reached favourable levels and this suggests that this flush is of limited value for
invertebrates in its current condition, with few specialists identified. It should be noted that part of this area
of the site has only recently been cleared of scrub.
4.8 Wet habitats in the east of the site (areas 8 and 9 – not including 8a and 9a) - ISIS recognised 8 SATs from
this area but none of them were strong communities. They were; seepage, open short sward, scrub edge,
reed-fen and pools, bark and sapwood decay, rich flower resource, scrub-heath and moorland, bare sand
and chalk. 8 BATs were recognised with the flowing water broad assembly noted as favourable.
4.9 Wet habitats in the east of the site (areas 8 and 9 – not including 8a and 9a) - BATs recognised were;
grassland and scrub matrix, permanent wet mire, flowing water, mineral marsh and open water, shaded
field and ground layer, unshaded early successional mosaic, wood decay and arboreal canopy.
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
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4.10 This analysis suggests no strong species assemblages occur within this wetland area, though it does
recognise that flowing water is important within the broader assemblage of species and is a key feature of
the site.
4.11 Scrubby, flower rich and drier habitats (areas 8a, 9a and 10) - ISIS showed 5 SATs within these areas (mostly
targeted for aculeate hymenoptera) and one species assembly type stands out as favourable – the bark and
sapwood decay. We had already noted that there was a higher than expected list of dead wood aculeate
hymenoptera, considering tree cover is limited, and few old or veteran trees occur at the site, so it was
pleasing to see ISIS agreeing with our own interpretation. ISIS suggests 26 species within the bark and
sapwood decay assembly. Other SATs listed were; rich flower resource, dung, scrub edge and open short
sward.
4.12 Scrubby, flower rich and drier habitats (areas 8a, 9a and 10) - 5 BATs were identified with three standing
out; unshaded early successional mosaic, wood decay and grassland and scrub matrix. The two remaining
and less strongly represented BATs were arboreal canopy and mineral marsh and open water.
4.13 Dry acid grassland and heather (areas 3, 4 and 6) - ISIS listed 4 SATs within these areas (mostly targeted for
coleoptera) and the scrub heath and moorland assembly was noted as favourable with 10 species within the
assembly type. The remaining three SATs were bare sand and chalk, scrub edge, and open short sward.
4.14 Dry acid grassland and heather (areas 3, 4 and 6) - 7 BATs were identified with the grassland and scrub
matrix assembly standing out far above all others listed. The others were; unshaded early successional
mosaic, mineral marsh and open water, shaded field and ground layer, permanent wet mire, flowing water.
This suggests that this area of the site on the western side is the most important area of acid grassland and
scrubby heath, but bare ground and short sward assemblies are poorly represented.
4.15 Damp grassland (areas 1 and 2) - ISIS was unable to identify any SATs for this habitat.
4.16 Damp grassland (areas 1 and 2) - 5 BATs. Of these the strongest broad assembly was amongst grassland
and scrub matrix species. The permanent wet mire broad assembly was not overly strong but nevertheless
worthy of mention whilst the mineral marsh and open water, shaded field and ground layer, and flowing
water broad assemblies were poorly represented. This data suggests that damp grassland habitats at the
site of the least productivity for species assemblages on the site.
5. Results – Summary of ISIS analysis
5.1 Analysis of the species and broad species assemblages of Penn Common using ISIS suggests that the scrub
matrix assembly, with associated dead wood, flowing water, dry acid grassland and flower rich grassland are
the key current habitat types at the site for invertebrates.
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 11
5.2 Heathland containing good quality heather and bare open ground seems in too short a supply to host
important invertebrate assemblages at present levels. Aculeate hymenoptera assemblages are more reliant
upon dead wood than traditionally suitable heathland habitat at the site.
5.3 The calcareous flush in the east of the site is currently not important in terms of species assemblages;
through it does host three broad wetland assemblages and a scrub matrix assemblage. This suggests that as
the flush scrubbed up it became less important for wetland species, though with recent and continued
habitat management may revert to hold more characteristic wetland assemblages of note in the future.
5.4 Other wet habitats at the site were not viewed as strongly important in terms of individual species
assemblages but flowing water as a broad assembly was noted as important.
5.5 Damp grassland at the north of the site is of little importance for species assemblages, which may reflect
the lack of grazing at the site, or simply the dry weather from 2011.
6. Management Suggestions
6.1 Current management of habitats needs to be continued and increased to maintain a mosaic of shady wet,
open wet, flower-rich (low growing and tussocky) grassland, and heathland habitats with heather and bare
ground. For example the aculeate hymenoptera at Penn Common need nesting opportunities (sandy bare
ground or dead wood), and foraging/hunting grounds in proximity to these nesting sites. Keeping the mix of
bare ground, short flower-rich grassland, rank patches of vegetation, scrub and woodland is ideal.
6.2 Wet areas and flushes;
Probably too many of the wet areas are, or have been until recently, scrubbed up to the detriment of
wetland diptera and other groups. These areas should be rotationally cut as has been done in area 5.
The main passage of water from north of south along the eastern boundary of the site (area 8) is
canalised and the grass is mown right up to the edge. Though there is probably not too much that can be
done about the canalization given the proximity to the fairway, some grading down the stream banks (to
make them more gently sloped) and a more relaxed approach to grass cutting may provide more
marginal habitat in this area.
The pond at the bottom of this ditch is becoming choked with vegetation and would benefit from some
clearance.
6.3 Heathland;
The current managed heather areas on the site are just too small to be of much use as heathland
pockets, and are often just a couple of straggly heather plants in the middle of a fairway. Areas should
be expanded and could be seeded with heather seeds from a local source.
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
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More bare ground would benefit nesting aggregations of bees. Bare ground is often seen as an erosion
problem or unsightly on golf courses and seeded with a grass mixture at the first opportunity. Some
decent sized scrapes on marginal areas of the fairways (as large as practicably possible) would provide
potentially a great deal of bare ground across the course. This sort of approach has recently been
undertaken at Highgate Common for the rare solitary bee Andrena nigrispina. The scrapes were
observed on a recent site visit there by two of our surveyors. Several of these bee scrapes are needed
around the site to encourage nesting aggregations providing horizontal and vertical surfaces into which
bees and wasps can burrow. These can cover a variety of aspects, but are probably the single most
important habitat management tool for heathland bees.
Solitary bees Andrena barbilabris and Crossocerus quadrimaculatus were observed using the bunkers for
nesting but only around the unraked sides, therefore whilst bunkers are a key aspect for the golfers at
the site, parts of them need not be regulary raked to maintain more breeding habitat for solitary bees.
This may be a matter of education of golfers rather than management per se.
6.4 Grassland;
The amount of acid grassland and species rich grassland should be increased at the site. This could be
done by relaxing mowing regimes on the fairways. Instead of a standard lozenge shape of a golf hole, the
fairways could be altered so that more of an hour glass shape exists filtering golfers down the middle of
the fairway in areas less commonly used on each hole (for instance between the tee and where the
average drive from that tee lands).
Flower rich areas need cutting and raking, with the arisings removed to restrict nutrient input. Cutting of
flowering areas needs to be timed so that they are cut after flowering (not during) and with areas left for
invertebrates to over-winter in seed heads or tussocks.
Where thick areas of bracken persist, some rotational cutting to open up the structure would add
another dimension to the habitat.
6.5 Dead wood;
ISIS analysis identified dead wood as a key feature of aculeate hymenoptera at the site, therefore as well
as ensuring existing amounts of dead wood are retained, more dead wood should be left available for
aculeate hymenoptera and coleoptera to utilise.
Standing dead wood is more valuable than fallen dead wood. Often standing dead wood is removed as
it is deemed to be deleterious to health and safety, however reducing the height of dead wood by
removing large branches or the upper half of the tree whilst leaving the trunk still standing to maintain
safety is still leaving a valuable resource in place.
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
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Standing dead wood could be further improved by drilling small holes into it with a battery powered drill
and bit. This method has worked well at sites in Shropshire where it has been adopted and used by a
variety of dead wood nesting bees and wasps (Cheeseborough pers. comm.).
The existing dead wood left on the ground is very quickly covered with vegetation making it unsuitable
quickly unsuitable. Fallen dead wood should therefore be kept clear of vegetation or kept higher up of
the ground if this is a problem.
7. Discussion
7.1 Penn Common was not considered to be of high conservation status in its current state. The vast majority of
species encountered were common in terms of nature conservation interest, though some can be
considered heathland, or wetland specialists.
7.2 The strongest species assemblages represented those of scrub edge, flower-rich grassland, and dead wood
assemblages of aculeate hymenoptera. Had heathland habitat been in better condition the species
assemblages would have reflected this with higher scores in the scrub-heath and bare sand assemblages.
7.3 The calcareous flush was found to be of less importance for wetland species than had been hoped based
upon historical botanical records.
7.4 One surprise was the large amount of dead wood aculeate hymenoptera found, considering the lack of
established veteran trees.
7.5 Clearly the impact of a very popular local amenity such as the Golf Course undoubtedly has had an effect
upon the habitat, though this is not to say the impact of more considered habitat management could
improve the existing situation under the Higher Level Scheme.
7.6 It should be noted that our results were affected by the lack of rain through the 2011 season, which led to
the drying out of most of the “wet” areas on the site by the end of the 2011 recording season, and clearly
had a detrimental effect on the numbers of wetland diptera and coleoptera sampled.
7.7 In terms of the aculeate hymenoptera recorded at the site, the species list achieved during the survey
compares favourably with other Staffordshire heathlands, and Penn Common could perhaps be listed as
fourth best Staffordshire heathland site (behind Highgate Common, Gentleshaw Common and Bevins
Birches) (Jukes, 2004) on its aculeate list.
7.8 It is noted that the better West Midlands heathland sites have decent amounts of bare or sparsely
vegetated ground, heather dominated areas and flower-rich heathy verge (Falk, 2010). Penn Common has
some flower-rich areas but not enough heather dominated areas and too little bare or sparsely vegetated
ground to host the level of aculeate hymenoptera that might improve its standing within the West Midlands
league table.
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 14
7.9 As already mentioned the amount of dead wood fauna was unexpected and this seems to be boosting the
site aculeate list. This is perhaps masking the lack of heathland specialists found during this survey. Although
above sites like Pelsall Common in the league table, Penn Common does however have fewer species of
national or regional interest than many other of the key West Midlands heathland sites, with only one very
rare species recorded (Anoplius caviventris) and two regionally scarce species (Tiphia minuta and Psenulus
concolor*) (Note – Psenulus concolor is viewed as common in Shropshire (Cheeseborough pers.comm.) but
is absent from the Staffordshire checklist (Webb, Bloxham & Slawson, 2011), and is also absent from Steven
Falk’s West Midlands region review of bee and wasp assemblages (Falk, 2010)). Both Anoplius caviventris
and Psenulus concolor appear to be new to Staffordshire. It should be noted that Anoplius caviventris was
listed as a general site record during our survey and as such was not included in the ISIS analysis for any
individual compartment, though was part of the whole site analysis.
7.10 Most coleoptera noted were common species with only three heathland specialists recorded (Bradycellus
ruficollis, Lochmaea suturalis and Micrelus ericae). This again illustrates that good quality heathland habitat
is currently at a premium on Penn Common. Approximately 27 species of beetle recorded were wetland
specialists, though all were considered common.
7.11 Wetland diptera were well represented but again most species were considered common. As stated earlier
the subsequent drying out of the wet areas, as the 2011 recording season progressed, did have a deleterious
effect upon our survey and towards the end of the season there were few wetland flies around, particularly
craneflies.
7.12 It would be hoped that further habitat management work could improve the wetland areas across the site.
ISIS analysis of the species assemblages of the calcareous flush showed how species associated with scrub
currently dominate the wetland assemblages. The continued removal of scrub here should begin to show a
change in the assemblages over time, as the vegetation responds to more available light.
7.13 Elsewhere, on the eastern side of the site, the flowing water species assemblages were identified by ISIS as
of importance and these should at least be maintained.
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 15
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Stubbs, A.E., & Drake, C.M., (2001); British Soldierflies and their allies: an illustrated guide to their identification and
ecology. BENHS. Dinton Pastures.
Trevis, G. (2004); Aculeate Hymenoptera in Worcestershire. Worcestershire Record No.16 p27-28
Webb, J., Bloxham, M. & Slawson C. (2005); A provisional atlas of the Bees and Wasps (Hymenoptera) of
Staffordshire. Staffordshire Ecological Record (web version).
Websites
www.data.nbn.org.uk – The NBN Gateway
www.hoverfly.org.uk – The Hoverfly Recording Scheme
www.staffs-ecology.org.uk – Staffordshire Ecological Record
Acknowledgements
The authors would like to thank Paul Mills at Penn Common Golf Course for access to the course and showing us
around on our first visit, and Jon Webb at Natural England for supplying ISIS. We would also like to thank for golfers
who were playing the course during our visits and fully expect that our attendance was blamed for more than one
bad shot or lost ball.
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 16
APPENDIX 1 – Full survey data for Penn Common Golf Course
Order Taxon Compartment Grid ref Date Determiner
Araneae Alopecosa pulverulenta 3 SO89529444 2011 Richard Wright
Araneae Alopecosa pulverulenta 4 SO89459440 2011 Richard Wright
Araneae Antistea elegans 5 SO89599442 2011 Richard Wright
Araneae Antistea elegans 8 SO90379448 2011 Richard Wright
Araneae Bathyphantes gracilis 6 SO89799476 2011 Richard Wright
Araneae Centromerita bicolor 2 SO89629479 2011 Richard Wright
Araneae Centromerita bicolor 6 SO89799476 2011 Richard Wright
Araneae Centromerita bicolor 8 SO90379448 2011 Richard Wright
Araneae Drassyllus pusillus 3 SO89529444 2011 Richard Wright
Araneae Drassyllus pusillus 7 SO90309478 2011 Richard Wright
Araneae Erigone atra 1 SO89709483 2011 Richard Wright
Araneae Erigone dentipalpis 3 SO89529444 2011 Richard Wright
Araneae Erigone dentipalpis 4 SO89459440 2011 Richard Wright
Araneae Gongylidiellum vivum 5 SO89599442 2011 Richard Wright
Araneae Gongylidiellum vivum 8 SO90379448 2011 Richard Wright
Araneae Lepthyphantes tenuis 2 SO89629479 2011 Richard Wright
Araneae Lepthyphantes tenuis 3 SO89529444 2011 Richard Wright
Araneae Lepthyphantes tenuis 4 SO89459440 2011 Richard Wright
Araneae Lepthyphantes tenuis 8 SO90379448 2011 Richard Wright
Araneae Leptorhoptrum robustum 4 SO89459440 2011 Richard Wright
Araneae Leptorhoptrum robustum 6 SO89799476 2011 Richard Wright
Araneae Meta mengei 1 SO89709483 2011 Richard Wright
Araneae Meta segmentata 2 SO89629479 2011 Richard Wright
Araneae Micaria pulicaria 3 SO89529444 2011 Richard Wright
Araneae Micaria pulicaria 4 SO89459440 2011 Richard Wright
Araneae Micaria pulicaria 8 SO90379448 2011 Richard Wright
Araneae Micrargus herbigradus 7 SO90309478 2011 Richard Wright
Araneae Neottiura bimaculata 4 SO89459440 2011 Richard Wright
Araneae Oedothorax gibbosus 1 SO89709483 2011 Richard Wright
Araneae Oedothorax gibbosus 2 SO89629479 2011 Richard Wright
Araneae Oedothorax gibbosus 6 SO89799476 2011 Richard Wright
Araneae Oedothorax gibbosus 8 SO90379448 2011 Richard Wright
Araneae Oedothorax gibbosus 9 SO90259456 2011 Richard Wright
Araneae Oedothorax retusus 1 SO89709483 2011 Richard Wright
Araneae Oedothorax retusus 5 SO89599442 2011 Richard Wright
Araneae Pachygnatha clercki 1 SO89709483 2011 Richard Wright
Araneae Pachygnatha clercki 2 SO89629479 2011 Richard Wright
Araneae Pachygnatha clercki 7 SO90309478 2011 Richard Wright
Araneae Pachygnatha clercki 9 SO90259456 2011 Richard Wright
Araneae Pachygnatha degeeri 1 SO89709483 2011 Richard Wright
Araneae Pachygnatha degeeri 2 SO89629479 2011 Richard Wright
Araneae Pachygnatha degeeri 3 SO89529444 2011 Richard Wright
Araneae Pachygnatha degeeri 4 SO89459440 2011 Richard Wright
Araneae Pardosa nigriceps 3 SO89529444 2011 Richard Wright
Araneae Pardosa nigriceps 6 SO89799476 2011 Richard Wright
Araneae Pardosa palustris 4 SO89459440 2011 Richard Wright
Araneae Pardosa prativaga 1 SO89709483 2011 Richard Wright
Araneae Pardosa prativaga 3 SO89529444 2011 Richard Wright
Araneae Pardosa prativaga 4 SO89459440 2011 Richard Wright
Araneae Pardosa prativaga 6 SO89799476 2011 Richard Wright
Araneae Pardosa prativaga 7 SO90309478 2011 Richard Wright
Araneae Pardosa prativaga 9 SO90259456 2011 Richard Wright
Araneae Pardosa pullata 1 SO89709483 2011 Richard Wright
Araneae Pardosa pullata 2 SO89629479 2011 Richard Wright
Araneae Pardosa pullata 4 SO89459440 2011 Richard Wright
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 17
Araneae Pardosa pullata 6 SO89799476 2011 Richard Wright
Araneae Pardosa pullata 8 SO90379448 2011 Richard Wright
Araneae Pardosa pullata 9 SO90259456 2011 Richard Wright
Araneae Pirata hygrophilus 9 SO90259456 2011 Richard Wright
Araneae Pirata piraticus 1 SO89709483 2011 Richard Wright
Araneae Pirata piraticus 5 SO89599442 2011 Richard Wright
Araneae Pirata piraticus 8 SO90379448 2011 Richard Wright
Araneae Pocadicnemis pumila 5 SO89599442 2011 Richard Wright
Araneae Pocadicnemis pumila 6 SO89799476 2011 Richard Wright
Araneae Robertus lividus 4 SO89459440 2011 Richard Wright
Araneae Robertus lividus 6 SO89799476 2011 Richard Wright
Araneae Walckenaeria vigilax 5 SO89599442 2011 Richard Wright
Araneae Walckenaeria vigilax 9 SO90259456 2011 Richard Wright
Araneae Xysticus cristatus 3 SO89529444 2011 Richard Wright
Araneae Xysticus cristatus 4 SO89459440 2011 Richard Wright
Araneae Xysticus cristatus 6 SO89799476 2011 Richard Wright
Coleoptera Acupalpus dubius 1 SO89709483 2011 Richard Wright
Coleoptera Acupalpus dubius 2 SO89629479 2011 Richard Wright
Coleoptera Acupalpus dubius 5 SO89599442 2011 Richard Wright
Coleoptera Acupalpus dubius 6 SO89799476 2011 Richard Wright
Coleoptera Acupalpus dubius 7 SO90309478 2011 Richard Wright
Coleoptera Acupalpus dubius 8 SO90379448 2011 Richard Wright
Coleoptera Acupalpus dubius 9 SO90259456 2011 Richard Wright
Coleoptera Adalia bipunctata site record SO898945 20/05/2011 Nigel Jones
Coleoptera Adalia bipunctata 9 SO90259456 29/06/2011 Pete Boardman
Coleoptera Agonum emarginatum 5 SO89599442 2011 Richard Wright
Coleoptera Agonum emarginatum 8 SO90379448 2011 Richard Wright
Coleoptera Agonum emarginatum 9 SO90259456 2011 Richard Wright
Coleoptera Agonum fuliginosum 5 SO89599442 2011 Richard Wright
Coleoptera Agonum fuliginosum 7 SO90309478 2011 Richard Wright
Coleoptera Agonum muelleri 8 SO90379448 2011 Richard Wright
Coleoptera Agonum viduum 5 SO89599442 2011 Richard Wright
Coleoptera Agonum viduum 8 SO90379448 2011 Richard Wright
Coleoptera Agriotes acuminatus 3 SO89529444 2011 Richard Wright
Coleoptera Agriotes lineatus 2 SO89629479 2011 Richard Wright
Coleoptera Agriotes pallidulus 3 SO89529444 2011 Richard Wright
Coleoptera Agriotes sputator 4 SO89459440 2011 Richard Wright
Coleoptera Amara communis 3 SO89529444 2011 Richard Wright
Coleoptera Amara communis 4 SO89459440 2011 Richard Wright
Coleoptera Amara communis 6 SO89799476 2011 Richard Wright
Coleoptera Amara lunicollis 3 SO89529444 2011 Richard Wright
Coleoptera Amara lunicollis 4 SO89459440 2011 Richard Wright
Coleoptera Amara lunicollis 6 SO89799476 2011 Richard Wright
Coleoptera Amara plebeja 1 SO89709483 2011 Richard Wright
Coleoptera Amara plebeja 2 SO89629479 2011 Richard Wright
Coleoptera Amara plebeja 5 SO89599442 2011 Richard Wright
Coleoptera Amara plebeja 8 SO90379448 2011 Richard Wright
Coleoptera Amara plebeja 9 SO90259456 2011 Richard Wright
Coleoptera Anacaena globulus 1 SO89709483 2011 Richard Wright
Coleoptera Anacaena globulus 5 SO89599442 2011 Richard Wright
Coleoptera Anacaena globulus 6 SO89799476 2011 Richard Wright
Coleoptera Anacaena globulus 8 SO90379448 2011 Richard Wright
Coleoptera Anacaena globulus 9 SO90259456 2011 Richard Wright
Coleoptera Anotylus rugosus 1 SO89709483 2011 Richard Wright
Coleoptera Anotylus rugosus 2 SO89629479 2011 Richard Wright
Coleoptera Anotylus rugosus 3 SO89529444 2011 Richard Wright
Coleoptera Anotylus rugosus 4 SO89459440 2011 Richard Wright
Coleoptera Anotylus rugosus 6 SO89799476 2011 Richard Wright
Coleoptera Anotylus rugosus 7 SO90309478 2011 Richard Wright
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 18
Coleoptera Anotylus rugosus 9 SO90259456 2011 Richard Wright
Coleoptera Anthobium unicolor 2 SO89629479 2011 Richard Wright
Coleoptera Anthobium unicolor 7 SO90309478 2011 Richard Wright
Coleoptera Aphodius prodromus 8 SO90379448 2011 Richard Wright
Coleoptera Aphthona euphorbiae 3 SO89529444 2011 Richard Wright
Coleoptera Apion frumentarium 5 SO89599442 20/04/2011 Pete Boardman
Coleoptera Apion frumentarium 7 SO90309478 20/04/2011 Pete Boardman
Coleoptera Athous haemorrhoidalis 7 SO90309478 2011 Richard Wright
Coleoptera Barypeithes pellucidus 2 SO89629479 2011 Richard Wright
Coleoptera Barypeithes pellucidus 3 SO89529444 2011 Richard Wright
Coleoptera Barypeithes pellucidus 4 SO89459440 2011 Richard Wright
Coleoptera Barypeithes pellucidus 7 SO90309478 2011 Richard Wright
Coleoptera Bembidion decorum 5 SO89599442 2011 Richard Wright
Coleoptera Bembidion guttula 5 SO89599442 2011 Richard Wright
Coleoptera Bembidion guttula 6 SO89799476 2011 Richard Wright
Coleoptera Bembidion lampros 3 SO89529444 2011 Richard Wright
Coleoptera Bembidion lampros 4 SO89459440 2011 Richard Wright
Coleoptera Bembidion lampros 9 SO90259456 2011 Richard Wright
Coleoptera Bembidion mannerheimi 1 SO89709483 2011 Richard Wright
Coleoptera Bembidion mannerheimi 2 SO89629479 2011 Richard Wright
Coleoptera Bembidion mannerheimi 6 SO89799476 2011 Richard Wright
Coleoptera Bembidion mannerheimi 7 SO90309478 2011 Richard Wright
Coleoptera Bembidion mannerheimi 8 SO90379448 2011 Richard Wright
Coleoptera Bembidion mannerheimi 9 SO90259456 2011 Richard Wright
Coleoptera Bembidion obtusum 1 SO89709483 2011 Richard Wright
Coleoptera Bembidion obtusum 3 SO89529444 2011 Richard Wright
Coleoptera Bembidion obtusum 4 SO89459440 2011 Richard Wright
Coleoptera Bembidion obtusum 6 SO89799476 2011 Richard Wright
Coleoptera Bembidion obtusum 9 SO90259456 2011 Richard Wright
Coleoptera Brachygluta haematica 4 SO89459440 2011 Richard Wright
Coleoptera Brachygluta haematica 8 SO90379448 2011 Richard Wright
Coleoptera Bradycellus ruficollis 4 SO89459440 2011 Richard Wright
Coleoptera Bradycellus ruficollis 6 SO89799476 2011 Richard Wright
Coleoptera Bradycellus verbasci 4 SO89459440 2011 Richard Wright
Coleoptera Calvia quattuordecimguttata 7 SO90029443 20/04/2011 Pete Boardman
Coleoptera Calvia quattuordecimguttata site record SO898945 20/05/2011 Nigel Jones
Coleoptera Cantharis nigra 3 SO89529444 2011 Richard Wright
Coleoptera Cassida rubiginosa 3 SO89529444 2011 Richard Wright
Coleoptera Ceratapion gibbirostre 3 SO89529444 2011 Richard Wright
Coleoptera Chaetocnema concinna 3 SO89529444 2011 Richard Wright
Coleoptera Chaetocnema concinna 4 SO89459440 2011 Richard Wright
Coleoptera Chaetocnema hortensis 3 SO89529444 2011 Richard Wright
Coleoptera Chilocorus renipustulatus site record SO898945 20/05/2011 Nigel Jones
Coleoptera Clivina fossor 8 SO90379448 2011 Richard Wright
Coleoptera Clytus arietis 12 SO90349456 18/04/2011 Ian Cheeseborough
Coleoptera Clytus arietis 8 SO90379448 20/05/2011 Nigel Jones
Coleoptera Coccidula rufa 1 SO89709483 2011 Richard Wright
Coleoptera Coccidula rufa 5 SO89599442 2011 Richard Wright
Coleoptera Coccinella septempunctata 4 SO89459440 2011 Richard Wright
Coleoptera Coccinella septempunctata site record SO899945 18/04/2011 Ian Cheeseborough
Coleoptera Coccinella septempunctata 7 SO90029443 20/04/2011 Pete Boardman
Coleoptera Coccinella septempunctata 7 SO90309478 20/04/2011 Pete Boardman
Coleoptera Coccinella septempunctata site record SO898945 20/05/2011 Nigel Jones
Coleoptera Corticarina minuta 3 SO89529444 2011 Richard Wright
Coleoptera Cortinicara gibbosa 4 SO89459440 2011 Richard Wright
Coleoptera Cortinicara gibbosa 6 SO89799476 2011 Richard Wright
Coleoptera Cortinicara gibbosa 7 SO90309478 2011 Richard Wright
Coleoptera Dryops ernesti 1 SO89709483 2011 Richard Wright
Coleoptera Dryops ernesti 5 SO89599442 2011 Richard Wright
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 19
Coleoptera Dryops ernesti 8 SO90379448 2011 Richard Wright
Coleoptera Dryops ernesti 9 SO90259456 2011 Richard Wright
Coleoptera Eutrichapion ervi 3 SO89529444 2011 Richard Wright
Coleoptera Eutrichapion viciae 3 SO89529444 2011 Richard Wright
Coleoptera Exochomus quadripustulatus 7 SO90309478 2011 Richard Wright
Coleoptera Gabrius breviventer 5 SO89599442 2011 Richard Wright
Coleoptera Gabrius breviventer 8 SO90379448 2011 Richard Wright
Coleoptera Gymnetron pascuorum 3 SO89529444 2011 Richard Wright
Coleoptera Gymnetron pascuorum 4 SO89459440 2011 Richard Wright
Coleoptera Harmonia axyridis site record SO898945 20/05/2011 Nigel Jones
Coleoptera Lathrobium geminum 5 SO89599442 2011 Richard Wright
Coleoptera Lathrobium geminum 8 SO90379448 2011 Richard Wright
Coleoptera Leiosoma deflexum 3 SO89529444 2011 Richard Wright
Coleoptera Lesteva sicula 5 SO89599442 2011 Richard Wright
Coleoptera Lesteva sicula 8 SO90379448 2011 Richard Wright
Coleoptera Lochmaea suturalis 4 SO89459440 2011 Richard Wright
Coleoptera Lochmaea suturalis 6 SO89799476 2011 Richard Wright
Coleoptera Malthinus flaveolus 8 SO90379448 29/06/2011 Pete Boardman
Coleoptera Mecinus pyraster 3 SO89529444 2011 Richard Wright
Coleoptera Micrelus ericae 4 SO89459440 2011 Richard Wright
Coleoptera Micrelus ericae 6 SO89799476 2011 Richard Wright
Coleoptera Nargus velox 4 SO89459440 2011 Richard Wright
Coleoptera Nargus velox 8 SO90379448 2011 Richard Wright
Coleoptera Nargus velox 9 SO90259456 2011 Richard Wright
Coleoptera Nebria brevicollis 3 SO89529444 2011 Richard Wright
Coleoptera Nebria brevicollis 4 SO89459440 2011 Richard Wright
Coleoptera Nebria brevicollis 9 SO90259456 2011 Richard Wright
Coleoptera Neocrepidodera ferruginea 9 SO90259456 2011 Richard Wright
Coleoptera Neocrepidodera transversa 3 SO89529444 2011 Richard Wright
Coleoptera Notaris acridulus 5 SO89599442 2011 Richard Wright
Coleoptera Notaris acridulus 8 SO90379448 2011 Richard Wright
Coleoptera Ochthebius minimus 9 SO90259456 2011 Richard Wright
Coleoptera Oedemera nobilis 9a SO90269452 20/05/2011 Ian Cheeseborough
Coleoptera Olibrus aeneus 5 SO89599442 2011 Richard Wright
Coleoptera Oxystoma craccae 3 SO89529444 2011 Richard Wright
Coleoptera Paradromius linearis 3 SO89529444 2011 Richard Wright
Coleoptera Perapion curtirostre 1 SO89709483 2011 Richard Wright
Coleoptera Perapion curtirostre 3 SO89529444 2011 Richard Wright
Coleoptera Perapion curtirostre 9 SO90259456 2011 Richard Wright
Coleoptera Perapion hydrolapathi 1 SO89709483 2011 Richard Wright
Coleoptera Philonthus cognatus 1 SO89709483 2011 Richard Wright
Coleoptera Philonthus cognatus 2 SO89629479 2011 Richard Wright
Coleoptera Philonthus cognatus 3 SO89529444 2011 Richard Wright
Coleoptera Philonthus cognatus 4 SO89459440 2011 Richard Wright
Coleoptera Philonthus cognatus 8 SO90379448 2011 Richard Wright
Coleoptera Phyllobius pomaceus 9 SO90259456 20/04/2011 Pete Boardman
Coleoptera Phyllobius roboretanus 7 SO90309478 2011 Richard Wright
Coleoptera Phyllotreta tetrastigma 5 SO89599442 2011 Richard Wright
Coleoptera Phyllotreta tetrastigma 6 SO89799476 2011 Richard Wright
Coleoptera Phyllotreta tetrastigma 8 SO90379448 2011 Richard Wright
Coleoptera Poecilus versicolor 3 SO89529444 2011 Richard Wright
Coleoptera Propylea quattuordecimpunctata 7 SO90309478 2011 Richard Wright
Coleoptera Propylea quattuordecimpunctata site record SO899945 18/04/2011 Ian Cheeseborough
Coleoptera Propylea quattuordecimpunctata 8a SO903945 28/07/2011 Ian Cheeseborough
Coleoptera Protapion assimile 3 SO89529444 2011 Richard Wright
Coleoptera Protapion fulvipes 3 SO89529444 2011 Richard Wright
Coleoptera Protapion trifolii 3 SO89529444 2011 Richard Wright
Coleoptera Psyllobora vigintiduopunctata 3 SO89529444 2011 Richard Wright
Coleoptera Pterostichus diligens 5 SO89599442 2011 Richard Wright
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 20
Coleoptera Pterostichus diligens 9 SO90259456 2011 Richard Wright
Coleoptera Pterostichus madidus 3 SO89529444 2011 Richard Wright
Coleoptera Pterostichus melanarius 1 SO89709483 2011 Richard Wright
Coleoptera Pterostichus melanarius 2 SO89629479 2011 Richard Wright
Coleoptera Pterostichus melanarius 8 SO90379448 2011 Richard Wright
Coleoptera Pterostichus melanarius 9 SO90259456 2011 Richard Wright
Coleoptera Pterostichus nigrita 5 SO89599442 2011 Richard Wright
Coleoptera Pterostichus nigrita 7 SO90309478 2011 Richard Wright
Coleoptera Pterostichus nigrita 8 SO90379448 2011 Richard Wright
Coleoptera Pterostichus nigrita 9 SO90259456 2011 Richard Wright
Coleoptera Pterostichus strenuus 1 SO89709483 2011 Richard Wright
Coleoptera Pterostichus strenuus 2 SO89629479 2011 Richard Wright
Coleoptera Pterostichus strenuus 4 SO89459440 2011 Richard Wright
Coleoptera Pterostichus strenuus 5 SO89599442 2011 Richard Wright
Coleoptera Pterostichus strenuus 6 SO89799476 2011 Richard Wright
Coleoptera Pterostichus strenuus 7 SO90309478 2011 Richard Wright
Coleoptera Pterostichus strenuus 8 SO90379448 2011 Richard Wright
Coleoptera Pterostichus strenuus 9 SO90259456 2011 Richard Wright
Coleoptera Pterostichus vernalis 5 SO89599442 2011 Richard Wright
Coleoptera Pterostichus vernalis 8 SO90379448 2011 Richard Wright
Coleoptera Pterostichus vernalis 9 SO90259456 2011 Richard Wright
Coleoptera Quedius curtipennis 1 SO89709483 2011 Richard Wright
Coleoptera Quedius curtipennis 2 SO89629479 2011 Richard Wright
Coleoptera Quedius curtipennis 3 SO89529444 2011 Richard Wright
Coleoptera Quedius curtipennis 6 SO89799476 2011 Richard Wright
Coleoptera Quedius curtipennis 7 SO90309478 2011 Richard Wright
Coleoptera Quedius fuliginosus 4 SO89459440 2011 Richard Wright
Coleoptera Quedius fuliginosus 9 SO90259456 2011 Richard Wright
Coleoptera Quedius maurorufus 5 SO89599442 2011 Richard Wright
Coleoptera Quedius maurorufus 6 SO89799476 2011 Richard Wright
Coleoptera Quedius maurorufus 9 SO90259456 2011 Richard Wright
Coleoptera Quedius molochinus 7 SO90309478 2011 Richard Wright
Coleoptera Rhagonycha fulva 3 SO89529444 2011 Richard Wright
Coleoptera Rhagonycha limbata 7 SO90309478 2011 Richard Wright
Coleoptera Rhyzobius litura 1 SO89709483 2011 Richard Wright
Coleoptera Rhyzobius litura 2 SO89629479 2011 Richard Wright
Coleoptera Rhyzobius litura 3 SO89529444 2011 Richard Wright
Coleoptera Rhyzobius litura 5 SO89599442 2011 Richard Wright
Coleoptera Rugilus rufipes 2 SO89629479 2011 Richard Wright
Coleoptera Rutpela maculata 9a SO90269452 20/06/2011 Ian Cheeseborough
Coleoptera Sitona cambricus 2 SO89629479 2011 Richard Wright
Coleoptera Sitona lineatus 3 SO89529444 2011 Richard Wright
Coleoptera Sitona lineatus 9 SO90259456 2011 Richard Wright
Coleoptera Sphaeroderma testaceum 3 SO89529444 2011 Richard Wright
Coleoptera Sphaeroderma testaceum 8 SO90379448 2011 Richard Wright
Coleoptera Stenus clavicornis 1 SO89709483 2011 Richard Wright
Coleoptera Stenus clavicornis 2 SO89629479 2011 Richard Wright
Coleoptera Stenus clavicornis 5 SO89599442 2011 Richard Wright
Coleoptera Stenus clavicornis 6 SO89799476 2011 Richard Wright
Coleoptera Stenus clavicornis 7 SO90309478 2011 Richard Wright
Coleoptera Stenus clavicornis 8 SO90379448 2011 Richard Wright
Coleoptera Stenus clavicornis 9 SO90259456 2011 Richard Wright
Coleoptera Stenus flavipes 2 SO89629479 2011 Richard Wright
Coleoptera Stenus flavipes 5 SO89599442 2011 Richard Wright
Coleoptera Stenus providus 5 SO89599442 2011 Richard Wright
Coleoptera Sunius propinquus 3 SO89529444 2011 Richard Wright
Coleoptera Sunius propinquus 6 SO89799476 2011 Richard Wright
Coleoptera Tachinus rufipes 2 SO89629479 2011 Richard Wright
Coleoptera Tachinus rufipes 3 SO89529444 2011 Richard Wright
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 21
Coleoptera Tachinus rufipes 4 SO89459440 2011 Richard Wright
Coleoptera Tachinus rufipes 7 SO90309478 2011 Richard Wright
Coleoptera Tachyporus chrysomelinus 1 SO89709483 2011 Richard Wright
Coleoptera Tachyporus chrysomelinus 3 SO89529444 2011 Richard Wright
Coleoptera Tachyporus chrysomelinus 6 SO89799476 2011 Richard Wright
Coleoptera Tachyporus dispar 5 SO89599442 2011 Richard Wright
Coleoptera Tachyporus dispar 8 SO90379448 2011 Richard Wright
Coleoptera Tachyporus dispar 9 SO90259456 2011 Richard Wright
Coleoptera Tachyporus hypnorum 1 SO89709483 2011 Richard Wright
Coleoptera Tachyporus hypnorum 2 SO89629479 2011 Richard Wright
Coleoptera Tachyporus hypnorum 3 SO89529444 2011 Richard Wright
Coleoptera Tachyporus hypnorum 4 SO89459440 2011 Richard Wright
Coleoptera Tachyporus hypnorum 5 SO89599442 2011 Richard Wright
Coleoptera Tachyporus hypnorum 6 SO89799476 2011 Richard Wright
Coleoptera Tachyporus hypnorum 7 SO90309478 2011 Richard Wright
Coleoptera Tachyporus hypnorum 9 SO90259456 2011 Richard Wright
Coleoptera Tachyporus solutus 2 SO89629479 2011 Richard Wright
Coleoptera Tachyporus solutus 7 SO90309478 2011 Richard Wright
Coleoptera Tachyporus solutus 9 SO90259456 2011 Richard Wright
Coleoptera Trachyphloeus asperatus 4 SO89459440 2011 Richard Wright
Coleoptera Trachyphloeus asperatus 6 SO89799476 2011 Richard Wright
Coleoptera Trixagus dermestoides 3 SO89529444 2011 Richard Wright
Coleoptera Trixagus dermestoides 4 SO89459440 2011 Richard Wright
Coleoptera Trixagus dermestoides 6 SO89799476 2011 Richard Wright
Coleoptera Xantholinus linearis 3 SO89529444 2011 Richard Wright
Coleoptera Xantholinus longiventris 2 SO89629479 2011 Richard Wright
Coleoptera Xantholinus longiventris 8 SO90379448 2011 Richard Wright
Diplopoda Cylindroiulus punctatus 6 SO89799476 2011 Richard Wright
Diplopoda Cylindroiulus punctatus 7 SO90309478 2011 Richard Wright
Diplopoda Ophyiulus pilosus 5 SO89599442 2011 Richard Wright
Diplopoda Ophyiulus pilosus 6 SO89799476 2011 Richard Wright
Diplopoda Ophyiulus pilosus 7 SO90309478 2011 Richard Wright
Diplopoda Ophyiulus pilosus 8 SO90379448 2011 Richard Wright
Diplopoda Polydesmus angustus 1 SO89709483 2011 Richard Wright
Diplopoda Polydesmus angustus 2 SO89629479 2011 Richard Wright
Diplopoda Polydesmus angustus 4 SO89459440 2011 Richard Wright
Diplopoda Polydesmus angustus 5 SO89599442 2011 Richard Wright
Diplopoda Polydesmus angustus 8 SO90379448 2011 Richard Wright
Diplopoda Tachypodoiulus niger 1 SO89709483 2011 Richard Wright
Diplopoda Tachypodoiulus niger 2 SO89629479 2011 Richard Wright
Diplopoda Tachypodoiulus niger 4 SO89459440 2011 Richard Wright
Diplopoda Tachypodoiulus niger 5 SO89599442 2011 Richard Wright
Diplopoda Tachypodoiulus niger 6 SO89799476 2011 Richard Wright
Diplopoda Tachypodoiulus niger 7 SO90309478 2011 Richard Wright
Diplopoda Tachypodoiulus niger 8 SO90379448 2011 Richard Wright
Diplopoda Tachypodoiulus niger 9 SO90259456 2011 Richard Wright
Diptera Agathomyia antennata 8 SO90379448 20/05/2011 Nigel Jones
Diptera Anomoia purmunda site record SO898945 28/07/2011 Nigel Jones
Diptera Anthomyia procellaris 9 SO90259456 20/05/2011 Nigel Jones
Diptera Argyra argentina 8 SO90379448 28/07/2011 Nigel Jones
Diptera Beris chalybata 5 SO89599442 20/05/2011 Nigel Jones
Diptera Beris chalybata 7 SO90309478 20/05/2011 Nigel Jones
Diptera Beris morrisii 8 SO90379448 28/07/2011 Nigel Jones
Diptera Beris vallata 8 SO90379448 28/07/2011 Nigel Jones
Diptera Bibio johannis 1 SO89709483 18/04/2011 Nigel Jones
Diptera Bibio johannis 5 SO89599442 18/04/2011 Nigel Jones
Diptera Bibio marci site record SO898945 18/04/2011 Nigel Jones
Diptera Bibio marci 7 SO90029443 20/04/2011 Pete Boardman
Diptera Neolimnomyia nemoralis 5 SO89599442 20/05/2011 Pete Boardman
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 22
Diptera Neolimnomyia nemoralis 8 SO90379448 20/05/2011 Pete Boardman
Diptera Neolimnomyia nemoralis 9 SO90259456 29/06/2011 Pete Boardman
Diptera Campsicnemus curvipes 8 SO90379448 18/04/2011 Nigel Jones
Diptera Campsicnemus curvipes 8 SO90379448 20/05/2011 Nigel Jones
Diptera Campsicnemus curvipes 8 SO90379448 28/07/2011 Nigel Jones
Diptera Chaetostomella cylindrica 1 SO89709483 20/05/2011 Nigel Jones
Diptera Chaetostomella cylindrica site record SO898945 28/07/2011 Nigel Jones
Diptera Chamaepsila rosae 5 SO89599442 20/05/2011 Nigel Jones
Diptera Chamaepsila rosae 8 SO90379448 20/05/2011 Nigel Jones
Diptera Cheilosia albitarsis 9 SO90259456 20/05/2011 Nigel Jones
Diptera Cheilosia illustrata 9a SO90269452 20/06/2011 Ian Cheeseborough
Diptera Cheilosia illustrata site record SO898945 28/07/2011 Nigel Jones
Diptera Cheilotrichia cinerascens 5 SO89599442 20/04/2011 Pete Boardman
Diptera Cheilotrichia cinerascens 7 SO90029443 20/04/2011 Pete Boardman
Diptera Cheilotrichia cinerascens 9 SO90259456 20/04/2011 Pete Boardman
Diptera Cheilotrichia cinerascens 7 SO90309478 20/05/2011 Pete Boardman
Diptera Chelifera precatoria 5 SO89599442 20/05/2011 Nigel Jones
Diptera Chrysogaster solstitialis site record SO898945 28/07/2011 Nigel Jones
Diptera Chrysopilus cristatus 5 SO89599442 20/05/2011 Nigel Jones
Diptera Chrysopilus cristatus 8 SO90379448 20/05/2011 Nigel Jones
Diptera Chrysopilus cristatus 9 SO90259456 29/06/2011 Pete Boardman
Diptera Chrysopilus cristatus 9 SO90259456 29/06/2011 Pete Boardman
Diptera Chrysotoxum festivum site record SO898945 28/07/2011 Nigel Jones
Diptera Chrysotus gramineus site record SO898945 28/07/2011 Nigel Jones
Diptera Cleigastra apicalis 9 SO90259456 18/04/2011 Nigel Jones
Diptera Clusiodes albimanus 8 SO90379448 28/07/2011 Nigel Jones
Diptera Clusiodes gentilis 8 SO90379448 20/05/2011 Nigel Jones
Diptera Coenia palustris 8 SO90379448 28/07/2011 Nigel Jones
Diptera Coenosia femoratus site record SO898945 28/07/2011 Nigel Jones
Diptera Coenosia infantula site record SO898945 28/07/2011 Nigel Jones
Diptera Coenosia tigrina site record SO898945 20/05/2011 Nigel Jones
Diptera Coenosia tigrina 9 SO90259456 28/07/2011 Nigel Jones
Diptera Coenosia tigrina site record SO898945 28/07/2011 Nigel Jones
Diptera Coenosia tigrina site record SO898945 28/07/2011 Nigel Jones
Diptera Conops quadrifasciatus site record SO898945 28/07/2011 Nigel Jones
Diptera Dasysyrphus venustus 8 SO90379448 20/05/2011 Nigel Jones
Diptera Dicranomyia affinis 5 SO89599442 20/05/2011 Pete Boardman
Diptera Dicranomyia chorea 5 SO89599442 20/04/2011 Pete Boardman
Diptera Dicranomyia chorea 7 SO90029443 20/04/2011 Pete Boardman
Diptera Dicranomyia chorea 7 SO90309478 20/05/2011 Pete Boardman
Diptera Dicranota pavida 5 SO89599442 20/05/2011 Pete Boardman
Diptera Dicranota subtilis 5 SO89599442 20/05/2011 Pete Boardman
Diptera Dioctria atricapilla 8 SO90379448 20/05/2011 Nigel Jones
Diptera Dioctria baumhaueri 8 SO90379448 29/06/2011 Pete Boardman
Diptera Ditrichophora calceata 8 SO90379448 28/07/2011 Nigel Jones
Diptera Dolichocephala irrorata 8 SO90379448 18/04/2011 Nigel Jones
Diptera Dolichopus brevipennis 2 SO89629479 20/05/2011 Nigel Jones
Diptera Dolichopus brevipennis 5 SO89599442 20/05/2011 Nigel Jones
Diptera Dolichopus brevipennis 9 SO90259456 20/05/2011 Nigel Jones
Diptera Dolichopus discifer 8 SO90379448 28/07/2011 Nigel Jones
Diptera Dolichopus picipes 5 SO89599442 20/05/2011 Nigel Jones
Diptera Dolichopus plumipes 5 SO89599442 20/05/2011 Nigel Jones
Diptera Dolichopus plumipes 8 SO90379448 20/05/2011 Nigel Jones
Diptera Dolichopus plumipes 9 SO90259456 20/05/2011 Nigel Jones
Diptera Dolichopus plumipes 8 SO90379448 28/07/2011 Nigel Jones
Diptera Dolichopus plumipes 9 SO90259456 28/07/2011 Nigel Jones
Diptera Dolichopus popularis 5 SO89599442 20/05/2011 Nigel Jones
Diptera Dolichopus popularis 8 SO90379448 28/07/2011 Nigel Jones
Diptera Dolichopus trivialis 5 SO89599442 20/05/2011 Nigel Jones
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 23
Diptera Dolichopus trivialis 8 SO90379448 28/07/2011 Nigel Jones
Diptera Dolichopus trivialis 9 SO90259456 28/07/2011 Nigel Jones
Diptera Dolichopus ungulatus 5 SO89599442 20/05/2011 Nigel Jones
Diptera Dolichopus ungulatus 8 SO90379448 28/07/2011 Nigel Jones
Diptera Elgiva cucularia 1 SO89709483 20/05/2011 Nigel Jones
Diptera Elgiva solicita 1 SO89709483 20/05/2011 Nigel Jones
Diptera Eloeophila maculata 5 SO89599442 20/05/2011 Pete Boardman
Diptera Eloeophila submarmorata 5 SO89599442 20/05/2011 Nigel Jones
Diptera Eloeophila submarmorata 5 SO89599442 20/05/2011 Pete Boardman
Diptera Empis aestiva 1 SO89709483 18/04/2011 Nigel Jones
Diptera Empis livida 7 SO90309478 28/07/2011 Nigel Jones
Diptera Empis livida site record SO898945 28/07/2011 Nigel Jones
Diptera Empis nigripes 9 SO90259456 18/04/2011 Nigel Jones
Diptera Empis nigripes 5 SO89599442 20/05/2011 Nigel Jones
Diptera Empis nigripes 8 SO90379448 20/05/2011 Nigel Jones
Diptera Empis praevia 5 SO89599442 20/05/2011 Nigel Jones
Diptera Empis stercorea 5 SO89599442 20/05/2011 Nigel Jones
Diptera Empis tessellata site record SO898945 20/05/2011 Nigel Jones
Diptera Epistrophe eligans 8 SO90379448 20/04/2011 Pete Boardman
Diptera Epistrophe eligans site record SO898945 20/05/2011 Nigel Jones
Diptera Epistrophe grossulariae 8 SO90379448 28/07/2011 Nigel Jones
Diptera Episyrphus balteatus site record SO898945 20/05/2011 Nigel Jones
Diptera Episyrphus balteatus 9 SO90259456 29/06/2011 Pete Boardman
Diptera Episyrphus balteatus 9 SO90259456 29/06/2011 Pete Boardman
Diptera Episyrphus balteatus 1 SO89709483 15/09/2011 Pete Boardman
Diptera Episyrphus balteatus 9 SO90259456 15/09/2011 Pete Boardman
Diptera Erioconopa trivialis 9 SO90259456 18/04/2011 Pete Boardman
Diptera Erioconopa trivialis 9 SO90259456 18/04/2011 Pete Boardman
Diptera Erioconopa trivialis 5 SO89599442 20/04/2011 Pete Boardman
Diptera Erioconopa trivialis 8 SO90379448 20/05/2011 Pete Boardman
Diptera Erioptera fusculenta 9 SO90259456 29/06/2011 Pete Boardman
Diptera Erioptera squalida 9 SO90259456 29/06/2011 Pete Boardman
Diptera Eriothrix rufomaculata site record SO898945 28/07/2011 Nigel Jones
Diptera Eristalis arbustorum site record SO898945 20/05/2011 Nigel Jones
Diptera Eristalis intricarius 9 SO90259456 20/04/2011 Pete Boardman
Diptera Eristalis pertinax 8 SO90379448 18/04/2011 Nigel Jones
Diptera Eristalis pertinax 9 SO90259456 20/04/2011 Pete Boardman
Diptera Eristalis tenax site record SO898945 20/05/2011 Nigel Jones
Diptera Eristalis tenax 8 SO90379448 28/07/2011 Nigel Jones
Diptera Eristalis tenax 1 SO89709483 15/09/2011 Pete Boardman
Diptera Eristalis tenax 9 SO90259456 15/09/2011 Pete Boardman
Diptera Eudasyphora cyanella site record SO898945 28/07/2011 Nigel Jones
Diptera Eumerus funeralis site record SO898945 20/06/2011 Nigel Jones
Diptera Eupeodes latifasciatus 1 SO89709483 20/05/2011 Nigel Jones
Diptera Eupeodes luniger site record SO898945 20/06/2011 Nigel Jones
Diptera Fannia armata 5 SO89599442 20/05/2011 Nigel Jones
Diptera Fannia armata 5 SO89599442 20/05/2011 Nigel Jones
Diptera Fannia lustrator 8 SO90379448 20/05/2011 Nigel Jones
Diptera Fannia serena 5 SO89599442 20/05/2011 Nigel Jones
Diptera Fannia similis 5 SO89599442 20/05/2011 Nigel Jones
Diptera Geomyza tripunctata 1 SO89709483 18/04/2011 Nigel Jones
Diptera Gloma fuscipennis 5 SO89599442 20/05/2011 Nigel Jones
Diptera Gloma fuscipennis 8 SO90379448 20/05/2011 Nigel Jones
Diptera Helina impuncta 5 SO89599442 20/05/2011 Nigel Jones
Diptera Helina impuncta 8 SO90379448 28/07/2011 Nigel Jones
Diptera Helina reversio 5 SO89599442 28/07/2011 Nigel Jones
Diptera Helina reversio 9 SO90259456 28/07/2011 Nigel Jones
Diptera Helophilus hybridus 1 SO89709483 28/07/2011 Nigel Jones
Diptera Helophilus pendulus 8 SO90379448 18/04/2011 Nigel Jones
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 24
Diptera Helophilus pendulus 8 SO90379448 20/05/2011 Nigel Jones
Diptera Helophilus pendulus site record SO898945 20/05/2011 Nigel Jones
Diptera Helophilus pendulus 9 SO90259456 29/06/2011 Pete Boardman
Diptera Hercostomus aerosus 8 SO90379448 28/07/2011 Nigel Jones
Diptera Hercostomus cupreus 1 SO89709483 20/05/2011 Nigel Jones
Diptera Hercostomus cupreus 5 SO89599442 20/05/2011 Nigel Jones
Diptera Hercostomus germanus 8 SO90379448 28/07/2011 Nigel Jones
Diptera Hercostomus parvilamellatus 5 SO89599442 20/05/2011 Nigel Jones
Diptera Herina frondescentiae 9 SO90259456 20/05/2011 Nigel Jones
Diptera Herina frondescentiae site record SO898945 28/07/2011 Nigel Jones
Diptera Hilara curtisi 8 SO90379448 20/05/2011 Nigel Jones
Diptera Hilara litorea 8 SO90379448 28/07/2011 Nigel Jones
Diptera Hilara lurida 8 SO90379448 28/07/2011 Nigel Jones
Diptera Hybos femoratus 8 SO90379448 28/07/2011 Nigel Jones
Diptera Hydrellia maura 5 SO89599442 20/05/2011 Nigel Jones
Diptera Hydrellia maura 9 SO90259456 20/05/2011 Nigel Jones
Diptera Hydrellia maura 9 SO90259456 20/05/2011 Nigel Jones
Diptera Hydrellia maura 8 SO90379448 28/07/2011 Nigel Jones
Diptera Hydrellia maura 9 SO90259456 28/07/2011 Nigel Jones
Diptera Hydromya dorsalis 8 SO90379448 18/04/2011 Nigel Jones
Diptera Hydromya dorsalis 9 SO90259456 18/04/2011 Nigel Jones
Diptera Hydromya dorsalis 5 SO89599442 20/04/2011 Pete Boardman
Diptera Hydromya dorsalis 9 SO90259456 28/07/2011 Nigel Jones
Diptera Hydrotaea cyrtoneurina 5 SO89599442 20/05/2011 Nigel Jones
Diptera Hydrotaea cyrtoneurina 8 SO90379448 28/07/2011 Nigel Jones
Diptera Ilisia maculata 8 SO90379448 20/05/2011 Nigel Jones
Diptera Ilisia maculata 9 SO90259456 20/05/2011 Pete Boardman
Diptera Jassidophaga villosa 9 SO90259456 18/04/2011 Nigel Jones
Diptera Lauxania cylindricornis site record SO898945 20/05/2011 Nigel Jones
Diptera Leptogaster cylindrica site record SO901944 20/06/2011 Ian Cheeseborough
Diptera Leucophora obtusa 8a SO903945 18/04/2011 Nigel Jones
Diptera Limnellia quadrata 9 SO90259456 18/04/2011 Nigel Jones
Diptera Limnia paludicloa 9 SO90259456 28/07/2011 Nigel Jones
Diptera Limnia unguicornis 8 SO90379448 20/05/2011 Nigel Jones
Diptera Limonia flavipes 5 SO89599442 20/05/2011 Pete Boardman
Diptera Limonia macrostigma 8 SO90379448 20/05/2011 Pete Boardman
Diptera Limonia nubeculosa 8 SO90379448 20/05/2011 Pete Boardman
Diptera Limonia nubeculosa 9 SO90259456 15/09/2011 Pete Boardman
Diptera Limonia phragmitidis 8 SO90379448 20/05/2011 Pete Boardman
Diptera Lonchaea chorea 8 SO90379448 20/05/2011 Nigel Jones
Diptera Lonchoptera lutea 5 SO89599442 18/04/2011 Nigel Jones
Diptera Lonchoptera lutea 7 SO90309478 18/04/2011 Nigel Jones
Diptera Loxocera albiseta 8 SO90379448 28/07/2011 Nigel Jones
Diptera Loxocera albiseta 9 SO90259456 28/07/2011 Nigel Jones
Diptera Lydella stabulans site record SO898945 28/07/2011 Nigel Jones
Diptera Machimus cingulatus site record SO898945 28/07/2011 Nigel Jones
Diptera Meigenia mutabilis 8 SO90379448 28/07/2011 Nigel Jones
Diptera Meiosimyza decipiens site record SO898945 28/07/2011 Nigel Jones
Diptera Meiosimyza rorida site record SO898945 28/07/2011 Nigel Jones
Diptera Melanogaster hirtella 9 SO90259456 20/05/2011 Nigel Jones
Diptera Melanostoma mellinum 9 SO90259456 28/07/2011 Nigel Jones
Diptera Melanostoma scalare 5 SO89599442 18/04/2011 Nigel Jones
Diptera Melanostoma scalare 5 SO89599442 20/05/2011 Nigel Jones
Diptera Melanostoma scalare 8 SO90379448 20/05/2011 Nigel Jones
Diptera Melanostoma scalare 5 SO89599442 28/07/2011 Nigel Jones
Diptera Melanostoma scalare 8 SO90379448 28/07/2011 Nigel Jones
Diptera Melanostoma scalare 9 SO90259456 28/07/2011 Nigel Jones
Diptera Merodon equestris 8 SO90379448 20/05/2011 Nigel Jones
Diptera Mesembrina meridiana 9 SO90259456 15/09/2011 Pete Boardman
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 25
Diptera Microchrysa flavicornis 8 SO90379448 28/07/2011 Nigel Jones
Diptera Microchrysa polita 8 SO90379448 20/05/2011 Nigel Jones
Diptera Microphor holosericeus 5 SO89599442 20/05/2011 Nigel Jones
Diptera Minettia fasciata site record SO898945 28/07/2011 Nigel Jones
Diptera Minettia inusta 8 SO90379448 28/07/2011 Nigel Jones
Diptera Minettia longipennis site record SO898945 20/05/2011 Nigel Jones
Diptera Minettia longipennis site record SO898945 28/07/2011 Nigel Jones
Diptera Molophilus cinereifrons 5 SO89599442 20/05/2011 Pete Boardman
Diptera Molophilus obscurus 1 SO89709483 20/05/2011 Pete Boardman
Diptera Molophilus obscurus 9 SO90259456 29/06/2011 Pete Boardman
Diptera Morellia hortorum 8 SO90379448 28/07/2011 Nigel Jones
Diptera Muscina levida 8 SO90379448 28/07/2011 Nigel Jones
Diptera Myathropa florea site record SO898945 20/05/2011 Nigel Jones
Diptera Nanna fasciata 2 SO89629479 18/04/2011 Nigel Jones
Diptera Nanna flavipes 2 SO89629479 18/04/2011 Nigel Jones
Diptera Neoascia podagrica 5 SO89599442 18/04/2011 Nigel Jones
Diptera Neoascia podagrica 5 SO89599442 28/07/2011 Nigel Jones
Diptera Nephrotoma appendiculata 8 SO90379448 20/05/2011 Pete Boardman
Diptera Nephrotoma appendiculata 9 SO90259456 29/06/2011 Pete Boardman
Diptera Nephrotoma quadrifaria 9 SO90259456 29/06/2011 Pete Boardman
Diptera Neria cibaria 8 SO90379448 20/05/2011 Nigel Jones
Diptera Neuroctena anilis 5 SO89599442 20/05/2011 Nigel Jones
Diptera Notiphila maculata 9 SO90259456 28/07/2011 Nigel Jones
Diptera Ocydromia glabricula 5 SO89599442 20/05/2011 Nigel Jones
Diptera Oedalea stigmatella 8 SO90379448 20/05/2011 Nigel Jones
Diptera Opomyza germinationis 5 SO89599442 20/05/2011 Nigel Jones
Diptera Opomyza petrei 8 SO90379448 28/07/2011 Nigel Jones
Diptera Ormosia lineata 11 SO89649435 20/04/2011 Pete Boardman
Diptera Ormosia lineata 5 SO89599442 20/05/2011 Pete Boardman
Diptera Ormosia lineata 7 SO90309478 20/05/2011 Pete Boardman
Diptera Oxycera rara 9 SO90259456 29/06/2011 Pete Boardman
Diptera Pachygaster leachii 8 SO90379448 28/07/2011 Nigel Jones
Diptera Palloptera scutellata 1 SO89709483 18/04/2011 Nigel Jones
Diptera Palloptera scutellata 5 SO89599442 18/04/2011 Nigel Jones
Diptera Palloptera scutellata 7 SO90309478 18/04/2011 Nigel Jones
Diptera Palloptera scutellata 9 SO90259456 18/04/2011 Nigel Jones
Diptera Parydra coarctata 5 SO89599442 20/05/2011 Nigel Jones
Diptera Parydra coarctata 9 SO90259456 28/07/2011 Nigel Jones
Diptera Pelatachina tibialis site record SO898945 28/07/2011 Nigel Jones
Diptera Phaonia errans 8 SO90379448 28/07/2011 Nigel Jones
Diptera Phaonia fuscata site record SO898945 28/07/2011 Nigel Jones
Diptera Phaonia subventa 8 SO90379448 28/07/2011 Nigel Jones
Diptera Phasia obesa site record SO898945 20/05/2011 Nigel Jones
Diptera Phasia pusilla 5 SO89599442 20/05/2011 Nigel Jones
Diptera Pherbellia schoenherri site record SO898945 28/07/2011 Nigel Jones
Diptera Pilaria discicollis 9 SO90259456 29/06/2011 Pete Boardman
Diptera Platycheirus clypeatus 9 SO90259456 28/07/2011 Nigel Jones
Diptera Platycheirus granditarsus 9 SO90259456 28/07/2011 Nigel Jones
Diptera Platycheirus manicatus 9 SO90259456 20/05/2011 Nigel Jones
Diptera Platycheirus peltatus 9 SO90259456 28/07/2011 Nigel Jones
Diptera Platycheirus tarsalis 9 SO90259456 20/05/2011 Nigel Jones
Diptera Platycheirus tarsalis 5 SO89599442 28/07/2011 Nigel Jones
Diptera Platypalpus longicornis 8 SO90379448 20/05/2011 Nigel Jones
Diptera Poecilobothrus nobilitatus 9 SO90259456 29/06/2011 Pete Boardman
Diptera Poecilobothrus nobilitatus 8 SO90379448 28/07/2011 Nigel Jones
Diptera Portevinia maculata 8 SO90379448 20/05/2011 Nigel Jones
Diptera Psila fimetaria site record SO898945 20/05/2011 Nigel Jones
Diptera Ptychoptera albimana 5 SO89599442 20/04/2011 Pete Boardman
Diptera Ptychoptera albimana 8 SO90379448 20/04/2011 Pete Boardman
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 26
Diptera Ptychoptera albimana 5 SO89599442 20/05/2011 Pete Boardman
Diptera Ptychoptera albimana 8 SO90379448 20/05/2011 Pete Boardman
Diptera Rhagio lineola 8 SO90379448 28/07/2011 Nigel Jones
Diptera Rhagio scolopaceus 8 SO90379448 20/05/2011 Nigel Jones
Diptera Rhamphomyia albipennis 1 SO89709483 18/04/2011 Nigel Jones
Diptera Rhamphomyia albohirta 5 SO89599442 18/04/2011 Nigel Jones
Diptera Rhamphomyia albohirta 8 SO90379448 20/05/2011 Nigel Jones
Diptera Rhamphomyia nigripennis 5 SO89599442 20/05/2011 Nigel Jones
Diptera Rhamphomyia subcinerascens 8 SO90379448 18/04/2011 Nigel Jones
Diptera Rhamphomyia subcinerascens 9 SO90259456 18/04/2011 Nigel Jones
Diptera Rhamphomyia sulcata 7 SO90309478 18/04/2011 Nigel Jones
Diptera Rhamphomyia umbripennis 5 SO89599442 18/04/2011 Nigel Jones
Diptera Rhamphomyia umbripennis 7 SO90309478 18/04/2011 Nigel Jones
Diptera Rhaphium brevicorne 8 SO90379448 28/07/2011 Nigel Jones
Diptera Rhaphium caliginosum 8 SO90379448 18/04/2011 Nigel Jones
Diptera Rhingia campestris 7 SO90029443 20/04/2011 Pete Boardman
Diptera Rhingia campestris 8 SO90379448 20/05/2011 Nigel Jones
Diptera Sapromyza quadricincta 8 SO90379448 20/05/2011 Nigel Jones
Diptera Sapromyza sexpunctata site record SO898945 28/07/2011 Nigel Jones
Diptera Scaeva pyrastri 8a SO903945 28/07/2011 Ian Cheeseborough
Diptera Scatella tenuicosta 9 SO90259456 28/07/2011 Nigel Jones
Diptera Scathophaga furcata site record SO898945 28/07/2011 Nigel Jones
Diptera Scathophaga lutaria 8 SO90379448 18/04/2011 Nigel Jones
Diptera Scathophaga stercoraria site record SO898945 20/05/2011 Nigel Jones
Diptera Sepedon sphegea 1 SO89709483 18/04/2011 Nigel Jones
Diptera Sepedon sphegea 1 SO89709483 20/04/2011 Pete Boardman
Diptera Sepsis fulgens site record SO898945 28/07/2011 Nigel Jones
Diptera Sepsis punctum site record SO898945 28/07/2011 Nigel Jones
Diptera Sicus ferrugineus site record SO898945 20/06/2011 Nigel Jones
Diptera Siphona geniculata 8 SO90379448 28/07/2011 Nigel Jones
Diptera Spania nigra 5 SO89599442 20/05/2011 Nigel Jones
Diptera Sphaerophoria interrupta site record SO898945 28/07/2011 Nigel Jones
Diptera Sphaerophoria scripta 9 SO90259456 15/09/2011 Pete Boardman
Diptera Sphaerophoria taeniata 2 SO89629479 28/07/2011 Nigel Jones
Diptera Sphegina clunipes 8 SO90379448 20/05/2011 Nigel Jones
Diptera Sybistroma obscurellum 8 SO90379448 28/07/2011 Nigel Jones
Diptera Sympycnus desoutteri 5 SO89599442 20/05/2011 Nigel Jones
Diptera Sympycnus desoutteri 9 SO90259456 20/05/2011 Nigel Jones
Diptera Syntormon macula 8 SO90379448 18/04/2011 Nigel Jones
Diptera Syntormon macula 8 SO90379448 20/05/2011 Nigel Jones
Diptera Syritta pipiens site record SO898945 20/05/2011 Nigel Jones
Diptera Syrphus vitripennis site record SO898945 28/07/2011 Nigel Jones
Diptera Tachina fera 9a SO90269452 20/05/2011 Ian Cheeseborough
Diptera Tachina fera 9 SO90259456 15/09/2011 Pete Boardman
Diptera Tephritis cometa 9 SO90259456 20/05/2011 Nigel Jones
Diptera Terellia ruficauda 8 SO90379448 28/07/2011 Nigel Jones
Diptera Terellia ruficauda 9 SO90259456 28/07/2011 Nigel Jones
Diptera Tetanocera elata site record SO898945 28/07/2011 Nigel Jones
Diptera Tetanocera ferruginea site record SO898945 28/07/2011 Nigel Jones
Diptera Tetanocera fuscinervis 8 SO90379448 20/05/2011 Nigel Jones
Diptera Teuchophorus calcaratus 8 SO90379448 28/07/2011 Nigel Jones
Diptera Teuchophorus monacanthus 9 SO90259456 20/05/2011 Nigel Jones
Diptera Teuchophorus monacanthus 8 SO90379448 28/07/2011 Nigel Jones
Diptera Teuchophorus nigricosta 8 SO90379448 28/07/2011 Nigel Jones
Diptera Teuchophorus simplex 8 SO90379448 28/07/2011 Nigel Jones
Diptera Themira annulipes site record SO898945 28/07/2011 Nigel Jones
Diptera Thricops diaphana 8 SO90379448 28/07/2011 Nigel Jones
Diptera Tipula fascipennis 9 SO90259456 29/06/2011 Pete Boardman
Diptera Tipula lateralis 11 SO89649435 20/04/2011 Pete Boardman
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 27
Diptera Tipula lateralis 9 SO90259456 29/06/2011 Pete Boardman
Diptera Tipula paludosa 1 SO89709483 15/09/2011 Pete Boardman
Diptera Tipula paludosa 9 SO90259456 15/09/2011 Pete Boardman
Diptera Tipula paludosa 9 SO90259456 15/09/2011 Pete Boardman
Diptera Tipula paludosa site record SO898945 15/09/2011 Pete Boardman
Diptera Tipula submarmorata site record SO89399444 20/05/2011 Pete Boardman
Diptera Tipula vittata 9 SO90259456 18/04/2011 Pete Boardman
Diptera Tipula vittata 5 SO89599442 20/04/2011 Pete Boardman
Diptera Tipula vittata 7 SO90309478 20/04/2011 Pete Boardman
Diptera Tipula vittata 9 SO90259456 20/04/2011 Pete Boardman
Diptera Tipula vittata 9 SO90259456 20/04/2011 Pete Boardman
Diptera Trichopsomyia flavitarsis 9 SO90259456 20/05/2011 Nigel Jones
Diptera Tricyphona immaculata 5 SO89599442 20/04/2011 Pete Boardman
Diptera Tricyphona immaculata 5 SO89599442 20/05/2011 Pete Boardman
Diptera Tricyphona immaculata 8 SO90379448 20/05/2011 Pete Boardman
Diptera Urophora cardui site record SO898945 28/07/2011 Nigel Jones
Diptera Urophora jaceana 8 SO90379448 29/06/2011 Pete Boardman
Diptera Urophora jaceana site record SO898945 28/07/2011 Nigel Jones
Diptera Vibrissina debilitata site record SO898945 28/07/2011 Nigel Jones
Diptera Volucella bombylans site record SO898945 20/05/2011 Nigel Jones
Diptera Xyphosia miliaria site record SO898945 28/07/2011 Nigel Jones
Hemiptera Acanthosoma haemorrhoidale 1 SO89709483 15/09/2011 Pete Boardman
Hemiptera Capsus ater 3 SO89529444 2011 Richard Wright
Hemiptera Coreus marginatus 9a SO90269452 20/06/2011 Ian Cheeseborough
Hemiptera Deraeocoris lutescens 7 SO90309478 2011 Richard Wright
Hemiptera Elasmostethus interstinctus site record SO898945 20/05/2011 Nigel Jones
Hemiptera Himacerus mirmicoides 4 SO89459440 2011 Richard Wright
Hemiptera Ischnodemus sabuleti 5 SO89599442 2011 Richard Wright
Hemiptera Leptopterna dolabrata 2 SO89629479 2011 Richard Wright
Hemiptera Leptopterna dolabrata 3 SO89529444 2011 Richard Wright
Hemiptera Liocoris tripustulatus 7 SO90309478 2011 Richard Wright
Hemiptera Nabis rugosus 2 SO89629479 2011 Richard Wright
Hemiptera Nabis rugosus 3 SO89529444 2011 Richard Wright
Hemiptera Nabis rugosus 4 SO89459440 2011 Richard Wright
Hemiptera Nabis rugosus 6 SO89799476 2011 Richard Wright
Hemiptera Nabis rugosus 7 SO90309478 2011 Richard Wright
Hemiptera Nabis rugosus 8 SO90379448 2011 Richard Wright
Hemiptera Notostira elongata 3 SO89529444 2011 Richard Wright
Hemiptera Orthotylus ericetorum 4 SO89459440 2011 Richard Wright
Hemiptera Orthotylus ericetorum 6 SO89799476 2011 Richard Wright
Hemiptera Palomena prasina 7 SO90309478 2011 Richard Wright
Hemiptera Palomena prasina site record SO899945 18/04/2011 Ian Cheeseborough
Hemiptera Palomena prasina 9a SO90269452 20/05/2011 Ian Cheeseborough
Hemiptera Piezodorus lituratus site record SO902948 18/04/2011 Ian Cheeseborough
Hemiptera Pithanus maerkelii 9 SO90259456 29/06/2011 Pete Boardman
Hemiptera Scolopostethus decoratus 4 SO89459440 2011 Richard Wright
Hemiptera Scolopostethus decoratus 6 SO89799476 2011 Richard Wright
Hemiptera Stenodema calcarata 3 SO89529444 2011 Richard Wright
Hemiptera Stenotus binotatus 3 SO89529444 2011 Richard Wright
Hemiptera Stygnocoris pedestris 3 SO89529444 2011 Richard Wright
Hemiptera Stygnocoris rusticus 4 SO89459440 2011 Richard Wright
Hemiptera Stygnocoris rusticus 8 SO90379448 2011 Richard Wright
Hemiptera Stygnocoris rusticus 9 SO90259456 2011 Richard Wright
Hemiptera Velia caprai 5 SO89599442 2011 Richard Wright
Hymenoptera Ancistrocerus gazella 8a SO903944 28/07/2011 Ian Cheeseborough
Hymenoptera Andrena barbilabris 11 SO89649435 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena bicolor site record SO899948 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena bicolor 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Andrena chrysosceles 12 SO90349456 18/04/2011 Ian Cheeseborough
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 28
Hymenoptera Andrena cineraria 11 SO89649435 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena cineraria site record SO899948 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena dorsata 8a SO903944 28/07/2011 Ian Cheeseborough
Hymenoptera Andrena fucata 8a SO903944 20/05/2011 Ian Cheeseborough
Hymenoptera Andrena fulva 11 SO89649435 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena fulva 12 SO90349456 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena fulva site record SO899945 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena haemorrhoa 11 SO89649435 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena haemorrhoa 12 SO90349456 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena haemorrhoa site record SO899945 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena haemorrhoa site record SO901944 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena haemorrhoa site record SO898945 20/05/2011 Ian Cheeseborough
Hymenoptera Andrena nigroaenea 7 SO90309478 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena nigroaenea site record SO901944 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena scotica 8a SO903945 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena scotica site record SO899945 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena scotica site record SO901944 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena subopaca 8a SO903944 18/04/2011 Ian Cheeseborough
Hymenoptera Andrena wilkella 10 SO89639441 28/07/2011 Ian Cheeseborough
Hymenoptera Andrena wilkella 8a SO903944 28/07/2011 Ian Cheeseborough
Hymenoptera Anoplius caviventris site record SO898945 20/05/2011 Ian Cheeseborough
Hymenoptera Anoplius nigerrimus 9a SO90269452 28/07/2011 Ian Cheeseborough
Hymenoptera Anthidium manicatum 10 SO89639441 28/07/2011 Ian Cheeseborough
Hymenoptera Anthophora furcata 8a SO903944 29/06/2011 Ian Cheeseborough
Hymenoptera Anthophora furcata 9a SO90269452 28/07/2011 Ian Cheeseborough
Hymenoptera Argogorytes mystaceus 8a SO903944 20/05/2011 Ian Cheeseborough
Hymenoptera Bombus hortorum site record SO899943 29/06/2011 Ian Cheeseborough
Hymenoptera Bombus hypnorum 9a SO90269452 20/05/2011 Ian Cheeseborough
Hymenoptera Bombus hypnorum site record SO898945 20/05/2011 Ian Cheeseborough
Hymenoptera Bombus hypnorum site record SO898945 20/05/2011 Nigel Jones
Hymenoptera Bombus hypnorum 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Bombus lapidarius site record SO901944 18/04/2011 Ian Cheeseborough
Hymenoptera Bombus lapidarius site record SO898945 20/05/2011 Ian Cheeseborough
Hymenoptera Bombus lapidarius 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Bombus lapidarius 10 SO89639441 28/07/2011 Ian Cheeseborough
Hymenoptera Bombus lucorum 12 SO90349456 18/04/2011 Ian Cheeseborough
Hymenoptera Bombus lucorum 9a SO90269452 20/05/2011 Ian Cheeseborough
Hymenoptera Bombus lucorum 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Bombus lucorum 10 SO89639441 28/07/2011 Ian Cheeseborough
Hymenoptera Bombus pascuorum 7 SO90309478 18/04/2011 Ian Cheeseborough
Hymenoptera Bombus pascuorum site record SO898945 20/05/2011 Ian Cheeseborough
Hymenoptera Bombus pascuorum 8a SO903944 29/06/2011 Ian Cheeseborough
Hymenoptera Bombus pascuorum 10 SO89639441 28/07/2011 Ian Cheeseborough
Hymenoptera Bombus pratorum 9a SO90269452 20/05/2011 Ian Cheeseborough
Hymenoptera Bombus pratorum site record SO898945 20/05/2011 Ian Cheeseborough
Hymenoptera Bombus pratorum 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Bombus sylvestris 9a SO90269452 20/05/2011 Ian Cheeseborough
Hymenoptera Bombus sylvestris site record SO898945 20/05/2011 Ian Cheeseborough
Hymenoptera Bombus terrestris 12 SO90349456 18/04/2011 Ian Cheeseborough
Hymenoptera Bombus terrestris site record SO899945 18/04/2011 Ian Cheeseborough
Hymenoptera Bombus terrestris 9a SO90269452 20/05/2011 Ian Cheeseborough
Hymenoptera Bombus terrestris 10 SO89639441 28/07/2011 Ian Cheeseborough
Hymenoptera Bombus vestalis site record SO899945 18/04/2011 Ian Cheeseborough
Hymenoptera Crossocerus annulipes 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Crossocerus capitosus 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Crossocerus cetratus 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Crossocerus megacephalus 9a SO90269452 28/07/2011 Ian Cheeseborough
Hymenoptera Crossocerus nigritus 8a SO903944 20/05/2011 Ian Cheeseborough
Hymenoptera Crossocerus podagricus 9a SO90269452 28/07/2011 Ian Cheeseborough
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 29
Hymenoptera Crossocerus quadrimaculatus 8 SO90379448 20/06/2011 Ian Cheeseborough
Hymenoptera Dolichovespula norwegica 8a SO903944 29/06/2011 Ian Cheeseborough
Hymenoptera Dolichovespula sylvestris 8a SO903944 29/06/2011 Ian Cheeseborough
Hymenoptera Ectemnius cavifrons 9a SO90269452 28/07/2011 Ian Cheeseborough
Hymenoptera Ectemnius continuus 8a SO903944 20/05/2011 Ian Cheeseborough
Hymenoptera Ectemnius continuus 8a SO903944 29/06/2011 Ian Cheeseborough
Hymenoptera Ectemnius continuus 9a SO90269452 28/07/2011 Ian Cheeseborough
Hymenoptera Halictus rubicundus 8a SO903944 29/06/2011 Ian Cheeseborough
Hymenoptera Halictus tumulorum 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Hylaeus communis 9 SO90259456 28/07/2011 Nigel Jones
Hymenoptera Hylaeus communis 9a SO90269452 28/07/2011 Ian Cheeseborough
Hymenoptera Lasioglossum calceatum 10 SO89639441 28/07/2011 Ian Cheeseborough
Hymenoptera Lasioglossum leucopus 8a SO903945 28/07/2011 Ian Cheeseborough
Hymenoptera Lasioglossum villosulum 8a SO903945 18/04/2011 Ian Cheeseborough
Hymenoptera Lasius flavus 7 SO90309478 18/04/2011 Ian Cheeseborough
Hymenoptera Lasius niger site record SO899948 18/04/2011 Ian Cheeseborough
Hymenoptera Lindenius albilabris site record SO899943 29/06/2011 Ian Cheeseborough
Hymenoptera Lindenius albilabris 8a SO903944 28/07/2011 Ian Cheeseborough
Hymenoptera Megachile ligniseca 8a SO903945 29/06/2011 Ian Cheeseborough
Hymenoptera Megachile willughbiella 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Megachile willughbiella 10 SO89639441 28/07/2011 Ian Cheeseborough
Hymenoptera Megachile willughbiella 8a SO903944 28/07/2011 Ian Cheeseborough
Hymenoptera Mimumesa dahlbomi 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Myrmosa atra site record SO899943 29/06/2011 Ian Cheeseborough
Hymenoptera Nomada fabriciana 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Nomada flava site record SO901944 18/04/2011 Ian Cheeseborough
Hymenoptera Nomada flava/panzeri 7 SO90309478 18/04/2011 Ian Cheeseborough
Hymenoptera Nomada goodeniana 7 SO90309478 18/04/2011 Ian Cheeseborough
Hymenoptera Nomada lathburiana 12 SO90349456 18/04/2011 Ian Cheeseborough
Hymenoptera Nomada ruficornis 11 SO89649435 18/04/2011 Ian Cheeseborough
Hymenoptera Nomada ruficornis 12 SO90349456 18/04/2011 Ian Cheeseborough
Hymenoptera Osmia caerulescens 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Osmia rufa 12 SO90349456 18/04/2011 Ian Cheeseborough
Hymenoptera Osmia rufa site record SO899945 18/04/2011 Ian Cheeseborough
Hymenoptera Osmia rufa site record SO901944 20/06/2011 Ian Cheeseborough
Hymenoptera Passaloecus gracilis 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Passaloecus singularis 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Pemphredon lugubris 8 SO90379448 20/05/2011 Nigel Jones
Hymenoptera Pemphredon lugubris 9a SO90269452 28/07/2011 Ian Cheeseborough
Hymenoptera Psenulus concolor 8a SO903944 20/05/2011 Ian Cheeseborough
Hymenoptera Psenulus pallipes 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Rhopalum clavipes 9a SO90269452 28/07/2011 Ian Cheeseborough
Hymenoptera Rhopalum clavipes site record SO898945 28/07/2011 Nigel Jones
Hymenoptera Sphecodes ephippius 12 SO90349456 18/04/2011 Ian Cheeseborough
Hymenoptera Stigmus solskyi 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Tiphia minuta 8a SO903945 29/06/2011 Ian Cheeseborough
Hymenoptera Trichrysis cyanea 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Trypoxylon attenuatum 9a SO90269452 20/05/2011 Ian Cheeseborough
Hymenoptera Trypoxylon attenuatum 9a SO90269452 20/06/2011 Ian Cheeseborough
Hymenoptera Trypoxylon attenuatum 8a SO903945 29/06/2011 Ian Cheeseborough
Hymenoptera Trypoxylon clavicerum 9a SO90269452 28/07/2011 Ian Cheeseborough
Hymenoptera Vespula germanica 7 SO90309478 18/04/2011 Ian Cheeseborough
Hymenoptera Vespula germanica 9a SO90269452 28/07/2011 Ian Cheeseborough
Hymenoptera Vespula rufa site record SO898945 20/05/2011 Ian Cheeseborough
Hymenoptera Vespula rufa 8a SO903944 29/06/2011 Ian Cheeseborough
Hymenoptera Vespula vulgaris 11 SO89649435 18/04/2011 Ian Cheeseborough
Hymenoptera Vespula vulgaris 8a SO903945 29/06/2011 Ian Cheeseborough
Isopoda Oniscus asellus 7 SO90309478 2011 Richard Wright
Isopoda Philoscia muscorum 1 SO89709483 2011 Richard Wright
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 30
Isopoda Philoscia muscorum 2 SO89629479 2011 Richard Wright
Isopoda Philoscia muscorum 3 SO89529444 2011 Richard Wright
Isopoda Philoscia muscorum 4 SO89459440 2011 Richard Wright
Isopoda Philoscia muscorum 6 SO89799476 2011 Richard Wright
Isopoda Philoscia muscorum 8 SO90379448 2011 Richard Wright
Isopoda Philoscia muscorum 9 SO90259456 2011 Richard Wright
Isopoda Trichoniscus pusillus 1 SO89709483 2011 Richard Wright
Isopoda Trichoniscus pusillus 2 SO89629479 2011 Richard Wright
Isopoda Trichoniscus pusillus 6 SO89799476 2011 Richard Wright
Isopoda Trichoniscus pusillus 9 SO90259456 2011 Richard Wright
Lepidoptera Aglais urticae 8 SO90379448 20/04/2011 Pete Boardman
Lepidoptera Aglais urticae 9a SO90269452 20/05/2011 Ian Cheeseborough
Lepidoptera Aglais urticae 9 SO90259456 29/06/2011 Pete Boardman
Lepidoptera Anthocharis cardamines site record SO902948 18/04/2011 Ian Cheeseborough
Lepidoptera Anthocharis cardamines 8 SO90379448 20/04/2011 Pete Boardman
Lepidoptera Anthocharis cardamines 9 SO90259456 20/04/2011 Pete Boardman
Lepidoptera Aphantopus hyperantus 9a SO90269452 20/06/2011 Ian Cheeseborough
Lepidoptera Aphantopus hyperantus 9 SO90259456 29/06/2011 Pete Boardman
Lepidoptera Aphantopus hyperantus 8a SO903945 28/07/2011 Ian Cheeseborough
Lepidoptera Callophrys rubi 9a SO90269452 20/05/2011 Ian Cheeseborough
Lepidoptera Coenonympha pamphilus 9a SO90269452 20/05/2011 Ian Cheeseborough
Lepidoptera Coenonympha pamphilus 9a SO90269452 20/06/2011 Ian Cheeseborough
Lepidoptera Inachis io 8 SO90379448 20/04/2011 Pete Boardman
Lepidoptera Inachis io 9a SO90269452 20/06/2011 Ian Cheeseborough
Lepidoptera Inachis io 8a SO903944 28/07/2011 Ian Cheeseborough
Lepidoptera Lycaena phlaeas 9a SO90269452 28/07/2011 Ian Cheeseborough
Lepidoptera Maniola jurtina 9 SO90259456 29/06/2011 Pete Boardman
Lepidoptera Ochlodes venata 9a SO90269452 20/06/2011 Ian Cheeseborough
Lepidoptera Ochlodes venata 9 SO90259456 29/06/2011 Pete Boardman
Lepidoptera Pararge aegeria 7 SO90309478 20/04/2011 Pete Boardman
Lepidoptera Pararge aegeria 9a SO90269452 20/05/2011 Ian Cheeseborough
Lepidoptera Pararge aegeria 9 SO90259456 29/06/2011 Pete Boardman
Lepidoptera Pararge aegeria 9 SO90259456 15/09/2011 Pete Boardman
Lepidoptera Pieris napi 8 SO90379448 20/04/2011 Pete Boardman
Lepidoptera Pieris napi 9a SO90269452 20/05/2011 Ian Cheeseborough
Lepidoptera Pieris rapae 9a SO90269452 20/06/2011 Ian Cheeseborough
Lepidoptera Pieris rapae 9a SO90269452 28/07/2011 Ian Cheeseborough
Lepidoptera Polygonia c-album 8 SO90379448 20/04/2011 Pete Boardman
Lepidoptera Pyronia tithonus 9a SO90269452 28/07/2011 Ian Cheeseborough
Lepidoptera Stigmella aurella 7 SO90029443 20/04/2011 Pete Boardman
Lepidoptera Thymelicus sylvestris 9a SO90269452 20/06/2011 Ian Cheeseborough
Odonata Coenagrion puella 9a SO90269452 20/05/2011 Ian Cheeseborough
Odonata Coenagrion puella 9a SO90269452 20/06/2011 Ian Cheeseborough
Odonata Ischnura elegans 9 SO90259456 29/06/2011 Pete Boardman
Odonata Pyrrhosoma nymphula 9a SO90269452 20/05/2011 Ian Cheeseborough
Odonata Sympetrum striolatum 1 SO89709483 15/09/2011 Pete Boardman
Opiliones Mitopus morio 2 SO89629479 2011 Richard Wright
Opiliones Nemastoma bimaculatum 2 SO89629479 2011 Richard Wright
Opiliones Nemastoma bimaculatum 5 SO89599442 2011 Richard Wright
Opiliones Nemastoma bimaculatum 7 SO90309478 2011 Richard Wright
Opiliones Nemastoma bimaculatum 9 SO90259456 2011 Richard Wright
Opiliones Paroligolophus agrestis 1 SO89709483 2011 Richard Wright
Opiliones Paroligolophus agrestis 3 SO89529444 2011 Richard Wright
Opiliones Paroligolophus agrestis 9 SO90259456 2011 Richard Wright
Opiliones Rilaena triangularis 1 SO89709483 2011 Richard Wright
Opiliones Rilaena triangularis 3 SO89529444 2011 Richard Wright
Opiliones Rilaena triangularis 4 SO89459440 2011 Richard Wright
Orthoptera Omocestus viridulus 9a SO90269452 20/06/2011 Ian Cheeseborough
Orthoptera Tetrix subulata 5 SO89599442 2011 Richard Wright
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 31
Orthoptera Tetrix subulata 8 SO90379448 2011 Richard Wright
Orthoptera Tetrix subulata 9 SO90259456 2011 Richard Wright
Orthoptera Tetrix subulata 5 SO89599442 20/04/2011 Pete Boardman
Appendix 2 – ISIS outputs for Penn Common
WHOLE SITE
SAT
code SAT name No. spp. Condition
% of national species
pool
Related BAT rarity
score
F001 scrub edge 19 fav 11
F002 rich flower resource 15 fav 6
A212 bark & sapwood decay 23 fav 5 131
F006 dung 3 3
F112 open short sward 4 2 120
W111 shingle banks 1 2 154
W126 seepage 1 2 154
W314 reedfen and pools 2 2 135
F003 scrub-heath & moorland 5 1
F111 bare sand & chalk 5 1 120
A211 heartwood decay 1 1 131
BAT code BAT name
Representation (1-100)
Rarity score Condition BAT species richness IEC
F2 grassland & scrub matrix 34 115 154
F1
unshaded early successional
mosaic 11 120 51
W3 permanent wet mire 10 135 44
W1 flowing water 8 154 fav 35
A2 wood decay 7 131 30 0
F3 shaded field & ground layer 6 130 29
W2 mineral marsh & open water 5 112 25
A1 arboreal canopy 3 13
CALCAREOUS FLUSH
SAT
code SAT name No. spp. Condition
% of national species
pool
Related BAT rarity
score
W111 shingle banks 1 2 132
BAT
code BAT name
Representation
(1-100)
Rarity
score Condition BAT species richness IEC
F2 grassland & scrub matrix 22 127 22
W1 flowing water 19 132 19
W2 mineral marsh & open water 18 122 18
F3 shaded field & ground layer 11 11
W3 permanent wet mire 10 10
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 32
WET HABITATS IN EAST OF SITE
SAT
code SAT name No. spp. Condition
% of national species
pool
Related BAT rarity
score
W126 seepage 2 4 177
F112 open short sward 3 2
F001 scrub edge 2 1
W314 reedfen and pools 1 1 121
A212 bark & sapwood decay 3 1
F002 rich flower resource 1 0
F003 scrub-heath & moorland 1 0
F111 bare sand & chalk 1 0
BAT
code BAT name
Representation
(1-100)
Rarity
score Condition BAT species richness IEC
F2 grassland & scrub matrix 29 116 73
W3 permanent wet mire 16 121 40
W1 flowing water 12 177 fav 31
W2 mineral marsh & open water 11 118 28
F3 shaded field & ground layer 6 15
F1 unshaded early successional mosaic 3 8
A2 wood decay 3 8 0
A1 arboreal canopy 1 3
DRY ACID GRASSLAND
SAT
code SAT name No. spp. Condition
% of national species
pool
Related BAT rarity
score
F003 scrub-heath & moorland 10 fav 3
F111 bare sand & chalk 3 1
F001 scrub edge 1 1
F112 open short sward 1 1
BAT code BAT name
Representation (1-100)
Rarity score Condition BAT species richness IEC
F2 grassland & scrub matrix 63 116 92
F1
unshaded early successional
mosaic 5 8
W2 mineral marsh & open water 4 6
F3 shaded field & ground layer 3 4
W3 permanent wet mire 3 4
W1 flowing water 1 1
DAMP GRASSLAND
SAT code SAT name No. spp. Condition
% of national species pool
Related BAT rarity score
BAT code BAT name
Representation (1-100)
Rarity score Condition BAT species richness IEC
F2 grassland & scrub matrix 47 114 43
W3 permanent wet mire 16 15
W2 mineral marsh & open water 4 4
F3 shaded field & ground layer 2 2
W1 flowing water 2 2
Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)
Page 33
ACULEATE HABITATS
SAT code SAT name No. spp. Condition
% of national species pool
Related BAT rarity score
A212 bark & sapwood decay 26 fav 5 119
F002 rich flower resource 8 3
F006 dung 3 3
F001 scrub edge 5 3
F112 open short sward 2 1 104
BAT code BAT name
Representation (1-100)
Rarity score Condition BAT species richness IEC
F1 unshaded early successional mosaic 29 104 28
A2 wood decay 28 119 27 0
F2 grassland & scrub matrix 21 100 20
A1 arboreal canopy 2 2
W2 mineral marsh & open water 2 2