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BIOL 205: COURSE INTRODUCTION & REVIEW
* COMMENTS ON ENROLLMENT:
* SYLLABUS:
A contract between you and meThe course webpageExams & gradingLecture sketches – cartoons & color highlight salient characteristics, e.g.:
* COURSE OBJECTIVES & OUTLINE:
A comprehensive look at the diversity of animals:Functional morphologyOntogenyPhylogeny
Learn the language of comparative organismal biology: The GlossaryBuild foundation for introspection, discovery, innovation & public policyLectures: Focus on introducing the building blocks of animal diversity;
frequent slide shows focused on putting the blocks together
* PARTIAL CLASSIFICATION OF INVERTEBRATES:
The taxa you will need to know – download this list from the course webpage.
* HOW TO USE THE TEXTBOOK:
B & B: Readings are listed on the syllabusChapters 2—4 & 24 are very important – consult them throughout the termFamiliarize yourself with the drawings & images in the remaining chaptersDO NOT overwhelm yourself with too many details – the text is a resourcePlease focus on the specific topics & terms highlighted in your lecture notes
pinna
pairedappendages
binocular eyehair on top of head
external naris
opposablethumb
labium
tool
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BIOL 205: COURSE INTRODUCTION & REVIEW
* THE LAB:
The ‘block learning’ component of the courseGet hands-on experience with a diverse array of invertebratesTake advantage of the virtual labs on the course webpageTreat all animals encountered with the utmost respect
* THE SCIENTIFIC METHOD:
“A” = Argument or Explanation
Scientific explanations:- must be logically consistent- must be naturalistic/empirical- must be demonstrative/repeatable- must be testable/falsifiable- are constrained by limitations of human perception (abilities & tools - microscopes)- accommodate/tolerate uncertainty- give rise to extremely accurate predictions- are outcomes of a self-correcting process
‘skyhook’ ‘crane’(non-science) (science)
OBSERVATION QUESTION HYPOTHESIS
EMPIRICALTEST
RESULTS
INDUCTION
THEORY DEDUCTION PREDICTIONS
Atomic TheoryUniversal Theory of GravitationTheory of Biological EvolutionTheory of Plate Tectonics
Support Hypothesis?Falsify Hypothesis?
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BIOL 205: COURSE INTRODUCTION & REVIEW
* OVERVIEW OF BIOLOGICAL EVOLUTION (VIA NATURAL SELECTION):
* CLASSICAL CRITERIA FOR HOMOLOGY:
HOMOLOGOUS CHARACTER: any trait present in two or more different lineages thatwas also present in their most recent common ancestor.
1. Unique similarity in form2. Corresponding position within a common body plan (‘bauplan’)3. The existence of intermediate states (developmental or paleontological)
* DISCOVERING PHYLOGENY:
Comparative analyses of biological information:morphological/behavioral traitsnucleotide sequences (RNA/DNA)amino acid sequences (proteins)fossils
Methods of phylogenetic analysis:parsimonymaximum likelihoodBayesian analysis
GENETIC MUTATIONSIN GERM LINE &RECOMBINATION
HERITABLEGENOTYPIC VARIATION
HERITABLE PHENOTYPICVARIATION
OVER-REPRODUCTIONLIMITED
RESOURCES
COMPETITIONDIFFERENTIAL
REPRODUCTION(NATURAL SELECTION)
PROPORTION OF ADVANTAGEOUSCHARACTERS INCREASES
POPULATIONS CHANGE OVERVAST GEOLOGICAL TIME
ORGANISMAL ADAPTATION
STASIS
GEOGRAPHIC/REPRODUCTIVEBARRIERS
CLADOGENETICSPECIATION
ANAGENETICSPECIATION
PATTERNS OFMACROEVOLUTION
& PHYLOGENY
GENETIC DRIFT
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BIOL 205: COURSE INTRODUCTION & REVIEW
* THE DIVERSITY OF EUKARYOTES – THE BIG PICTURE:
TELEOLOGY: An archaic, non-scientific (anthropocentric) idea asserting that thediversity of organisms is purposeful and is directed toward optimum design orperfection. Teleological thinking gave rise to the SCALA NATURAE (advocated byAristotle, 384-322 B.C.) – a viewpoint that is pervasive even today:
Please avoid using teleological language (e.g., “higher”, “lower”, “purpose”)when expressing yourself on exams or in class.
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INTRODUCTION TO ANIMAL ORGANIZATION
* WHAT IS AN ANIMAL?
1. EUKARYOTES2. MULTICELLULAR3. INTERNALLY DIGESTIVE HETEROTROPHS4. LOCOMOTOR ABILITY5. SENSE ORGANS & RAPID RESPONSES6. SEXUAL REPRODUCTION (DIPLOID LIFE CYLES)7. FUNDAMENTAL LEVELS OF BIOLOGICAL ORGANIZATION:
MOLECULES
ORGANELLES CELLS
TISSUES ORGANS
ORGAN SYSTEMS ORGANISMS
POPULATIONS COMMUNITIES ECOSYSTEMS
BIOMES BIOSPHERE
* OVERVIEW OF LECTURE THEMES – SYSTEM COMPATIBILITY:
1. MAJOR ANIMAL PHYLA / GENERAL BODY FORMS2. BODY WALL, SKELETAL SYSTEMS & LOCOMOTION3. FEEDING & DIGESTIVE SYSTEMS4. CIRCULATORY SYSTEMS & GAS EXCHANGE5. EXCRETORY & OSMOREGULATORY SYSTEMS6. NERVOUS SYSTEMS7. REPRODUCTION8. LIFE HISTORIES & ONTOGENY9. PHYLOGENETIC CONTEXT & CHARACTER EVOLUTION
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INTRODUCTION TO ANIMAL ORGANIZATION
* BODY SYMMETRY:1. Radial
2. Bilateral
* EMBRYONIC GERM LAYERS:1. None2. Diploblastic
A. EctodermB. Endoderm (Entoderm)
3. TriploblasticA. EctodermB. MesodermC. Endoderm (Entoderm)
* BODY CAVITIES:
Gut epithelium(Endoderm)
Gut epithelium(Endoderm)
Epidermis(Ectoderm)
Acoelomate Blastocoelomate(Pseudocoelomate)
Coelomate
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PHYLOGENETICS
* BRIEF REVIEW OF THE TERMS & CONCEPTS OF CLADISTICS:
1. PATTERNS OF DIVERSITY ARE HIERARCHICAL:A. Phylogenies
I. Matrices of comparative dataB. Classifications
I. Storage & retrieval systemII. CommunicationIII. Reflect evolutionary relationshipsVI. Taxa vs. ranks
2. PHYLOGENETIC SYSTEMATICS:A. ParsimonyB. Cladogram
I. MonophyleticII. PolyphyleticIII. Paraphyletic
C. SynapomorphiesD. SymplesiomorphiesE. Polarity & outgroupsF. Morphostasis
MAMMALS LIZARDS SNAKES BIRDS CROCODILES TURTLES
A
B
D E
1
2
3
5
6
7
8 9
10
4C
Homologies
8
INTRODUCTION TO ANIMAL ORGANIZATION
* FEEDING & DIGESTION (NUTRITION):INGESTION
DIGESTIONABSORPTION DEFECATION
ASSIMILATION
1. FEEDING STRATEGIES:A. SuspensionB. Deposit
I. DirectII. Selective
C. PredatoryI. CarnivoryII. ScavengingIII. Parasitism
D. Herbivory
2. NUTRIENTS:A. MineralsB. VitaminsC. CarbohydratesD. ProteinsE. LipidsF. Nucleic Acids
3. ORGANS:A. Buccal Cavity / PharynxB. StomachC. Digestive GlandsD. IntestinesE. Rectum
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INTRODUCTION TO ANIMAL ORGANIZATION
* CIRCULATION & GAS EXCHANGE (INTERNAL TRANSPORT):
1. OPEN:
2. CLOSED:HEART
VEINS ARTERIES
VENULES ARTERIOLES
CAPILLARIES
HEMOCOEL
INCREASE
HEART ARTERIES ARTERIOLES CAPILLARIES VENULES VEINS HEART HEMOCOEL
TOTAL X-SECTIONALAREA
BLOOD ORHEMOLYMPHVELOCITY
10
INTRODUCTION TO ANIMAL ORGANIZATION
* EXCRETION & OSMOREGULATION:
1. DEAMINATION WASTE PRODUCTS:A. Ammonia – NH3 (Ammonotelic)B. Detoxification
I. Urea (e.g. vertebrates)II. Uric acid (Uricotelic)
2. OSMOTIC CONDITIONS:A. Isotonic (e.g. marine)B. Hypotonic (e.g. freshwater)C. Hypertonic (e.g. brines)D. TerrestrialE. Osmoregulators vs. Osmoconformers
3. NEPHRIDIA:A. ProtonephridiaB. Metanephridia
* NERVOUS SYSTEM & COORDINATION:
1. FUNDAMENTAL UNITS:A. NeuronsB. Nerves
2. REFLEX ARCS:
STIMULUS RECEPTOR SITE SENSORY NERVES
NERVE CORDS & GANGLIA
RESPONSE EFFECTOR SITE MOTOR NERVES
3. RECEPTORS: GEO-, PHOTO-, PHONO-, CHEMO-, PROPRIO-,THERMO-, TACTILE, PRESSURE, PAIN
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INTRODUCTION TO ANIMAL ORGANIZATION
* REPRODUCTION:
1. GENERAL ANIMAL LIFE CYCLE:
ADULTS
MITOSIS MEIOSIS
MATURATION HAPLOID GAMETES
SPERM OVA
JUVENILE ZYGOTE
EMBRYOLOGY
2. ASEXUAL REPRODUCTION:
ADVANTAGES DISADVANTAGES • No need to find mates • No variation• Clones (if well adapted)• Faster reproduction• Physiologically cheaper
3. CONDITIONS OF SEXUAL REPRODUCTION:A. Gonochoristic = DioeciousB. Hermaphroditic = Monoecious
4. FERTILIZATION:A. External (Free spawning)B. Internal
SEXUALREPRODUCTION
ASEXUALREPRODUCTION
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INTRODUCTION TO ANIMAL ORGANIZATION
* LIFE HISTORIES:
1. Indirect
2. Direct
3. Mixed
ADULTS
JUVENILE GAMETES
METAMORPHOSIS FREE SPAWN
INTERNALFERTILIZATION
LARVAE EXTERNALFERTILIZATION
ZYGOTE
4. BASIC LARVAL TYPES:
A. Planktotrophic
B. Lecithotrophic
Indirect Low cost/zygoteLow survivalWide distribution (dispersal)Low parental care
Direct High cost/zygoteHigh survivalNarrow distribution
High parental care