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READING THE PASTORAL LANDSCAPE: PALYNOLOGICAL AND HISTORICAL
EVIDENCE FOR THE IMPACTS OF LONG-TERM GRAZING ON WETHER HILL, INGRAM,
NORTHUMBERLAND
Althea L. Davies1 and Piers Dixon2
1 Department of Environmental Science, University of Stirling, Stirling, Scotland, UK, FK9 4LA. Current
address: AHRC Centre for Environmental History, University of Stirling, Stirling, Scotland, UK, FK9
4LA; email: [email protected] Royal Commission on the Ancient and Historical Monuments of Scotland, John Sinclair House, 16
Bernard Terrace, Edinburgh, EH8 9NX; email: [email protected]
Biographies
Althea Davies studied botany and then environmental archaeology, specialising in pollen analysis. She
has a particular interest in the interactions between upland farming communities and their environment,
and most of her work has been undertaken in the Scottish Highlands. She is pursuing interdisciplinary
research involving archaeology, history, ecology and conservation management.
Piers Dixon studied archaeology and history, completing a doctorate on 'The Deserted Medieval Villages
of North Northumberland' in 1985. He has continued to develop his interests in rural settlement,
publishing the results of excavations in the Scottish Borders and Northumberland, a book on rural
settlement in lowland Scotland entitled Puir labourers and busy husbandmen, as well as synthetic articles.
Abstract
Many upland environments are valued for their openness, which is often actively maintained by extensive
pastoral agriculture. Documentary sources indicate the complexity and longevity of regulations designed
to protect the hill grazing resource from over-exploitation but these systems leave relatively few traces on
the ground. Consequently pollen analysis is an important method for establishing the impact of centuries
of grazing on the quality of hill pastures. This is demonstrated at Wether Hill, Northumberland, where a
pollen sequence details changes in vegetation composition and diversity over the last c.1500 years. These
are correlated with historical evidence over the last c.800 years for a more complete understanding of the
socio-economic context that governed the use of hill grazing. Changes in grazing regimes had a profound
influence on these hill pastures, contributing to permanent changes in the relative abundance of heather,
grasses and herbs, and causing a severe decline in habitat diversity within the last c.200 years. The results
have many regional parallels, indicating extensive reductions in the biodiversity of upland habitats. This
1
has implications for future management and conservation policies and shows the contribution that an
understanding of environmental and land-use history can make to debates surrounding current
environmental issues.
Keywords
Uplands, conservation, environmental history, heathland, grazing history, archaeology; ridge and furrow.
Introduction
To many, the openness of upland landscapes in Britain is an intrinsic part of their attraction and value, as
seen, for example, in the perceived ‘wildness’ of the Scottish Highlands or concerns that scrub could
invade the Cumbrian hills following the decline in livestock numbers during the Foot and Mouth crisis of
2001 (e.g. Luxmoore & Fenton 2005). Many upland areas have been treeless for centuries or millennia
but, far from being static, open hill habitats have experienced significant changes in vegetation
composition over this period (e.g. Stevenson & Birks 1995, Tipping 2000). The hills of Northumberland
provide a good example of the dynamics and variability of vegetation composition and related socio-
economic conditions over the last 1500 years.
There is a wealth of archaeological evidence for occupation in the Cheviot Hills and adjacent coastal
plains over the last 9000 years (Figure 1), including well-preserved field remains of cultivation from the
last c.4000 years (Topping 1989, 1991, Frodsham 2004). However, as in many upland areas, pastoralism
was probably always the most extensive form of agriculture. Documentary sources from the fifteenth
century A.D. onwards can provide detailed records of the multitude of laws governing grazing practices in
the western Borders of Cumbria (Winchester 2000). In the eastern uplands of the Cheviots, the most
significant historical aspect of this is the dominance of grazing by the landowning classes as communal
forms of transhumance were finally expunged (Dixon 1985, Vol. I). However, tracing the links between
changing patterns of use and regulation, and pasture type on the ground is a harder prospect since long-
term pastoralism may cause relatively few structural changes in the landscape and, indeed, often
contributes to the good preservation of older archaeological evidence, including cultivation ridges.
Palaeoenvironmental techniques such as pollen analysis are rarely applied in the historic period (Tipping
1998, 2004), but can provide an additional source of evidence for understanding changing patterns of
grazing. By choosing pollen sampling sites that are sensitive to changes in vegetation cover within a
radius of c.50-100 m, pollen analysts can reconstruct landscape history on local scales which are
comparable with site-focussed historical and archaeological records (Tipping 1997a, Davies & Tipping
2
2004). This is the approach adopted at Wether Hill, in the eastern foothills of the Cheviots, where the fill
of an abandoned Iron Age ditch provides a record of changing plant communities on the surrounding hill
pastures since c. A.D. 430.
The purpose of this paper is to combine pollen-based (palynological), documentary and archaeological
evidence for the impact of pastoralism on plant composition and diversity in an area of the Cheviot
foothills that today lies in the Northumberland National Park (Figure 1). The record has implications for
the presentation of landscape history to visitors, future land management and conservation.
Wether Hill: context of research and methods
Northumberland Archaeological Group has undertaken excavations on Wether Hill since 1993 as part of
the Ingram and Upper Breamish Valley Landscape Project, which also involves Northumberland National
Park and the University of Durham (Figure 1). The results indicate that these hills have been in use for
around 8000 years (Frodsham 2004). The area includes a well-preserved Iron Age hillfort on Wether Hill
(National Grid Ref. NU 013 144; 1º 59”W, 55º 25”N; 290 m OD) (Figure 2, Plate 1). The fort and
adjacent prehistoric cord rig cultivation are located near a c.300 m long, linear cross-ridge dyke that lies to
the south-west of the fort (Topping 1994). The dyke consists of a ditch flanked by two banks and was
constructed around c.200 BC (all dates are calibrated), but after c. A.D. 430 it ceased to be maintained
(Topping 1996). There is little archaeological evidence for post-Roman and Anglo-Saxon activity, but
mixed farming was conducted in the medieval period, as indicated by nearby evidence for ridge and
furrow cultivation (Topping 1995).
Damp conditions within the ditch delimiting the hill fort allowed the accumulation of a 50 cm deep,
mineral-rich peat deposit which was sampled for pollen analysis. The ditch fill is ideally located to
provide a record of the changes in vegetation associated with land-use on the hill since it lies at the centre
of a multi-period archaeological landscape (Figure 2). The base of the ditch at 50 cm is dated to 1590
60 BP (calibrated to A.D. 260-620, mid-point A.D. 430). A second radiocarbon assay from 41.5 cm
produced a very similar date (1640 60 BP, cal A.D. 260-560, mid-point cal A.D. 420), indicating that
organic material is likely to have been reworked, possibly due to the erosion of soil from the adjacent
banks by grazing livestock. However, well-drained soils provide poor conditions for pollen preservation
and there is no evidence that pollen was also reworked. Dating evidence for the upper portion of the
sequence is provided by the appearance of (1) pollen from exotic trees and conifers in the post-
Agricultural Improvement landscape (Carter et al. 1997) and (2) spheroidal carbonaceous particles
(SCPs), a by-product of fossil fuel combustion since around 1900 (Rose et al. 1995), which were counted
3
on pollen slides. The sequence therefore covers c.1500 years, from A.D. 430 to the present. Sediment
accumulation was very slow, at around 30 years/cm, and consequently the temporal resolution of the
sequence is relatively coarse.
Pollen data are presented in Figure 3. The record is divided into zones (WH1-3) that reflect periods of
similar vegetation composition. The record was analysed using a statistical technique called palynological
richness, which provides a proxy for changes in the diversity and heterogeneity of plant communities
through time (Birks & Line 1992). This curve is presented in Figure 4. Latin names are used in the text for
clarity since not all pollen types can be identified to a single plant species (see Bennett 1994), but a
representative common name is included the first time a pollen type is mentioned in the text.
Results and Interpretation
A rich pastoral resource: species-rich grassland
After the abandonment of the cross-ridge dyke at around A.D. 430, the vegetation on Wether Hill was
dominated by open, herb-rich, dry grassland (Figure 3: zone WH1). Extensive clearance during the late
Iron Age had already removed most trees and shrubs from the Cheviot uplands and much of
Northumberland (Davies & Turner 1979, Macklin et al. 1991, Tipping 1997b). Many of the herbs
recorded on Wether Hill are favoured by disturbance, particularly grazing. This includes Plantago
lanceolata (ribwort plantain), Ranunculus acris-type (buttercups), Rumex acetosa (common sorrel),
Cichorium intybus-type (e.g. dandelions) and Succisa pratensis (devil’s bit scabious), and possibly
Trifolium-type (clovers). Farmers were thus using high quality, species-rich grassland as a pastoral
resource. The ditch contained a damp Cyperaceae (sedge)-Filipendula (e.g. meadowsweet) community.
Cereal cultivation also formed part of the agricultural system, contributing pollen from barley and oats or
wheat (Boyd 1988, van der Veen 1992; the pollen of oats and wheat cannot be separated), as well as
arable weeds such as Papaver rhoeas-type (poppies), Brassicaceae (e.g. shepherd’s purse, kale) and
Chenopodiaceae (e.g. fat hen). Cereal pollen only travels very short distances from fields (Hall 1989), so
the cereal-weed assemblage suggests that some of the land around the hill was being cultivated (cf.
Archaeological Services, University of Durham 1997) (Figure 2).
Pollen diagrams from Northumberland and more widely across Northern England and the Scottish
Cheviots all indicate a long history of agricultural land-use. There are no obvious post-Roman or Anglo-
Saxon field monuments in the Breamish Valley around Wether Hill, but pastoral and arable land-use were
widespread by the seventh century A.D., when both monastic and Anglo-Saxon communities were
4
established in Northumberland. At lower altitude sites to the east, woodland regeneration is recorded
between the fifth and eighth centuries A.D. (Davies & Turner 1979). However, in the Cheviots, both
around Wether Hill and across the Border in the Bowmont Valley (Tipping 1998) (Figure 1), agriculture
continued uninterrupted by political changes during the proto-historic period.
The hills in transition: heather and changing grazing regimes
Calluna vulgaris (heather) was initially only a minor component of the rich pastures on Wether Hill, but a
change in conditions allowed it to invade areas of grassland, forming a mixed grass-heath in zone WH2
(Figure 3). A change in the grazing regime is the inferred cause. Intermediate-level grazing pressures can
favour heather, particularly if this involves summer grazing, when stock preferentially consume grasses,
followed by the removal of the animals from hill-pasture during the winter, to limit grazing of evergreen
heaths (Hunter 1962, Welch 1984, Grant et al. 1987, Hobbs & Gimingham 1987, Milne et al. 1998). This
regime did not affect the range or diversity of herbs present in the pastures around Wether Hill, suggesting
that the new system of land management did not involve high grazing pressures. Higher charcoal values
(Figure 3) may indicate that fire was used to manage the pastures once heather had begun to spread, since
burning prevents heather from becoming too woody, so helping to maintain younger, more nutritious
growth (Hobbs & Gimingham 1987). This combination of burning and grazing maintained a patchy
heather canopy, providing open areas for herbs and grass to flourish and preventing Calluna from
permanently replacing the herb-rich grass community. It is possible that the pastoral regime became more
structured, thus giving rise to and maintaining a grass-heath mosaic. This also raises questions about the
nature of earlier, probably proto-historic grazing systems, which are more difficult to answer from
archaeological and historical evidence.
The chronology for the pollen sequence is poor between the basal radiocarbon date and appearance of
eighteenth to nineteenth century dating indicators, but palynological and documentary evidence from the
Cheviots, Northumberland and Cumbria suggests that the expansion of heather on Wether Hill may have
been a response to widespread changes in grazing practice from the twelfth to thirteenth centuries A.D. In
some instances this may reflect the expansion of monastic-controlled sheep rearing (Barber et al. 1994,
RCAHMS 1994, Tipping 1997a, 1997b, 2000), as occurred in parts of the Forest of Redesdale to the
south, but this cannot be the case in Ingram parish since no monasteries had grazing rights there (Dixon
1985, Vol. I). In addition, cattle remained the most common stock type in this area until the mid- to late
eighteenth century A.D.. However, it is likely that the regulation of summer grazing was established at the
same time as the medieval nuclear villages, which were certainly in place by the twelfth century A.D. at
the latest. Furthermore, Ingram parish possessed extensive rough grazing, which ran into the heart of the
5
Cheviot hills, bordering upon the hunting forests of Chevy Chase to the north-west, Alnham to the south,
and probably Ilderton to the north, since Uhtred the forester of Ilderton was apprehended by the lord of
Hartside, which lay in the west of Ingram parish, in A.D. 1256 (NCH XIV, 471-6). It is not known for
certain if Ingram was within a hunting forest, but being surrounded by them makes it highly likely. In this
context, the regulation of grazing becomes especially sensitive, since the maintenance of the vert or
habitat of the deer was crucial to the successful management of the forest (Gilbert 1983).
Population expansion is thought to have been the driving force behind the spread of agriculture even
further into uplands during the twelfth and thirteenth centuries A.D. in many parts of northern England
and southern Scotland (e.g. Weardale, Co. Durham (Roberts et al. 1973), the forest of Bowland,
Lancashire (Anson & Tallis 1994) and Liddesdale, Roxburghshire (Dixon 1997)). North Northumberland
is no exception (Dixon 1985). Here it pushed into the fringes of the Cheviots massif. Despite the use of
the upper part of the Breamish as a hunting forest by the Lords of Alnwick, settlement had spread to the
cultivable parts of the valley, as high as 380 m OD at Bromley in the neighbouring parish of Alnham by
1300 A.D.. As elsewhere, rough grazing was being progressively taken into cultivation through the
benefits of summer grazing, as happened at Greenside further up the valley; while Hartside township to
the west marks the limits of the medieval expansion in Ingram during the thirteenth century A.D. (Dixon
1985, Vol. II, No. 102). Indeed rig cultivation had visibly spread very close to this part of Wether Hill in
the medieval period (Dixon 1985, Vol. I), leaving only a relatively small area of unimproved rough
grazing in the township (Figure 2). The changes in vegetation cover and inferred grazing management in
zone WH2 may result from these processes of expansion and consolidation.
From the thirteenth to seventeenth centuries A.D., the Anglo-Scottish Borders were affected by successive
outbreaks of war and plague (Dixon 1985, Lomax 1996). Instability caused by Border warfare extended to
the valleys around Wether Hill with the burning of nearby Ingram church in A.D. 1296 (NCH, 1893-1940,
XIV, 465). Agricultural recession is indicated by the extensive remains of abandoned rig and deserted
settlements in the upper parts of Ingram parish (which includes Wether Hill), in neighbouring Alnham
parish, as well as elsewhere in the Cheviots. This is supported by contemporary late medieval
documentary sources (Dixon 1985) and the evidence of excavation at Alnhamsheles (Dixon & Brown
1981). The reduced population and the decline in the amount of land under cultivation had several
consequences that are relevant here. First, there was the creation of enclosed pastures on former arable
land, called ‘ox pastures’ on the Percy estates, where the cattle of the plough team were grazed. This
increased access to good pasture and reduced the traditional reliance upon shieling for summer grazing - a
practice that landowners finally eradicated in the early seventeenth century A.D. in this part of
6
Northumberland, in order to exploit the grazing of the summer pastures for their own cattle and sheep.
Second, some former communal arable was converted to demesne land and was leased to a single farmer,
who maintained it as pasture in upland parishes like Alnham (Dixon 1985).
Through this period, the palynological evidence indicates the continued use of pastures accompanied by
reduced arable production and there are numerous fluctuations in the balance between heather and grass,
as well as changes in the amount of burning (zone WH2). It is unfortunate that the slow rate of sediment
accumulation and relatively poor dating evidence do not allow these vegetation changes to be interpreted
in their appropriate historical context. It is not possible to establish whether they reflect the impact of
medieval expansion, later medieval insecurities and population decline, or the effects of subsequent
changes in land-holding patterns. It could be speculated that a combination of summer pasturing with
expanded arable during the twelfth to thirteenth centuries A.D. are reflected in the continuous record for
cereal-type pollen in the transition from zones WH1-2, preceding the subsequent recession in cultivation
with continued grazing recorded in zone WH2. More closely regulated management of hill pastures
following late medieval agricultural recession may have contributed to the maintenance of grass-heath.
However, it remains problematic that the present pollen record cannot be more closely compared with
other records of agrarian and social change.
At a regional level, palynological evidence for the medieval period onwards is scarce and chronologies are
poor (Young 2004), but there is evidence for both continuity and woodland regeneration across the north
of England around this time. Widespread but patchy woodland regeneration recorded elsewhere in
Northumberland, Cumbria and Weardale during the fourteenth century A.D. has been attributed to war
and plague, especially the decline in population density due to disease (Roberts et al. 1973, Davies &
Turner 1979, Barber et al. 1994). However, more securely dated pollen sequences from the Scottish
Cheviots indicate the continuity of grazing activities through this period (Tipping 1997b, 2000),
comparable with the evidence from Wether Hill. Livestock may have provided a form of ‘moveable
wealth’, which allowed farmers the flexibility to maintain the grazing regime through periods of social
and political upheaval (cf. Topping 1999, Campbell et al. 2002).
Landscape degradation: over-exploitation of the resource?
Changes in peasant land tenure in the seventeenth century A.D. were one stage in the process that led to
the final demise of communal agriculture in Northumberland. As late as A.D. 1604 there were still twelve
small tenant farmers at Ingram village who worked 1600 acres that were still intermixed townfields in
A.D. 1663 (Dixon 1985, Vol. II, 368-9). A survey of A.D. 1734 indicates that the inhabitants of Ingram
7
were poor, but still engaged in mixed farming. By A.D. c.1800 the set-up had changed. There were a few
enclosed rectilinear fields used for cultivation near the farmstead, which had replaced the village, and the
rest of the farmland was converted to pasture (Dixon 1985, Vol. II, No. 125 Ingram) (Figure 2). The
implication is that the clearance of the village was carried out in the mid-to-late eighteenth century A.D..
This appears to be associated with the most significant vegetation changes of the last 1500 years (the
transition to zone WH3, Figure 3), when heather virtually disappeared, grasses dominated the pollen rain
on Wether Hill, and the small amount of remaining tree and shrub growth was lost, resulting in the
formation of a predominantly treeless landscape. Exotic species like Fagus (beech), Juglans (walnut) and
Acer (e.g. sycamore) appear, followed by Picea (spruce) and the expansion of Pinus sylvestris (Scots
pine). These park and plantation species suggest that heather had disappeared by the mid- to late
nineteenth century A.D., an inference supported by the subsequent appearance of soot particles from the
widespread combustion of fossil fuels during the late nineteenth century A.D..
By A.D.c.1900 the landscape had altered radically. The diverse range of herbs that had characterised the
pastures had disappeared (Figure 4), resulting in the formation of the present species-poor dry grassland.
While the disappearance of many species could be interpreted as an absence of grazing pressure (Tipping
2000), the two most common herbs remain ribwort plantain, which survives grazing pressure, and
Potentilla-type, possibly tormentil, which is unpalatable to stock. This suggests that the pollen sequence
reflects intensified grazing pressure. Bedstraw pollen (Rubiaceae) is the only herb pollen type to show
increased representation and grazing experiments have shown that heath bedstraw (Galium saxatile) is
favoured by high grazing pressure from sheep (Welch 1984). This species is common in dry grassland and
may be the source of the Rubiaceae pollen recorded at Wether Hill. This would correlate well with the
evidence for a move from mixed farming to a predominantly sheep grazing regime that was focussed on a
few enclosed fields near the farmstead on the site of Ingram village, which was specifically designed as a
sheep-farm. The combined impact of this new, intensified grazing regime, the cessation of burning and the
senescence of heather (Hunter 1962), are likely to have culminated in the loss of first the heather and then
the herbs from these hills.
The ecological changes recorded on Wether Hill during the past few centuries were not confined to this
part of Northumberland: similar changes have been observed across many upland areas of Britain and
Ireland (Roberts et al. 1973, Stevenson & Thompson 1993, Stevenson & Birks 1995, Chambers et al.
1999, Davies 1999, Stevenson & Rhodes 2000, Tipping 2000, Tipping et al. 2001). The deterioration of
upland pastures is linked with a significant intensification in land-use over the last c.500 years,
8
particularly since the age of Agricultural Improvement (Dodgshon & Olsson 2006). At a national level, no
single cause can be identified (Stevenson & Rhodes 2000), but higher grazing pressures and the
introduction of heavier breeds of sheep to satisfy the demands of expanded markets are significant factors
in the degradation of upland plant communities. This evidence indicates a need to question the origin and
antiquity of current ‘degraded’ heaths, since both the formation of these upland heaths and their
subsequent contraction are a direct result of changes in management; they are not remnants of natural
communities (cf. Chambers et al. 1999).
Conclusions: the legacy of pastoral exploitation
The sequence of vegetation changes recorded at Wether Hill is by no means unique and this adds to
existing evidence that, far from being a natural wilderness, open upland habitats in Britain and Ireland
have undergone significant changes over the last 1500 years. Using historical evidence it is possible to
establish how decisions made within changing social and economic contexts have shaped the landscapes
reflected in palaeoenvironmental sequences. Agriculture, specifically grazing, has been the most important
factor shaping the present landscape in this area of Northumberland. It is also significant at a national
scale because of the widespread use of upland and Highland areas for sheep grazing. As a consequence of
this long exploitation, and especially the more intensive management of the last c.200 years, many upland
landscapes have suffered a decline in biodiversity. Although perhaps less ecologically damaging, the data
also indicate a need to examine more closely the processes and effects of agrarian and social changes
during the twelfth to seventeenth centuries A.D.. This must raise questions about the sustainability of
current land-use practices and the conservation value of the plant communities in these ancient, cultural
landscapes.
The environment is a fundamental resource, yet without understanding and acknowledging the extent to
which it has been influenced by humans, it is difficult to evaluate the possible impacts of climatic change
or establish sustainable land-management and conservation strategies. Combining pollen records with
historical and archaeological sources provides a valuable insight into the relationship between society,
economics and landscape development.
Acknowledgements
The authors would like to thank Paul Frodsham (Northumberland National Parks) for funding the second
radiocarbon date, Tim Gates for permission to use the aerial photograph, and Peter Topping, Richard
Tipping, Erlend Hindmarch and James Hepher for advice and assistance.
9
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12
Captions for figures and plate
Figure 1. Location of Wether Hill within Northumberland, NE England.
Figure 2. Plan of the Iron Age hillfort on Wether Hill, showing the location of the pollen core from the
adjacent cross-ridge dyke, and the extent of the surrounding ridge and furrow cultivation throughout the
township of Ingram.
Figure 3. Pollen data from Wether Hill, with estimated ages for transitions between pollen assemblage
zones. The oldest pollen lies at the base, progressing to recent pollen rain at the top of the diagram. The
clear curve represents a ten times exaggeration to show more clearly the trends in rarer pollen types.
Figure 4. Estimated palynological richness or diversity around Wether Hill, with 95% confidence
intervals, mean diversity and pollen zone boundaries. Note the long-term decline in diversity within zone
WH3 after a period of relative stability.
Plate 1. Aerial photograph of Wether Hill from the south-east, showing the hill fort, the cross-ridge dyke,
the cultivation ridges and present mosaic of grass, bracken and heath. Photo by Tim Gates, copyright
reserved.
13
Figure 2. Plan of the Iron Age hillfort on Wether Hill, showing the location of the pollen core from the
adjacent cross-ridge dyke, and the extent of the surrounding ridge and furrow cultivation throughout the
township of Ingram.
15
5
10
15
20
25
30
35
40
45
50
Depth
below
surfa
ce (c
m)
c. AD 1900
c. AD 1750
c. AD 1100-1400
c. AD 430
Acer c
ampe
stre-t
ype (e
.g. sy
camore
)
Alnus
glutinosa
(alder)
Betul
a (birc
h)
Fagus
(bee
ch)
Frax
inus e
xcelsio
r (ash
)
Juglan
s (walnut)
Picea (
spruc
e)
Pinus
sylve
st ris
(Sco
ts pine)
Quercus
(oak
)
Sorbus
aucu
paria
(rowan)
Ulmus (
elm)
Corylus
avella
na-typ
e (cf. h
azel)
Myrica
gale (b
og myrt
le)
Salix
(willo
w)
Viburn
um opulus
(gueld
er rose
)
20 40 60
Callun
a vulgari
s (he
ather)
Erica
(hea
th)
Vacc
inium
-type
(e.g. b
ilberr
y)
20 40 60 80 100
Poac
eae (w
ild gras
ses)
Avena
/Triticu
m grou
p (oa
ts/whea
t)
Hordeum
group (e
.g. barl
ey)
20
Cyperace
ae (s
edge
s)
Achille
a-type
(e.g. y
arrow)
Agroste
mma gith
ago (c
orn co
ckle)
Apiac
eae (
e.g. co
w parsley
)
Artemisi
a-typ
e (e.g.
mugw
ort)
Aster
aceae
(dais
y/dan
delio
n family
)
Bras
sicace
ae (mus
tard/cabb
age fa
mily)
Campa
nula-ty
pe (h
arebe
lls)
Caryop
hylla
ceae
(pink
/campion
family
)
Centau
rea nigr
a (kn
apweed
)
Cerasti
um-type (
e.g. ch
ickwee
d)
Cheno
podiac
eae (
fat hea
n fam
ily)
Cichori
um in
tybus-t
ype (
e.g. da
ndeli
ons)
Cirsium
-type
(thist
les)
Epilo
bium-ty
pe (w
illowhe
rbs)
Fabac
eae (p
ea/cl
over fa
mily)
20 40
Filipen
dula (e
.g. mea
dowsw
eet)
Helian
themum (ro
ck ro
se)
Heracle
um (h
ogwee
d)
Hyperic
um (St. J
ohn's w
ort)
Lamiace
ae (mint fa
mily)
Lotus (
bird's-
foot tre
foils)
Lych
nis f lo
si-cu
culi (
ragged
robin
)
Lych
nis vi
scaria
-type (
e.g. ca
tchfly
)
Melampy
rum (c
ow-whe
at)
Mentha
-type
(e.g. m
int)
Zone
WH3
WH2
WH1
HerbsHeathsTrees & shrubs
Analyst: A.L.DaviesClear curve = x10 exaggeration
Stratig
raphy
Turf Black peat Brown peat Clay
Figure 3a. Pollen data from Wether Hill, with estimated ages for transitions between pollen assemblage zones. The oldest pollen lies at the base,
progressing to recent pollen rain at the top of the diagram. The clear curve represents a ten times exaggeration to show more clearly the trends in
rarer pollen types.
16
5
10
15
20
25
30
35
40
45
50
Depth
below
surfa
ce (c
m)
c. AD 1900
c. AD 1750
c. AD 1100-1400
c. AD 430
Papa
verac
eae (p
oppy
family)
Pedic
ularis
(louse
wort)
Persi
caria
macu
losa-ty
pe (k
notw
eed)
Planta
go co
ronop
us (b
uck's-
horn plan
tain)
20
Planta
go la
nceola
ta (ri
bwort
plantai
n)
Planta
go m
edia/m
ajor (g
reater
/hoary
plantai
n)
Polyg
ala (m
ilkwort)
Polyg
onum
(kno
tgrass
)
20
Poten
tilla-ty
pe (e
.g. torm
entil)
Ranun
culus
acris-t
ype (
butte
rcups
)
Rosace
ae (ros
e fam
ily)
Rubiace
ae (bed
st raw fa
mily)
Rumex
aceto
sella
(shee
p's so
rrel)
Rumex
aceto
sa-ty
pe (c
ommon sorre
l)
Rumex
obtus
ifoliu
s-typ
e (doc
k)
Rumex
undiff.
(doc
k/sorre
l)
Sang
uisorb
a mino
r (bu
rnet )
Serra
tula-t
ype (
e.g. sa
w-wort)
Silen
e dioi
ca-ty
pe (ca
mpions)
Solid
ago v
irgaure
a-type
(e.g.
daisie
s)
Sperg
ula-typ
e (e.g. s
purry)
Stac
hys s
ylvatic
a-typ
e (e.g.
woun
dwort
)
Stell
aria holo
stea (
stitch
wort)
Succ
isa praten
sis (d
evil's
-bit s
cabious
)
Thalict
rum (mea
dow ru
e)
Trifo
lium-ty
pe (e
.g. clov
ers)
Valer
ianella
(corn
salad
)
Valer
iana of
ficinali
s-typ
e (vale
rian)
Vicia
sylva
tica-t
ype (
vetch
es)
Botry
chium
lunari
a (moo
nwort
)
Cryptogra
mma cris
pa (p
arsley
fern)
Ophioglos
sum (a
dder's
-tongu
e fern
)
Osmun
da re
galis (r
oyal fe
rn)
Polyp
odium
vulgare
(poly
pody
)
Pteri
dium aq
uilinu
m (brack
en)
Ptero
psida (
monole
te) in
det. (fe
rns)
Lyco
podiu
m clava
tum (stag
's-ho
rn cl
ubmoss
)
Spha
gnum
(bog
mos
s)
Equis
etum (h
orsetai
l)
Sparg
anium erectu
m-type (
bur r
eed)
50 100 150
Charco
al su
m
Sphe
roidal c
arbonac
eous p
art icles
20 40 60 80 100
Organic
conte
nt
20 40 60 80 100
Tree
sSh
rubs
Heaths
Herbs
Zone
WH3
WH2
WH1
Ferns, mosses & aquaticsHerbs
Analyst: A.L.DaviesClear curve = x10 exaggeration
Stratig
raphy
Turf Black peat Brown peat Clay
Figure 3b. Pollen data continued.
17
4.0
9.0
14.0
19.0
24.0
29.0
34.0
39.0
44.0
49.0
0 5 10 15 20 25 30 35 40
Estimated pollen diversity
Dep
th b
elow
sur
face
(cm
)
Zone WH3
Zone WH2
Zone WH1
Figure 4. Estimated palynological richness or diversity around Wether Hill, with 95% confidence
intervals, mean diversity and pollen zone boundaries. Note the long-term decline in diversity within zone
WH3 after a period of relative stability.
18