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PLANT SCIENCE 184. Cellular Respiration: Harvesting Chemical Energy. Bacteria are used to produce yogurt, sour cream, pepperoni, and cheese. Both carbon monoxide and cyanide kill by disrupting cellular respiration. All the energy in all the food you eat can be traced back to sunlight. - PowerPoint PPT Presentation
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Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
PowerPoint® Lecture Slides for Essential Biology, Second Edition & Essential Biology with Physiology
Neil Campbell, Jane Reece, and Eric Simon
Presentation prepared by Chris C. Romero
PLANT SCIENCE 184PLANT SCIENCE 184
Cellular Respiration: Harvesting Chemical Energy
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Bacteria are used to produce yogurt, sour cream, pepperoni, and cheese
• Both carbon monoxide and cyanide kill by disrupting cellular respiration
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• All the energy in all the food you eat can be traced back to sunlight
• If you exercise too hard, your muscles shut down from a lack of oxygen
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• When you exercise
BIOLOGY AND SOCIETY: FEELING THE “BURN”
– Muscles need energy in order to perform work
– Your cells use oxygen to release energy from the sugar glucose
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Aerobic metabolism
– When enough oxygen reaches cells to support energy needs
• Anaerobic metabolism
– When the demand for oxygen outstrips the body’s ability to deliver it
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Anaerobic metabolism
– Without enough oxygen, muscle cells break down glucose to produce lactic acid
– Lactic acid is associated with the “burn” associated with heavy exercise
– If too much lactic acid builds up, your muscles give out
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Physical conditioning allows your body to adapt to increased activity
– The body can increase its ability to deliver oxygen to muscles
• Long-distance runners wait until the final sprint to exceed their aerobic capacity
Figure 6.1
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ENERGY FLOW AND CHEMICAL CYCLING IN THE BIOSPHERE
• Fuel molecules in food represent solar energy
– Energy stored in food can be traced back to the sun
• Animals depend on plants to convert solar energy to chemical energy
– This chemical energy is in the form of sugars and other organic molecules
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Photosynthesis
Producers and Consumers
– Light energy from the sun powers a chemical process that makes organic molecules
– This process occurs in the leaves of terrestrial plants
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• Autotrophs
– “Self-feeders”
– Plants and other organisms that make all their own organic matter from inorganic nutrients
• Heterotrophs
– “Other-feeders”
– Humans and other animals that cannot make organic molecules from inorganic ones
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Producers
– Biologists refer to plants and other autotrophs as the producers in an ecosystem
• Consumers
– Heterotrophs are consumers, because they eat plants or other animals
Figure 6.2
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• The ingredients for photosynthesis are carbon dioxide and water
– CO2 is obtained from the air by a plant’s leaves
– H2O is obtained from the damp soil by a plant’s roots
• Chloroplasts rearrange the atoms of these ingredients to produce sugars (glucose) and other organic molecules
– Oxygen gas is a by-product of photosynthesis
Chemical Cycling Between Photosynthesis and Cellular Respiration
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Both plants and animals perform cellular respiration
– Cellular respiration is a chemical process that harvests energy from organic molecules
– Cellular respiration occurs in mitochondria
• The waste products of cellular respiration, CO2 and H2O, are used in photosynthesis
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Figure 6.3
Sunlightenergy
Ecosystem
Photosynthesis(in chloroplasts)
Glucose
Oxygen
Carbon dioxide
Cellular respiration(in mitochondria)
Water
for cellular work
Heat energy
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Cellular respiration
CELLULAR RESPIRATION: AEROBIC HARVEST OF FOOD ENERGY
– The main way that chemical energy is harvested from food and converted to ATP
– This is an aerobic process—it requires oxygen
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Cellular respiration and breathing are closely related
– Cellular respiration requires a cell to exchange gases with its surroundings
– Breathing exchanges these gases between the blood and outside air
The Relationship Between Cellular Respiration and Breathing
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Figure 6.4
Breathing
Lungs
Musclecells
Cellularrespiration
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• A common fuel molecule for cellular respiration is glucose
– This is the overall equation for what happens to glucose during cellular respiration
The Overall Equation for Cellular Respiration
Unnumbered Figure 6.1
Glucose Oxygen Carbondioxide
Water Energy
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• During cellular respiration, hydrogen and its bonding electrons change partners
– Hydrogen and its electrons go from sugar to oxygen, forming water
The Role of Oxygen in Cellular Respiration
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Chemical reactions that transfer electrons from one substance to another are called oxidation-reduction reactions
Redox Reactions
– Redox reactions for short
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• The loss of electrons during a redox reaction is called oxidation
• The acceptance of electrons during a redox reaction is called reduction
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Unnumbered Figure 6.2
[Oxygen gains electrons (and hydrogens)]
Oxidation[Glucose loses electrons (and hydrogens)]
Glucose Oxygen Carbondioxide
Water
Reduction
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
•RS IS NECESSARY IN ALL LIVING CELLS.
•PLANTS ARE WELL KNOWN FOR PS, BUT THEY MUST ALSO REPIRE IN ORDER TO
SURVIVE.
• PS - OCCURS ONLY IN PLANT CELLS CONTAINING CHLOROPHYLL DURING THE DAYLIGHT HOURS.
•RS - OCCURS IN ALL OF A PLANT’S LIVING CELLS 24 -7.
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
PLANTS NEED ENERGY TO PERFORM MANY ESSENTIAL FUNCTIONS OF LIFE:
GROWTH,
REPAIR,
NUTRIENT MOVEMENT,
REPRODUCTION, &
NUTRIENT TRANSPORT.
WHY IS RS NECESSARY?
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Cellular respiration is an example of a metabolic pathway
– A series of chemical reactions in cells –building or degradation process
• All of the reactions involved in cellular respiration can be grouped into three main stages
– Glycolysis
– The Krebs cycle
– Electron transport
– * WHAT IS METABOLISM?
The *Metabolic Pathway of Cellular Respiration
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
A Road Map for Cellular Respiration
Cytosol
Mitochondrion
High-energyelectronscarriedby NADH
High-energyelectrons carriedmainly byNADH
Glycolysis
Glucose2
Pyruvicacid
KrebsCycle
ElectronTransport
Figure 6.7
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Glycolysis
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Glycolysis breaks a six-carbon glucose into two three-carbon molecules
– These molecules then donate high energy electrons to NAD+, forming NADH
• A molecule of glucose is split into two molecules of pyruvic acid
Stage 1: Glycolysis
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Figure 6.8
Glucose
2 Pyruvic acid
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Krebs Cycle
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Stage 2: The Krebs Cycle
• The Krebs cycle completes the breakdown of sugar
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• In the Krebs cycle, pyruvic acid from glycolysis is first “prepped” into a usable form, Acetyl-CoA
Figure 6.10
CoA
1
2
3Pyruvic
acid
Aceticacid
Coenzyme A
Acetyl-CoA(acetyl-coenzyme A)
CO2
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• The Krebs cycle extracts the energy of sugar by breaking the acetic acid molecules all the way down to CO2
– The cycle uses some of this energy to make ATP
– The cycle also forms NADH and FADH2
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Figure 6.11
Input
Acetic acid
ADP
3 NAD
FAD
KrebsCycle
Output
2 CO2
1 2
3
4
5
6
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Electron Transport
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Stage 3: Electron Transport
• Electron transport releases the energy your cells need to make the most of their ATP
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• The molecules of electron transport chains are built into the inner membranes of mitochondria
– The chain functions as a chemical machine that uses energy released by the “fall” of electrons to pump hydrogen ions across the inner mitochondrial membrane
– These ions store potential energy
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Figure 6.12
Proteincomplex
Electroncarrier
Innermitochondrialmembrane
Electronflow
Electron transport chain ATP synthase
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
The Versatility of Cellular Respiration
• Cellular respiration can “burn” other kinds of molecules besides glucose
– Diverse types of carbohydrates
– Fats
– Proteins
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Figure 6.13
Food
Polysaccharides Fats Proteins
Sugars Glycerol Fatty acids Amino acids
Amino groups
Glycolysis Acetyl-CoA
KrebsCycle Electron
Transport
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Adding Up the ATP from Cellular Respiration
Figure 6.14
Cytosol
Mitochondrion
Glycolysis
Glucose2
Pyruvicacid
2Acetyl-
CoA
KrebsCycle Electron
Transport
bydirectsynthesis
by directsynthesis
byATPsynthase
Maximumper
glucose:
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
FERMENTATION: ANAEROBIC HARVEST OF FOOD ENERGY
• Some of your cells can actually work for short periods without oxygen
– For example, muscle cells can produce ATP under anaerobic conditions
• Fermentation
– The anaerobic harvest of food energy
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Human muscle cells can make ATP with and without oxygen
– They have enough ATP to support activities such as quick sprinting for about 5 seconds
– A secondary supply of energy (creatine phosphate) can keep muscle cells going for another 10 seconds
– To keep running, your muscles must generate ATP by the anaerobic process of fermentation
Fermentation in Human Muscle Cells
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Glycolysis is the metabolic pathway that provides ATP during fermentation
– Pyruvic acid is reduced by NADH, producing NAD+, which keeps glycolysis going
– In human muscle cells, lactic acid is a by-product
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Figure 6.15a
2 ADP+ 2
Glycolysis
Glucose
2 NAD
2 Pyruvicacid
+ 2 H
2 NAD
2 Lacticacid
(a) Lactic acid fermentation
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Various types of microorganisms perform fermentation
– Yeast cells carry out a slightly different type of fermentation pathway
– This pathway produces CO2 and ethyl alcohol
Fermentation in Microorganisms
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
Figure 6.15b
2 ADP+ 2
2 ATPGlycolysis
Glucose
2 NAD
2 Pyruvicacid
2 CO2 released
+ 2 H
2 NAD
2 Ethylalcohol
(b) Alcoholic fermentation
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• The food industry uses yeast to produce various food products
Figure 6.16
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• Ancient bacteria probably used glycolysis to make ATP long before oxygen was present in Earth’s atmosphere
EVOLUTION CONNECTION:LIFE ON AN ANAEROBIC EARTH
– Glycolysis is a metabolic heirloom from the earliest cells that continues to function today in the harvest of food energy
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
SUMMARY OF KEY CONCEPTS
• Chemical Cycling Between Photosynthesis and Cellular Respiration
Visual Summary 6.1
Sunlight
Heat
PhotosynthesisCellular
respiration
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• The Overall Equation for Cellular Respiration
Visual Summary 6.2
Oxidation:Glucose loses electrons(and hydrogens)
Glucose Carbon dioxide
Electrons(and hydrogens) Energy
Oxygen
Reduction:Oxygen gainselectrons (andhydrogens)
Copyright © 2004 Pearson Education, Inc. publishing as Benjamin Cummings
• The Metabolic Pathway of Cellular Respiration
Visual Summary 6.3
Glucose Oxygen Water Energy