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What are Ruminants?
Multi-compartment stomach.
Specialized digestive system
capable of digesting
cellulose.
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?
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Cattle
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Sheep
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Goats
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Deer
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Parts and functions of the
ruminant digestive system
Ruminants are
characterized by
having a stomachwith four
compartments
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1. Anus 7. Reticulum
2. Rectum 8. Esophagus3. Cecum 9. Abomasum
4. Colon 10. Omasum
5. Duodenum 11. Small Intestine
6. Rumen
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Parts and functions of the
ruminant digestive systemSmall Intestine
-enzymatic digestion and absorption
-Functions of the small intestine: digestion of
proteins, carbohydrates, and fats; absorption of the
end products of digestion.
1.duodenum
2.jejunum3.ileum
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Mouth
-contains dental pad, teeth, tongue and saliva-saliva contains no salivary amylase
Esophagus
-tube from mouth to stomach-tube from stomach to mouth
Large Intestine
-water absorption-waste storage
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um nan
stomachFour compartments or chambers1) Reticulum
2) Rumen
3) Omasum
4) Abomasum
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Compartment
Capacity Reticulum 5% of capacity
Rumen 80% of capacity
Omasum 7% of capacity
Abomasum 8% of capacity
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Reticulum
http://courses.washington.edu
Honeycomb lining
Smallest compartment
Lies close to the heart
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Functions of
Reticulum Holding area for feed after it passes down esophagus. Provides additional area for fermentation. Contains microorganisms, like rumen
Collection compartment for foreign objects.
Helps open and close rumen. Minimal separation with rumen.
Can contract to a fraction of its resting size.
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Omasu
m Third compartment Globe-shaped
Lining called many plies
Butchers Bible
many piles
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unc ons o
Omasum Reduces feed particle size
Absorbs some water, nutrients and dries out
ingeta .
Absorbs volatile fatty acids
http://www.dpi.nsw.gov.au/agriculture/livestock/goats/mgt/general/rearing
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Abomasum
http://www.cvmbs.colostate.edu
Final Compartment
True Stomach
Glandular Stomach
-Monogastric
Tubular Design
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Secretes enzymes & hydrochloric acid
-Chemical digestion
Dissolves minerals
Breaks down proteins
Passes ingesta to small intestine
Reduces pH to 2.5- Dissolves minerals- Kills rumen bacteria- Breaks down proteins
Functions of
Abomasum
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Rumen Largest of the four components.
On the left side of the animal.
Storage site and fermentation vat.
Houses millions of microorganisms.
Lined with millions of finger-like projections (papillae) thatare needed for absorption.
paunch
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Storage and Soaking
Physical mixing
Breaks down fibrous feeds
Fermentation
Synthesizes:
-Some vitamins-Amino Acids & Proteins
Breaks down fibrous feeds into VFAs
Functions of
Rumen
R i ti ( d
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Rumination (cud
chewing) Rumination- the regurgitation,rechewing and reswallowingof
ingested food
Cud- mass of regurgitated
ingesta; bolus
Process of rumination
1. regurgitate bolus from
rumen
2. rechew and reinsalivate
3. reswallow
4. repeat with another bolus
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Functions of Microorganisms
-digest roughages to make Volatile Fatty Acids
-make protein
-make vitamins K and B complex
(Very similar to cecum of rabbit and horse)
The function of the rumen is to house microorganisms.
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Rumen Environment
rumen contents in mature beef cattleranges from 68 litres to 86 litres withincreasing levels of forage in the diet
rumen microorganisms may account for7 to 14% of the total rumen weight
Ruminal fermentation requires strictlyanaerobic
rumen microorganisms produce volatilefatty acids (i.e., acetate, propionate,butyrate)
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Changes in Rumen
Environment
ruminant's diet is the major influence on
the nature of the rumen environment.
Factors such as composition of the feed,
the degree of physical processing andthe presence of feed additives all affect
the numbers, proportions and digestive
activity of rumen microorganisms.
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occurs during the transition from a forage toa grain based diet
the principal substrates for microbialfermentation change from the components ofplant cell walls (ie. cellulose, hemicellulose,pectin) to cereal starch
The development of a stable microbialpopulation upon transition from a forage to agrain diet is not an immediate process
the numbers of bacteria which produce lacticacid (the acid involved in lactic acidosis)increase with the introduction of grain intothe diet
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the transition from a forage to a grain diet istoo abrupt or if the particle size of the grain istoo small, the microbial population may
become unstable pH of the rumen drops below 5.0 and the
ruminant suffers from lactic acidosis.
the rumen contents may become viscouswith the formation of a stable foam in therumen
The inherently stable nature of rumen
populations makes it extremely difficult toalter the rumen environment through the useof feed additives
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some of these additives may cause short
term changes in the rumen environment,
microbial adaptation often results in therumen environment reverting back to
pretreatment conditions
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FermentationSymbiotic relationship between animal and microorganisms
1) Fermentation of structural
carbohydrates into readily
absorbable and utilizable
sources of energy.
2) Conversion of non-protein
nitrogen into utilizable
protein sources.
3) Synthesis of vitamin K
and B vitamins.
V l til f tt id
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Volatile fatty acids
(VFAs) Main end products of
microbial fermentation.
Provides up to 80% ofthe energy needs of
the animal.
Absorbed by rumen.
Proportion of VFAs
greatly influenced by
diet.
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Volatile fatty acids95% of VFAs
Propionic acid Usually 18 to 20%
Provides energy for
conversion to glucose.
Used in lactose
synthesis.
Some metabolized to
lactic acid.
Favored in a high-
concentrate diet.
Increased by adding
ionophores to diet
Propionic acid
Milk production
Acetic acid
Usually 50 to 60%
Most abundant VFA in
general circulation. Prime metabolic substrate
Used in fat synthesis
Predominates in high
forage diet.
Mammary gland is most
important user of acetate.
Acetic acidMilk fat
Butyric acid
Usually 12 to 18%
Metabolized and
oxidized toketones.
Provides energy to
rumen wall.
Increased in high
grain rations.
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Protein Degradable intake
protein (DIP) Broken down by rumen
microbes into ammonia,amino acids, and
peptides. Microbe growth and
reproduction
Absorbed as ammonia
Undegradable intakeprotein (UIP) Not degraded in rumen
ver e w t or a rupt s t
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ver e w t , or a rupt s tto grain
or other readily-fermented carbohydrate
Lactic acid
accumulation in rumen
(and blood).
Ruminal pH drops.
Drastic shift in microbial
population from normal
rumen microbes to moreacid-tolerant lactobacilli.
can be fatal!
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Eructation reflex
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Eructation reflex -
belching- Quiet, almost inaudible,well-developed reflex
mechanism.
Ruminal contractions
Increased intra-esophageal
pressure
Closure of nasopharyneal
sphincter.
Failure to expel gas can
results in bloat and death.
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Bloat
Gas cant escape
Animal dies from suffocation because of distended
rumen
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Humans, swine, rabbits, chickens and horses all
have a simple stomach, which is also known as a
monogastric digestive system.
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Known as a modifiedmonogastric system
Parts and functions of the monogastric avian
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Mouth/Beak- gather and break down feed
Esophagus- tube from mouth to stomach that is open at th
mouth end
Crop- feed storage and moistening
Proventriculus - glandular stomach (HCI and gastri
juices); enzymatic
Gizzard- muscular stomach; mechanical breakdown
Parts and functions of the monogastric avian
digestive system
P t d f ti f th t i i
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Parts and functions of the monogastric avian
digestive system
Small Intestine- enzymatic digestion and absorption
-Functions of the small intestine: digestion of
proteins, carbohydrates, and fats; absorption of
the end products of digestion
-Enzymes in the small intestine
Ceca- essentially non functioning in monogastrics
P t d f ti f th t i i
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Parts and functions of the monogastric avian
digestive system
Large Intestine
-bacterial activity
-water absorption-waste storage
Cloaca- common chamber for GI and urinary tracts
Vent- common exit for GI and urinary tracts
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Parts and functions of the monogastric mammal digestive
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Parts and functions of the monogastric mammal digestive
system
Stomach-muscular gland lined sac that receives ingesta from the
esophagus and conducts both physical and chemical digestion
-primary secretions:
pepsin- enzyme that digests protein;
hydrochlorides- acids that aid in protein digestion
Small Intestine-enzymatic digestion and absorption
Parts and functions of the monogastric mammal digestive
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-Functions of the small intestine: digestion of proteins, carbohydrates,and fats; absorption of the end products of digestion
1. duodenum- most digestion occurs here
2. jejunum- some digestion and some absorption occur
3. ileum- mostly absorption
-Bile - made in liver, stored in gall bladder, active in the small intestine, emulsifiesfat to aid in digestion
Parts and functions of the monogastric mammal digestive
system
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Parts and functions of the monogastric mammal
digestive system
Enzymes in the small intestine
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Parts and functions of the monogastric mammal
digestive system
Cecum
essentially non functioning in many monogastrics.
Rabbits and horses have an enlarged cecum that acts like a rumen and is
involved with microbial digestion (fermentation)
Large Intestine
-bacterial activity
-water absorption
-waste storage
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Simple stomach, but very large and
complex large intestine.
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Advantage
Soluble carbohydrates, such as glycogen,are available to the animal before they areavailable to the microbes
Disadvantage
Microbial protein is not available to hindgutfermenters
Microbes in the hindgut dont synthezisevitamins, which are also available to theanimal further on in the digestive tract