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    What are Ruminants?

    Multi-compartment stomach.

    Specialized digestive system

    capable of digesting

    cellulose.

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    ?

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    Cattle

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    Sheep

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    Goats

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    Deer

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    Parts and functions of the

    ruminant digestive system

    Ruminants are

    characterized by

    having a stomachwith four

    compartments

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    1. Anus 7. Reticulum

    2. Rectum 8. Esophagus3. Cecum 9. Abomasum

    4. Colon 10. Omasum

    5. Duodenum 11. Small Intestine

    6. Rumen

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    Parts and functions of the

    ruminant digestive systemSmall Intestine

    -enzymatic digestion and absorption

    -Functions of the small intestine: digestion of

    proteins, carbohydrates, and fats; absorption of the

    end products of digestion.

    1.duodenum

    2.jejunum3.ileum

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    Mouth

    -contains dental pad, teeth, tongue and saliva-saliva contains no salivary amylase

    Esophagus

    -tube from mouth to stomach-tube from stomach to mouth

    Large Intestine

    -water absorption-waste storage

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    um nan

    stomachFour compartments or chambers1) Reticulum

    2) Rumen

    3) Omasum

    4) Abomasum

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    Compartment

    Capacity Reticulum 5% of capacity

    Rumen 80% of capacity

    Omasum 7% of capacity

    Abomasum 8% of capacity

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    Reticulum

    http://courses.washington.edu

    Honeycomb lining

    Smallest compartment

    Lies close to the heart

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    Functions of

    Reticulum Holding area for feed after it passes down esophagus. Provides additional area for fermentation. Contains microorganisms, like rumen

    Collection compartment for foreign objects.

    Helps open and close rumen. Minimal separation with rumen.

    Can contract to a fraction of its resting size.

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    Omasu

    m Third compartment Globe-shaped

    Lining called many plies

    Butchers Bible

    many piles

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    unc ons o

    Omasum Reduces feed particle size

    Absorbs some water, nutrients and dries out

    ingeta .

    Absorbs volatile fatty acids

    http://www.dpi.nsw.gov.au/agriculture/livestock/goats/mgt/general/rearing

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    Abomasum

    http://www.cvmbs.colostate.edu

    Final Compartment

    True Stomach

    Glandular Stomach

    -Monogastric

    Tubular Design

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    Secretes enzymes & hydrochloric acid

    -Chemical digestion

    Dissolves minerals

    Breaks down proteins

    Passes ingesta to small intestine

    Reduces pH to 2.5- Dissolves minerals- Kills rumen bacteria- Breaks down proteins

    Functions of

    Abomasum

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    Rumen Largest of the four components.

    On the left side of the animal.

    Storage site and fermentation vat.

    Houses millions of microorganisms.

    Lined with millions of finger-like projections (papillae) thatare needed for absorption.

    paunch

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    Storage and Soaking

    Physical mixing

    Breaks down fibrous feeds

    Fermentation

    Synthesizes:

    -Some vitamins-Amino Acids & Proteins

    Breaks down fibrous feeds into VFAs

    Functions of

    Rumen

    R i ti ( d

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    Rumination (cud

    chewing) Rumination- the regurgitation,rechewing and reswallowingof

    ingested food

    Cud- mass of regurgitated

    ingesta; bolus

    Process of rumination

    1. regurgitate bolus from

    rumen

    2. rechew and reinsalivate

    3. reswallow

    4. repeat with another bolus

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    Functions of Microorganisms

    -digest roughages to make Volatile Fatty Acids

    -make protein

    -make vitamins K and B complex

    (Very similar to cecum of rabbit and horse)

    The function of the rumen is to house microorganisms.

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    Rumen Environment

    rumen contents in mature beef cattleranges from 68 litres to 86 litres withincreasing levels of forage in the diet

    rumen microorganisms may account for7 to 14% of the total rumen weight

    Ruminal fermentation requires strictlyanaerobic

    rumen microorganisms produce volatilefatty acids (i.e., acetate, propionate,butyrate)

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    Changes in Rumen

    Environment

    ruminant's diet is the major influence on

    the nature of the rumen environment.

    Factors such as composition of the feed,

    the degree of physical processing andthe presence of feed additives all affect

    the numbers, proportions and digestive

    activity of rumen microorganisms.

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    occurs during the transition from a forage toa grain based diet

    the principal substrates for microbialfermentation change from the components ofplant cell walls (ie. cellulose, hemicellulose,pectin) to cereal starch

    The development of a stable microbialpopulation upon transition from a forage to agrain diet is not an immediate process

    the numbers of bacteria which produce lacticacid (the acid involved in lactic acidosis)increase with the introduction of grain intothe diet

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    the transition from a forage to a grain diet istoo abrupt or if the particle size of the grain istoo small, the microbial population may

    become unstable pH of the rumen drops below 5.0 and the

    ruminant suffers from lactic acidosis.

    the rumen contents may become viscouswith the formation of a stable foam in therumen

    The inherently stable nature of rumen

    populations makes it extremely difficult toalter the rumen environment through the useof feed additives

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    some of these additives may cause short

    term changes in the rumen environment,

    microbial adaptation often results in therumen environment reverting back to

    pretreatment conditions

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    FermentationSymbiotic relationship between animal and microorganisms

    1) Fermentation of structural

    carbohydrates into readily

    absorbable and utilizable

    sources of energy.

    2) Conversion of non-protein

    nitrogen into utilizable

    protein sources.

    3) Synthesis of vitamin K

    and B vitamins.

    V l til f tt id

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    Volatile fatty acids

    (VFAs) Main end products of

    microbial fermentation.

    Provides up to 80% ofthe energy needs of

    the animal.

    Absorbed by rumen.

    Proportion of VFAs

    greatly influenced by

    diet.

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    Volatile fatty acids95% of VFAs

    Propionic acid Usually 18 to 20%

    Provides energy for

    conversion to glucose.

    Used in lactose

    synthesis.

    Some metabolized to

    lactic acid.

    Favored in a high-

    concentrate diet.

    Increased by adding

    ionophores to diet

    Propionic acid

    Milk production

    Acetic acid

    Usually 50 to 60%

    Most abundant VFA in

    general circulation. Prime metabolic substrate

    Used in fat synthesis

    Predominates in high

    forage diet.

    Mammary gland is most

    important user of acetate.

    Acetic acidMilk fat

    Butyric acid

    Usually 12 to 18%

    Metabolized and

    oxidized toketones.

    Provides energy to

    rumen wall.

    Increased in high

    grain rations.

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    Protein Degradable intake

    protein (DIP) Broken down by rumen

    microbes into ammonia,amino acids, and

    peptides. Microbe growth and

    reproduction

    Absorbed as ammonia

    Undegradable intakeprotein (UIP) Not degraded in rumen

    ver e w t or a rupt s t

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    ver e w t , or a rupt s tto grain

    or other readily-fermented carbohydrate

    Lactic acid

    accumulation in rumen

    (and blood).

    Ruminal pH drops.

    Drastic shift in microbial

    population from normal

    rumen microbes to moreacid-tolerant lactobacilli.

    can be fatal!

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    Eructation reflex

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    Eructation reflex -

    belching- Quiet, almost inaudible,well-developed reflex

    mechanism.

    Ruminal contractions

    Increased intra-esophageal

    pressure

    Closure of nasopharyneal

    sphincter.

    Failure to expel gas can

    results in bloat and death.

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    Bloat

    Gas cant escape

    Animal dies from suffocation because of distended

    rumen

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    Humans, swine, rabbits, chickens and horses all

    have a simple stomach, which is also known as a

    monogastric digestive system.

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    Known as a modifiedmonogastric system

    Parts and functions of the monogastric avian

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    Mouth/Beak- gather and break down feed

    Esophagus- tube from mouth to stomach that is open at th

    mouth end

    Crop- feed storage and moistening

    Proventriculus - glandular stomach (HCI and gastri

    juices); enzymatic

    Gizzard- muscular stomach; mechanical breakdown

    Parts and functions of the monogastric avian

    digestive system

    P t d f ti f th t i i

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    Parts and functions of the monogastric avian

    digestive system

    Small Intestine- enzymatic digestion and absorption

    -Functions of the small intestine: digestion of

    proteins, carbohydrates, and fats; absorption of

    the end products of digestion

    -Enzymes in the small intestine

    Ceca- essentially non functioning in monogastrics

    P t d f ti f th t i i

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    Parts and functions of the monogastric avian

    digestive system

    Large Intestine

    -bacterial activity

    -water absorption-waste storage

    Cloaca- common chamber for GI and urinary tracts

    Vent- common exit for GI and urinary tracts

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    Parts and functions of the monogastric mammal digestive

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    Parts and functions of the monogastric mammal digestive

    system

    Stomach-muscular gland lined sac that receives ingesta from the

    esophagus and conducts both physical and chemical digestion

    -primary secretions:

    pepsin- enzyme that digests protein;

    hydrochlorides- acids that aid in protein digestion

    Small Intestine-enzymatic digestion and absorption

    Parts and functions of the monogastric mammal digestive

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    -Functions of the small intestine: digestion of proteins, carbohydrates,and fats; absorption of the end products of digestion

    1. duodenum- most digestion occurs here

    2. jejunum- some digestion and some absorption occur

    3. ileum- mostly absorption

    -Bile - made in liver, stored in gall bladder, active in the small intestine, emulsifiesfat to aid in digestion

    Parts and functions of the monogastric mammal digestive

    system

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    Parts and functions of the monogastric mammal

    digestive system

    Enzymes in the small intestine

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    Parts and functions of the monogastric mammal

    digestive system

    Cecum

    essentially non functioning in many monogastrics.

    Rabbits and horses have an enlarged cecum that acts like a rumen and is

    involved with microbial digestion (fermentation)

    Large Intestine

    -bacterial activity

    -water absorption

    -waste storage

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    Simple stomach, but very large and

    complex large intestine.

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    Advantage

    Soluble carbohydrates, such as glycogen,are available to the animal before they areavailable to the microbes

    Disadvantage

    Microbial protein is not available to hindgutfermenters

    Microbes in the hindgut dont synthezisevitamins, which are also available to theanimal further on in the digestive tract