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Annales de la Socit entomologique de France(N.S.): International Journal of EntomologyPublication details, including instructions for authors and subscription information:http://www.tandfonline.com/loi/tase20
First Baltic amber megapodagrionid damselfly(Odonata: Zygoptera)Dany Azar a & Andr Nel ba Department of Natural Sciences , Lebanese University , Fanar - Matn - P. O. box26110217 , Lebanonb Entomologie , CNRS UMR 5202, Museum National dHistoire Naturelle , CP 50, 45 rueBuffon, Paris , F-75005 , FrancePublished online: 31 May 2013.
To cite this article: Dany Azar & Andr Nel (2008) First Baltic amber megapodagrionid damselfly (Odonata: Zygoptera),Annales de la Socit entomologique de France (N.S.): International Journal of Entomology, 44:4, 451-457, DOI:10.1080/00379271.2008.10697580
To link to this article: http://dx.doi.org/10.1080/00379271.2008.10697580
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Ann. soc. entomol. Fr. (n.s.), 2008, 44 (4) : 451-457
First Baltic amber megapodagrionid damselfl y (Odonata: Zygoptera)
Abstract. Electropodagrion szwedoi n. gen., n. sp., fi rst Baltic amber megapodagrionid damselfl y, is described. The European and North American fossils document a very high diversity and a much wider distribution of this group of damselfl ies during the Cenozoic than today. A checklist of described fossil species of damselfl ies of the family Megapodagrionidae is given.
Rsum. Le premier Megapodagrionidae de lambre de la Baltique (Odonata : Zygoptera). Electropodagrion szwedoi n. gen., n. sp., premier Megapodagrionidae de lambre balte, est dcrit. Les fossiles europens et nord-amricains dmontrent une trs forte diversit et une distribution beaucoup plus large pour ce groupe pendant le Cnozoque que dans lactuel. Une liste des espces fossiles de Zygoptera: Megapodagrionidae est donne. Keywords: Fossil; Tertiary; Eocene; Baltic Amber.
Dany Azar (1) & Andr Nel (2)*(1) Lebanese University / Faculty of Sciences II / Department of Natural Sciences/ Fanar - Matn - P. O. box 26110217 / Lebanon
(2) CNRS UMR 5202, Musum National dHistoire Naturelle, CP 50, Entomologie, 45 rue Buff on, F-75005, Paris, France * Corresponding author
E-mail: email@example.com, firstname.lastname@example.orgAccept le 23 octobre 2007
The damselfl y taxon Megapodagrionidae sensu Bechly 1996 currently includes 34 extant genera and shows an essentially pantropical distribution (Petruleviius et al. 2008). Relationships among the known living species, are however unclear, and it is unknown whether the fossil species are allied to those that are living today in the pantropical regions. Th e fossil record of the megapodagrionid damselfl ies indicates that this group have a long history extending well beyond the pantropical region.
Th e higher classifi cation in this work follows the phylogenetic system of fossil and extant Odonata of Bechly (1996, 2005). Megapodagrionid damselfl ies are classifi ed as Euzygoptera: Coenagriomorpha. In addition to Megapodagrionidae, the Coenagriomorpha comprises Hypolestidae and Coenagrioniformia in an unresolved phylogenetic relationship; their phylogenetic positions might be subject to change (sedis mutabilis) (Bechly 2005). Included in Megapodagrionidae are Argiolestinae and Megapodagrioninae. Until now, no synapomorphies for all Megapodagrionidae are known, so they could well be a paraphyletic group in the present composition (Bechly 1996, 2005; Rehn 2003).
We describe the fi rst megapodagrionid from the Baltic amber, which also represents the fi rst record of the family in amber.
Material and methodsTh e specimen was examined with an Olympus SZX9 stereomicroscope. Drawings were made using a camera lucida. All measurements were produced using an ocular micrometer; with structures measured as preserved. Photographs were made with an Olympus C5060 digital camera.Th e nomenclature of the dragonfl y wing venation is based on the interpretations of Riek (1976), Riek & Kukalov-Peck (1984), amended by Kukalov-Peck (1991), Nel et al. (1993) and Bechly (1996).
Family Megapodagrionidae Calvert 1913
Genus Electropodagrion n. gen.Type species. Electropodagrion szwedoi n. sp.
Etymology. Named after Elektron, Greek name for amber and Podagrion.Diagnosis. Pterostigma short, covering one cell; pterostigmal brace weak and not oblique; no secondary longitudinal veins between main longitudinal veins except for two weak ones between IR2 and RP3/4 and RP3/4 and MA; CuP exactly opposite base of AA.
Electropodagrion szwedoi n. sp.(Figs 15)
Material. Holotype specimen number 4995, Museum Inkluzji W Bursztynie, Katedra Zoologii Bezkrgocw, Uniwersytet Gdaski, Poland. Etymology. Named after our friend and colleague Dr Jacek Szwedo.Description. Th e thorax with one leg, three wing bases and a distal two third of another wing are preserved. With all the
D. Azar & A. Nel
preserved structures of the wings, we were able to reconstruct partially the forewing and entirely the hindwing (see fi gs 15). Wings nearly similar, hyaline, length 20 mm; width at nodus 3.3 mm; max. width (between nodus and pterostigma) 4.8 mm; distance from base to nodus 7.2 mm (nodus situated at about 36% of wing length); distance from nodus to pterostigma 10.1 mm; distance from base to arculus 4.1 mm; Ax1 and Ax2 well defi ned; Ax1 1.5 mm basal of arculus and Ax2 opposite arculus; no secondary antenodal cross-veins; nine postnodal cross-veins between nodus and pterostigma, rather well aligned with their corresponding postsubnodal cross-veins; pterostigma 1.5 mm long and max. 0.7 mm wide, covering one cell and a half; anterior and posterior margins of pterostigma not distinctly
broadened; basal side of pterostigma not oblique; fi ve cross-veins distal of pterostigma between C and RA; pterostigmal brace vein very slightly oblique; posterior part of arculus (= basal discoidal cross-vein) present, so discoidal cell basally closed, quadrangular, with distal side very oblique, and free of cross-veins, basal margin 0.17 mm long, costal margin 1.04 mm long, posterior margin 1.46 mm, distal margin 0.67 mm long; posterior margin of discoidal cell (= MP+CuA) exactly aligned with MP; origins of RP and MA strongly approximate in arculus; cubital cell free (except for CuP-crossing, 0.5 mm basal of arculus); anal area max. 0.25 mm wide with one row of cells; basal part of AA and AP fused, so that anal area not reaching wing base; CuP exactly opposite base of AA; cubito-
Figure 1 Electropodagrion szwedoi n. gen., n. sp., holotype 4995, photograph of forewing base.
Figure 2 Electropodagrion szwedoi n. gen., n. sp., holotype 4995, photograph of hind wing base.
First Baltic amber megapodagrionid
anal area max. 0.75 mm wide with one row of cells; CuA zigzagged; nodal Cr and subnodus very oblique; nodal Cr branching from ScP 0.2 mm basal of fusion of ScP with costal margin; MP, MA, RP3/4, and IR2 parallel; MP straight, not distally zigzagged, and ending between nodus and pterostigma; postdiscoidal area narrow; MA slightly zigzagged and ending two postnodal cells basal of pterostigma; RP3/4 and MA with one row of cells between them but with area distally widened; RP3/4 ending opposite basal side of pterostigma; base of RP3/4 0.9 mm basal of subnodus; IR2 aligned with subnodus; RP2 originating about three cells and 2.9 mm distal of subnodus; base of IR1 about three cells distally; no lestine oblique vein O between RP2 and IR2; RP2 and IR1, IR1 and RP1, and RP2 and IR2 with only one row of cells between them; RP1 without distinct kink at pterostigmal bra