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Journal of Biomechanics Affiliated with the American Society of Biomechanics, the International Society of Biomechanics, the European Society of Biomechanics, the Japanese Society for Clinical Biomechanics and the Australian and New Zealand Society of Biomechanics. Editor-in-Chief Farshid Guilak Depts. of Orthopaedic Surgery, BME, and MEMS, Duke University Medical Center, Box 3093, Med. Sci. Res. Bldg. RM 375, Durham, NC 27710, USA Editorial Board Associate Editors Frank P.T. Baaijens, Eindhoven University of Technology, Eindhoven, The Netherlands Aurelio Cappozzo, University of Rome “Foro Italico” , Roma, Italy Jonathan Dingwell, University of Texas at Austin, Autsin, TX, USA Marcus Pandy, University of Melbourne, Australia Saeed A. Shirazi-Adl, Ecole Polytechnique, Montreal, Canada Melody A. Swartz, Swiss Institute of Experimental Cancer Research, Lausanne, Switzerland ASB Representatives Richard N. Hinrichs, Arizona State University, Arizona, USA Nicholas Stergiou, University of Nebraska at Omaha, NE, USA ESB Representatives Luca Cristofolini, University of Bologna, Italy Hans Van Oosterwyck, K.U.Leuven, Belgium ISB Representatives David Lloyd, Griffith Health Institute, QLD, Australia Ton van den Bogert, Orchard Kinetics, OH, USA ANZSB Representatives Peter Hunter, University of Auckland, Auckland, New Zealand David Lloyd, University of Western Australia, Australia Independent Members John D. Currey, University of York, UK Walter Herzog, University of Calgary, Alberta, Canada John P. Paul, University of Strathclyde, Glasgow, UK Masaaki Sato, Tohoku University, Japan Nico Westerhof, VU Amsterdam, The Netherlands Author enquiries: For enquiries relating to the submission of articles (including electronic submission) please visit this journal’s homepage at http://www.elsevier.com/locate/biomech. For detailed instructions on the preparation of electronic artwork, please visit http://www.elsevier.com/artworkinstructions. Contact details for questions arising after acceptance of an article, especially those relating to proofs, will be provided by the publisher. You can track accepted articles at http://www.elsevier.com/trackarticle. You can also check our Author FAQs at http://www.elsevier.com/authorFAQ and/or contact Customer Support via http://support.elsevier.com. Journal of Biomechanics has no page charges. Publication Information: Journal of Biomechanics (ISSN 0021-9290). For 2013, volume 46 (16 issues) is scheduled for publication. Subscription prices are available upon request from the Publisher or from the Elsevier Customer Service Department nearest you or from this journal’s website (http://www.elsevier.com/locate/jbiomech). Further information is available on this journal and other Elsevier products through Elsevier’s website (http://www.elsevier.com). Subscriptions are accepted on a prepaid basis only and are entered on a calendar year basis. Issues are sent by standard mail (surface within Europe, air delivery outside Europe). Priority rates are available upon request. Claims for missing issues should be made within six months of the date of dispatch. Advertising information. Advertising orders and enquiries can be sent to: USA, Canada and South America: Elsevier Inc., 360 Park Avenue South, New York, NY 10010-1710, USA; phone: (+1) (212) 633 3974; Europe and ROW: Nathan Wright, Business Development Executive, Commercial Sales Department, Elsevier Ltd, The Boulevard, Langford Lane, Kidlington, Oxford OX5 1GB, UK; phone: (+44) 1865 843652; fax: (+44) 1865 843976; e-mail: [email protected] Orders, claims, and journal enquiries: Please contact the Elsevier Customer Service Department nearest you: St. Louis: Elsevier Customer Service Department, 3251 Riverport Lane, Maryland Heights, MO 63043, USA; phone: (800) 6542452 [toll free within the USA]; (+1) (314) 4478871 [outside the USA]; fax: (+1) (314) 4478029; e-mail: [email protected] Oxford: Elsevier Customer Service Department, The Boulevard, Langford Lane, Kidlington, Oxford, OX5 1GB, UK; phone: (+44) (1865) 843434; fax: (+44) (1865) 843970; e-mail: [email protected] Tokyo: Elsevier Customer Service Department, 4F Higashi-Azabu, 1-Chome Bldg, 1-9-15 Higashi-Azabu, Minato-ku, Tokyo 106-0044, Japan; phone: (+81) (3) 5561 5037; fax: (+81) (3) 5561 5047; e-mail: [email protected] Singapore: Elsevier Customer Service Department, 3 Killiney Road, #08-01 Winsland House I, Singapore 239519; phone: (+65) 63490222; fax: (+65) 67331510; e-mail: [email protected] USA mailing notice: Journal of Biomechanics (ISSN 0021-9290) is published sixteen issues per year by Elsevier Ltd. (The Boulevard, Langford Lane, Kidlington, Oxford OX5 1GB, UK). Periodicals postage paid at Jamaica, NY 11431 and additional mailing offices. USA POSTMASTER: Send change of address to Journal of Biomechanics, Elsevier Customer Service Department, 3251 Riverport Lane, Maryland Heights, MO 63043, USA. AIRFREIGHT AND MAILING in USA by Air Business Ltd., c/o Worldnet Shipping Inc., 156-15, 146th Avenue, 2nd Floor, Jamaica, NY 11434, USA. Sponsored supplements and/or commercial reprints: For more information please contact Elsevier Life Sciences Commercial Sales, Radarweg 29, 1043 NX Amsterdam, The Netherlands; phone: (+31) (20) 4852939/2059; e-mail: [email protected] Editorial Advisory Board Kai-Nan An, USA Thomas P. Andriacchi, USA Thomas D. Brown, USA David B. Burr, USA David Butler, USA Lutz Claes, Germany Stephen C. Cowin, USA Cyril B. Frank, Canada Vijay K. Goel, USA Steven A. Goldstein, USA Roger C. Haut, USA Maury Hull, USA Paavo V. Komi, Finland Van C. Mow, USA Nikos Stergiopolous, Switzerland Ray Vito, USA Peter S. Walker, USA Savio L. Woo, USA Peter Zioupos, UK Consulting Editors Taiji Adachi, Japan Adamantios Arampatzis, Germany Richard M. Aspden, UK Gerard Ateshian, USA Joan E. Bechtold, USA Maarten F. Bobbert, The Netherlands Thomas Buchanan, USA Peter Butler, USA Krishnan B Chandran, USA Jacek Cholewicki, USA Joseph J. Crisco III, USA Tim David, New Zealand Scott Delp, USA Paul DeVita, USA Jonathan Dingwell, USA Manuel Doblare, Spain Dawn Elliott, USA David P. Fyhrie, USA Denis Gagnon, Canada Amit Gefen, Israel Mark D. Grabiner, USA Mansoor Haider, USA Tammy Haut-Donahue, USA Jeffrey Holmes, USA Jay D. Humphrey, USA James C. Iatridis, USA Keita Ito, Netherlands Jukka Jurvelin, Finland Ghassan Kassab, USA Joyce H. Keyak, USA Melissa L. Knothe Tate, USA Sean S. Kohles, USA Alberto Leardini, Italy Amy Lerner, USA Marc Levenston, USA Shu Liu, USA Quan Long, UK Karol Miller, Australia Mohammad K. Mofrad, USA Glen Niebur, USA Maury Nussbaum, USA Fergal O’Brien, Ireland Michael R. Pierrynowski, Canada Domique Pioletti, Switzerland Patrick J. Prendergast, Ireland Antonius Rohlmann, Germany Michael S. Sacks, USA Joan E. Sanders, USA Lori A. Setton, USA Louis J. Soslowsky, USA David Steinman, Canada Ian A. F. Stokes, USA Eiji Tanaka, Japan Scott Tashman, USA Huub M. Toussaint, Netherlands Frans van de Vosse, Netherlands Antonie Van den Bogert, USA Jos Vander Sloten, Belgium Pascal Verdonck, Belgium Marco Viceconti, Italy William R. Walsh, Australia James H.C. Wang, USA Takami Yamaguchi, Japan Philippe K. Zysset, Switzerland

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Journal of BiomechanicsAffiliated with the American Society of Biomechanics, the International Society of Biomechanics, the European Society of Biomechanics, the Japanese Society for Clinical Biomechanics and the Australian and New Zealand Society of Biomechanics.

Editor-in-Chief

Farshid GuilakDepts. of Orthopaedic Surgery, BME, and MEMS, Duke University Medical Center, Box 3093, Med. Sci. Res. Bldg. RM 375, Durham, NC 27710, USA

Editorial Board

Associate EditorsFrank P.T. Baaijens, Eindhoven University of Technology, Eindhoven, The NetherlandsAurelio Cappozzo, University of Rome “Foro Italico”, Roma, ItalyJonathan Dingwell, University of Texas at Austin, Autsin, TX, USAMarcus Pandy, University of Melbourne, Australia Saeed A. Shirazi-Adl, Ecole Polytechnique, Montreal, CanadaMelody A. Swartz, Swiss Institute of Experimental Cancer Research, Lausanne, SwitzerlandASB RepresentativesRichard N. Hinrichs, Arizona State University, Arizona, USANicholas Stergiou, University of Nebraska at Omaha, NE, USAESB RepresentativesLuca Cristofolini, University of Bologna, ItalyHans Van Oosterwyck, K.U.Leuven, Belgium

ISB RepresentativesDavid Lloyd, Griffi th Health Institute, QLD, AustraliaTon van den Bogert, Orchard Kinetics, OH, USAANZSB RepresentativesPeter Hunter, University of Auckland, Auckland, New ZealandDavid Lloyd, University of Western Australia, AustraliaIndependent MembersJohn D. Currey, University of York, UKWalter Herzog, University of Calgary, Alberta, CanadaJohn P. Paul, University of Strathclyde, Glasgow, UKMasaaki Sato, Tohoku University, JapanNico Westerhof, VU Amsterdam, The Netherlands

Author enquiries:For enquiries relating to the submission of articles (including electronic submission) please visit this journal’s homepage at http://www.elsevier.com/locate/biomech. For detailed instructions on the preparation of electronic artwork, please visit http://www.elsevier.com/artworkinstructions. Contact details for questions arising after acceptance of an article, especially those relating to proofs, will be provided by the publisher. You can track accepted articles at http://www.elsevier.com/trackarticle. You can also check our Author FAQs at http://www.elsevier.com/authorFAQ and/or contact Customer Support via http://support.elsevier.com.Journal of Biomechanics has no page charges.

Publication Information:Journal of Biomechanics (ISSN 0021-9290). For 2013, volume 46 (16 issues) is scheduled for publication. Subscription prices are available upon request from the Publisher or from the Elsevier Customer Service Department nearest you or from this journal’s website (http://www.elsevier.com/locate/jbiomech). Further information is available on this journal and other Elsevier products through Elsevier’s website (http://www.elsevier.com). Subscriptions are accepted on a prepaid basis only and are entered on a calendar year basis. Issues are sent by standard mail (surface within Europe, air delivery outside Europe). Priority rates are available upon request. Claims for missing issues should be made within six months of the date of dispatch.

Advertising information. Advertising orders and enquiries can be sent to: USA, Canada and South America: Elsevier Inc., 360 Park Avenue South, New York, NY 10010-1710, USA; phone: (+1) (212) 633 3974; Europe and ROW: Nathan Wright, Business Development Executive, Commercial Sales Department, Elsevier Ltd, The Boulevard, Langford Lane, Kidlington, Oxford OX5 1GB, UK; phone: (+44) 1865 843652; fax: (+44) 1865 843976; e-mail: [email protected]

Orders, claims, and journal enquiries: Please contact the Elsevier Customer Service Department nearest you: St. Louis: Elsevier Customer Service Department, 3251 Riverport Lane, Maryland Heights, MO 63043, USA; phone: (800) 6542452 [toll free within the USA]; (+1) (314) 4478871 [outside the USA]; fax: (+1) (314) 4478029; e-mail: [email protected]: Elsevier Customer Service Department, The Boulevard, Langford Lane, Kidlington, Oxford, OX5 1GB, UK; phone: (+44) (1865) 843434; fax: (+44) (1865) 843970; e-mail: [email protected]: Elsevier Customer Service Department, 4F Higashi-Azabu, 1-Chome Bldg, 1-9-15 Higashi-Azabu, Minato-ku, Tokyo 106-0044, Japan; phone: (+81) (3) 5561 5037; fax: (+81) (3) 5561 5047; e-mail: [email protected]: Elsevier Customer Service Department, 3 Killiney Road, #08-01 Winsland House I, Singapore 239519; phone: (+65) 63490222; fax: (+65) 67331510; e-mail: [email protected]

USA mailing notice: Journal of Biomechanics (ISSN 0021-9290) is published sixteen issues per year by Elsevier Ltd. (The Boulevard, Langford Lane, Kidlington, Oxford OX5 1GB, UK). Periodicals postage paid at Jamaica, NY 11431 and additional mailing offi ces.

USA POSTMASTER: Send change of address to Journal of Biomechanics, Elsevier Customer Service Department, 3251 Riverport Lane, Maryland Heights, MO 63043, USA.AIRFREIGHT AND MAILING in USA by Air Business Ltd., c/o Worldnet Shipping Inc., 156-15, 146th Avenue, 2nd Floor, Jamaica, NY 11434, USA.Sponsored supplements and/or commercial reprints: For more information please contact Elsevier Life Sciences Commercial Sales, Radarweg 29, 1043 NX Amsterdam, The Netherlands; phone: (+31) (20) 4852939/2059; e-mail: [email protected]

Editorial Advisory Board

Kai-Nan An, USAThomas P. Andriacchi, USA Thomas D. Brown, USA David B. Burr, USA David Butler, USALutz Claes, Germany

Stephen C. Cowin, USA Cyril B. Frank, Canada Vijay K. Goel, USA Steven A. Goldstein, USA Roger C. Haut, USAMaury Hull, USAPaavo V. Komi, Finland

Van C. Mow, USA Nikos Stergiopolous, Switzerland Ray Vito, USAPeter S. Walker, USA Savio L. Woo, USAPeter Zioupos, UK

Consulting Editors

Taiji Adachi, JapanAdamantios Arampatzis, GermanyRichard M. Aspden, UK Gerard Ateshian, USAJoan E. Bechtold, USA Maarten F. Bobbert, The Netherlands Thomas Buchanan, USA Peter Butler, USAKrishnan B Chandran, USAJacek Cholewicki, USA Joseph J. Crisco III, USA Tim David, New Zealand Scott Delp, USA Paul DeVita, USAJonathan Dingwell, USAManuel Doblare, Spain Dawn Elliott, USA David P. Fyhrie, USA Denis Gagnon, CanadaAmit Gefen, Israel

Mark D. Grabiner, USA Mansoor Haider, USATammy Haut-Donahue, USAJeffrey Holmes, USAJay D. Humphrey, USA James C. Iatridis, USA Keita Ito, Netherlands Jukka Jurvelin, Finland Ghassan Kassab, USAJoyce H. Keyak, USA Melissa L. Knothe Tate, USA Sean S. Kohles, USA Alberto Leardini, ItalyAmy Lerner, USAMarc Levenston, USAShu Liu, USAQuan Long, UK Karol Miller, Australia Mohammad K. Mofrad, USAGlen Niebur, USA Maury Nussbaum, USA

Fergal O’Brien, IrelandMichael R. Pierrynowski, CanadaDomique Pioletti, SwitzerlandPatrick J. Prendergast, IrelandAntonius Rohlmann, GermanyMichael S. Sacks, USA Joan E. Sanders, USA Lori A. Setton, USA Louis J. Soslowsky, USA David Steinman, CanadaIan A. F. Stokes, USA Eiji Tanaka, JapanScott Tashman, USA Huub M. Toussaint, Netherlands Frans van de Vosse, NetherlandsAntonie Van den Bogert, USAJos Vander Sloten, Belgium Pascal Verdonck, BelgiumMarco Viceconti, Italy William R. Walsh, Australia James H.C. Wang, USA Takami Yamaguchi, JapanPhilippe K. Zysset, Switzerland