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Publication information: Biochimie (ISSN 0300-9084). For 2014, volumes 96 – 107 is scheduled for publication. Subscription prices are available upon request from the Publisher or from the Regional Sales Office nearest you or from this journal’s website (http://www.elsevier.com/ locate/biochi). Further information is available on this journal and other Elsevier products through Elsevier’s website: (http://www.elsevier.com). Subscriptions are accepted on a prepaid basis only and are entered on a calendar year basis. Issues are sent by standard mail (surface within Europe, air delivery outside Europe). Priority rates are available upon request. Claims for missing issues should be made within six months of the date of disptach. Orders, claims, and journal enquiries: please contact the Customer Service Department nearest you: St. Louis: Elsevier Customer Service Department, 3251 Riverport Lane, Maryland Heights, MO 63043, USA; phone: (877) 8397126 [toll free within the USA]; (+1) (314) 4478878 [outside the USA]; fax: (+1) (314) 4478077; e-mail: [email protected] Oxford: Elsevier Customer Service Department, The Boulevard, Langford Lane, Kidlington, Oxford OX5 1GB, UK; phone: (+44) (1865) 843434; fax: (+44) (1865) 843970; e-mail: [email protected] Tokyo: Elsevier Customer Service Department, 4F Higashi-Azabu, 1-Chome Bldg, 1-9-15 Higashi-Azabu, Minato-ku, Tokyo 106-0044, Japan; phone: (+81) (3) 5561 5037; fax: (+81) (3) 5561 5047; e-mail: [email protected] Singapore: Elsevier Customer Service Department, 3 Killiney Road, #08-01 Winsland House I, Singapore 239519; phone: (+65) 63490222; fax: (+65) 67331510; e-mail: [email protected] Author enquiries For enquiries relating to the submission of articles (including electronic submission) please visit this journal’s homepage at http://www.elsevier.com/ locate/biochi. For detailed instructions on the preparation of electronic artwork, please visit http://www.elsevier.com/artworkinstructions. Contact de- tails for questions arising after acceptance of an article, especially those relating to proofs, will be provided by the publisher. You can track accepted articles at http://www.elsevier.com/trackarticle. You can also check our Author FAQs at http://www.elsevier.com/authorFAQ and/or contact Customer Support via http://support.elsevier.com. Aims and Scope This multitopical journal publishes original work as well as review articles and mini-reviews in all areas of biology (enzymology, genetics, immunology, microbiology, structure of macromolecules, etc) provided that the approach adopted stems from biochemistry, biophysics or molecular biology. Editorial Board: Chief Editor: Richard H. Buckingham (Paris, France) Editors: Olga A. Dontsova (Moscow, Russia), Claude Forest (Paris, France), Jeffrey M. Gimble (New Orleans, LA, USA), Jean-Louis Mergny (Pessac, France), Yoshikazu Nakamura (Tokyo, Japan) Associate Editors: Nada A. Abumrad (St. Louis, MO, USA), Paola B. Arimondo (Paris, France), Giuseppe Baldacci (Paris, France), Michel Caboche (Versailles, France), Jonathan B. Chaires (Louisville, KY, USA), Alain J. Cozzone (Lyon, France), Antoine Danchin (Paris, France), Muriel Delepierre (Paris, France), Jay W. Fox (Charlottesville, VA, USA), Jacques Haiech (Illkirch, France), Albert Jeltsch (Stuttgart, Germany), Just Justesen (Aarhus, Denmark), Jack F. Kirsch (Berkeley, CA, USA), Leif A. Kirsebom (Uppsala, Sweden), Andrei A. Korostelev (Worcester, MA, USA), Guido Kroemer (Villejuif, France), Anne-Marie Lambeir (Antwerp, Belgium), Wouter H. Lamers (Amsterdam, Netherlands), Michel Lazdunski (Valbonne, France), David Leys (Manchester, UK), Bruno Lomonte (San José, Costa Rica), Sergio Papa (Bari, Italy), Athanasios G. Papavassiliou (Athens, Greece), Anh Tuân Phan (Singapore), Pierre Plateau (Palaiseau, France), Catherine Postic (Paris, France), Pascale Romby (Strasbourg, France), Alain Sarasin (Villejuif, France), Jiří Šponer (Brno, Czech Republic), Jean-François Savouret (Paris, France), Eric Westhof (Strasbourg, France), Huan-Ming Yang (Beijing, China) Published under the auspices of: SOCIÉTÉ FRANÇAISE DE BIOCHIMIE ET BIOLOGIE MOLÉCULAIRE Site web : www.sfbbm.fr Siège Social : 4, avenue de l’Observatoire, 75006 Paris Adresse Postale : 45, rue des Saints-Pères, 75270 Paris cedex 06 Président : Frédéric Dardel - Secrétaire Général : Alain Krol - Secrétaire aux Relations Internationales : Jacques-Henri Weil - Secrétaire des Réunions Scientifiques : Guy Branlant - Trésorier : Philippe Fossé - Regard sur la Biochimie : Alain Krol - Responsable du site internet : Christophe Geourjon - Relations avec l’Industrie : Danielle Lando © 2014 Société française de biochimie et biologie moléculaire / Elsevier Masson SAS Printed by Henry Ling Ltd., The Dorset Press, Dorchester, UK Directeur de la publication : Daniel Rodriguez Dépôt légal : 409242k - À parution Périodicité : 12 n os / an

Editorial Board

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Publication information: Biochimie (ISSN 0300-9084). For 2014, volumes 96 – 107 is scheduled for publication. Subscription prices are available upon request from the Publisher or from the Regional Sales Office nearest you or from this journal’s website (http://www.elsevier.com/ locate/biochi). Further information is available on this journal and other Elsevier products through Elsevier’s website: (http://www.elsevier.com). Subscriptions are accepted on a prepaid basis only and are entered on a calendar year basis. Issues are sent by standard mail (surface within Europe, air delivery outside Europe). Priority rates are available upon request. Claims for missing issues should be made within six months of the date of disptach.

Orders, claims, and journal enquiries: please contact the Customer Service Department nearest you:St. Louis: Elsevier Customer Service Department, 3251 Riverport Lane, Maryland Heights, MO 63043, USA; phone: (877) 8397126 [toll free within the USA]; (+1) (314) 4478878 [outside the USA]; fax: (+1) (314) 4478077; e-mail: [email protected]: Elsevier Customer Service Department, The Boulevard, Langford Lane, Kidlington, Oxford OX5 1GB, UK; phone: (+44) (1865) 843434; fax: (+44) (1865) 843970; e-mail: [email protected]: Elsevier Customer Service Department, 4F Higashi-Azabu, 1-Chome Bldg, 1-9-15 Higashi-Azabu, Minato-ku, Tokyo 106-0044, Japan; phone: (+81) (3) 5561 5037; fax: (+81) (3) 5561 5047; e-mail: [email protected]: Elsevier Customer Service Department, 3 Killiney Road, #08-01 Winsland House I, Singapore 239519; phone: (+65) 63490222; fax: (+65) 67331510; e-mail: [email protected]

Author enquiriesFor enquiries relating to the submission of articles (including electronic submission) please visit this journal’s homepage at http://www.elsevier.com/locate/biochi. For detailed instructions on the preparation of electronic artwork, please visit http://www.elsevier.com/artworkinstructions. Contact de-tails for questions arising after acceptance of an article, especially those relating to proofs, will be provided by the publisher. You can track accepted articles at http://www.elsevier.com/trackarticle. You can also check our Author FAQs at http://www.elsevier.com/authorFAQ and/or contact Customer Support via http://support.elsevier.com.

Aims and Scope This multitopical journal publishes original work as well as review articles and mini-reviews in all areas of biology (enzymology, genetics, immunology, microbiology, structure of macromolecules, etc) provided that the approach adopted stems from biochemistry, biophysics or molecular biology.

Editorial Board:

Chief Editor:Richard H. Buckingham (Paris, France)

Editors:Olga A. Dontsova (Moscow, Russia), Claude Forest (Paris, France), Jeffrey M. Gimble (New Orleans, LA, USA), Jean-Louis Mergny (Pessac, France), Yoshikazu Nakamura (Tokyo, Japan)

Associate Editors:Nada A. Abumrad (St. Louis, MO, USA), Paola B. Arimondo (Paris, France), Giuseppe Baldacci (Paris, France), Michel Caboche (Versailles, France), Jonathan B. Chaires (Louisville, KY, USA), Alain J. Cozzone (Lyon, France), Antoine Danchin (Paris, France), Muriel Delepierre (Paris, France), Jay W. Fox (Charlottesville, VA, USA), Jacques Haiech (Illkirch, France), Albert Jeltsch (Stuttgart, Germany), Just Justesen (Aarhus, Denmark), Jack F. Kirsch (Berkeley, CA, USA), Leif A. Kirsebom (Uppsala, Sweden), Andrei A. Korostelev (Worcester, MA, USA), Guido Kroemer (Villejuif, France), Anne-Marie Lambeir (Antwerp, Belgium), Wouter H. Lamers (Amsterdam, Netherlands), Michel Lazdunski (Valbonne, France), David Leys (Manchester, UK), Bruno Lomonte (San José, Costa Rica), Sergio Papa (Bari, Italy), Athanasios G. Papavassiliou (Athens, Greece), Anh Tuân Phan (Singapore), Pierre Plateau (Palaiseau, France), Catherine Postic (Paris, France), Pascale Romby (Strasbourg, France), Alain Sarasin (Villejuif, France), Jiří Šponer (Brno, Czech Republic), Jean-François Savouret (Paris, France), Eric Westhof (Strasbourg, France), Huan-Ming Yang (Beijing, China)

Published under the auspices of:SOCIÉTÉ FRANÇAISE DE BIOCHIMIE ET BIOLOGIE MOLÉCULAIRESite web : www.sfbbm.frSiège Social : 4, avenue de l’Observatoire, 75006 ParisAdresse Postale : 45, rue des Saints-Pères, 75270 Paris cedex 06Président : Frédéric Dardel - Secrétaire Général : Alain Krol - Secrétaire aux Relations Internationales : Jacques-Henri Weil - Secrétaire des Réunions Scientifiques : Guy Branlant - Trésorier : Philippe Fossé - Regard sur la Biochimie : Alain Krol - Responsable du site internet :Christophe Geourjon - Relations avec l’Industrie : Danielle Lando

© 2014 Société française de biochimie et biologie moléculaire / Elsevier Masson SAS

Printed by Henry Ling Ltd., The Dorset Press, Dorchester, UK Directeur de la publication : Daniel RodriguezDépôt légal : 409242k - À parution Périodicité : 12 nos / an