Estuarine, Coastal and Shelf Science Editors N. C. Flemming Institute of Oceanographic Sciences, Wormley, Godalming, Surrey, U.K.

E. Naylor School of Animal Biology, University College of North Wales, Bangor, Gwynedd, North Wales,

U.K.

M. Grant Gross Box 1497, Baltimore, Maryland 21203, U.S.A.

Book Review Editor J. B. Wilson Institute of Oceanographic Sciences, Wormley, Godalming, Surrey, U.K.

S. R. Aston IAEA, Monaco.

R. J. H. Beverton UWIST, Cardiff, U.K.

A. J. Bowen Dalhousie University, Canada.

J. S. Bunt Australian Institute of Marine

Science, Australia.

J. D. Burton Southampton University, U.K.

L. Cabioch Centre National de la Recherche

Scientifique, France.

R. B. Clark University of Newcastle, U.K.

L. E. Cronin University of Maryland, U.S.A.

L. Draper Institute of Oceanographic

Sciences, U.K.

K. Dyer Institute of Oceanographic Sciences,

U.K.

K. 0. Emery Woods Hole Oceanographic

Institution, U.S.A.

B. W. Flemming University of Capetown,

South Africa.

R. S. Glover IMER, Plymouth, U.K.

J. S. Gray Universitetet i Oslo, Norway.

L. W. Haas College of William and Mary,

U.S.A.

D. V. Hansen Atlantic Oceanographic and

Meteorological Laboratory, Miami, U.S.A.

R. G. Hartnoll University of Liverpool, U.K.

N. S. Heaps Institute of Oceanographic

Sciences, Bidston, U.K.

J. B. Jillett University of Otago, New Zealand.

C. Kidson University College of Wales, U.K.

0. Kinne Biologische Anstalt Helgoland,

Germany.

R. B. Krone University of California (Davis),

U.S.A.

P. Lasserre Station Biologique, Roscoff,

France.

C. Little University of Bristol, U.K.

A. McLachlan University of Port Elizabeth,

South Africa.

S.W.Nixon UniversityofRhode Island, U.S.A.

C. B. Officer Dartmouth College, U.S.A.

J. Onoszko Instytut of Hydro-Engineering,

Poland.

S. Z. Qasim Department of Ocean Develop-

ment, New Delhi, India.

M. Rattray, Jr. University of Washington,

U.S.A.

G. Riley Dalhousie University, Canada.

J. P. Riley University of Liverpool, U.K.

M. Schwartz Western Washington Univer-

sity, U.S.A.

J. H. Simpson University College of North

Wales, U.K.

A. D. Short University of Sydney, Australia.

J. H. Steele Woods Hole Oceanographic

Institution, U.S.A.

R. L. Wiegel University of California

(Berkeley), U.S.A.

J. D. Woods University of Kiel, Germany.

Published monthly [January-December) at 24-28 Oval Road, London NW1 7DX, England by Academic Press Inc. (London) Limited. 1985: Volumes 2&21 (12 issues). Inland Ll27.00 inclusive of postage and packing, abroad $315.00 inclusive of postage and packing. Subscription orders should be sent to Academic Press Inc. (London) Limited, High Street, Foots Gay, Sidcup, Kent DA14 5HI’, with the exception of those originating in the U.S.A., Canada and Central and South America which should be sent to Academic Press Inc., 111 Fifth Avenue, New York, N.Y. 10003, U.S.A. Send notices of change of address to the office of the Publishers at least 6-8 weeks in advance. Please include both old and new addresses. U.S.A. Postmaster, send changes of address to Esruarine, Coastal and Shelf Science, Academic Press Inc., 111 Fifth Avenue, New York, N.Y. 10003, U.S.A. Second-class postage paid at Jamaica, N.Y. 11431, U.S.A. Air freight and mailing in the U.S.A. by Publications Expediting Inc., 200 Meacham Avenue, Elmont, N.Y. 10003, U.S.A. Copyright 0 1985 Academic Press Inc. (London) Limited. The appearance of the code at the bottom of the first page in this journal indicates the copyright owner’s consent that copies of the paper may be made for personal or internal use, or for the personal or internal use of specific clients in the U.S.A. This consent is given on the condition, within the U.S.A., that the copier pay the stated per-copy fee through the Copyright Clearance Center, Inc., 21 Congress Street, Salem, MA 01970, for copying beyond that permitted by Sections 107 or 108 of the U.S. Copyright Law. This consent does not extend to other kinds of copying, such as copying for general distribution, for advertising or promotional purposes, for creating new collective works, for resale or for copying or distributing copies outside the U.S.A.