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Editor in Chief Raoul Poupon (Paris) Associate Editors Lionel Arrivé (Paris), Charles Balabaud (Bordeaux), François Ballet (Paris), Yvon Calmus (Paris), Massimo Conio (Genoa), Dominique Debray (Paris), Christèle Desbois-Mouthon (Paris), Christian Florent (Paris), Chantal Housset (Paris), Mehdi Karoui (Paris), Patrick Marcellin (Clichy), Vlad Ratziu (Paris), Marika Rudler (Paris), Lawrence Serfaty (Paris), Julien Taieb (Paris), Christian Trépo (Lyon) Consultant Editors Paulette Bioulac-Sage (Bordeaux), Christian Bréchot (Paris), Fabrice Carrat (Paris), Daniel Dhumeaux (Paris), Serge Erlinger (Aix-en-Provence), Jean-Michel Pawlotsky (Créteil), Thierry Poynard (Paris), Kristina Schoonjans (Lausanne) Editorial assistant Yves Chrétien (Paris) Copyediting Dale Roche-Lebrec (Paris), Gerald Pope (Rennes), Sharyn Wong (Little Rock) Editorial Board Domenico Alvaro (Rome), Tony Andréani (Paris), Mario Angelico (Rome), Vicente Arroyo (Barcelone), Tarik Asselah (Clichy), Adolfo F Attili (Rome), Johan Auwerx (Lausanne), Charles Balabaud (Bordeaux), Jean-Claude Barbare (Amiens), Véronique Barbu (Paris), Margaret Bassendine (Newcastle), Antonio Benedetti (Ancone), Ulrich Beuers (Amsterdam), Montgomery Bissell (San Francisco), Jaime Bosch (Barcelone), James Boyer (New Haven), Jordi Bruix (Barcelone), Jean-François Cadranel (Creil), Jacqueline Capeau (Paris), Nicolas Carbonell (Paris), Martin C Carey (Boston), Xavier Causse (Orléans), Roger W Chapman (Oxford), Olivier Chazouillères (Paris), Nicolas Chignard (Paris), Christophe Corpechot (Paris), Aimery de Gramont (Paris), Françoise Degos (Clichy), Victor de Ledinghen (Bordeaux), Yves Deugnier (Rennes), Jacques Devière (Bruxelles), François Durand (Clichy), Shahin Emami (Paris), Claude Eugène (Poissy), Peter Ferenci (Vienne), Johan Fevery (Louvain), Peter Fickert (Graz), Jean-François Fléjou (Paris), Pere Gines (Barcelone), Pierre-Marie Girard (Paris), Odile Goria (Rouen), Olivier Goulet (Paris), Jérôme Guéchot (Paris), Antoine Hadengue (Genêve), Dieter Haüssinger (Düsseldorf), Alan F Hofmann (San Diego), Pietro Invernizzi (Milan), Harry LA Janssen (Rotterdam), Neil Kaplowitz (Los Angeles), François Lacaine (Paris), Karine Lacombe (Paris), Didier Lebrec (Clichy), Sophie Lotersztajn (Créteil), Ariane Mallat (Créteil), Marco Marzioni (Ancone), Andrew Mason (Edmonton), Michèle Maurice (Paris), Juan F Medina (Pamplone), Yves Menu (Paris), Giorgina Mieli-Vergani (Londres), Richard Moreau (Paris), Antonio Moschetta (Bari), Sylvie Naveau (Clamart), Albert Pares (Barcelone), Gustav Paumgartner (München), Arnaud Pauwels (Paris), Dominique Pessayre (Paris), Massimo Pinzani (Florence), Jesus Prieto (Pamplone), Eric Raymond (Clichy), Mario Rizetto (Turin), Cecilia Rodriguez (Lisbonne), Jean Rosenbaum (Bordeaux), Olivier Rosmorduc (Paris), Didier Samuel (Villejuif), Daniel Shouval (Jerusalem), Olivier Soubrane (Paris), Mario Strazzabosco (New Haven), Claudio Tiribelli (Trieste), Thierry Tordjmann (Orsay), Michaël Trauner (Graz), Christian Trautwein (Aachen), Christian Trepo (Lyon), Jean-Michel Tubiana (Paris), Henk van Buuren (Rotterdam), Karel J van Erpecum (Utrecht), Michel Vaubourdolle (Paris), Dominique Wendum (Paris), Fabien Zoulim (Lyon), Jessica Zucman-Rossi (Paris) Clinics and Research in Hepatology and Gastroenterology (ISSN 2210-7401) 2014 (volume 38), monthly publication: 6 issues. France : 526 euros (TTC). See complete rates on www.elsevier-masson.fr/revue/CLINRE. Address order and payment to Elsevier Masson SAS, Service Abonnements, 62, rue Camille-Desmoulins, 92442 Issy-les-Moulineaux cedex: payment by check or credit card (CB, EuroCard, MasterCard or Visa: indicate no, and expiration date); La Banque Postale, centre de Paris, n° RIB 20041 00001 1904540H020 95. Subscriptions begin 4 weeks after receipt of payment and start with the rst issue of the calendar year. Back issues and volumes are available from the publisher. Claims for missing issues should be made within 6 months of publication. Includes air delivery. Journal manager – Pierre Vautherin. Tél. : + 33 (0)1 71 16 54 18. Fax : + 33 (0)1 71 16 51 67 - E-mail : [email protected] Sales Director - Marie-Pierre Cancel. Tél. : +33 (0)1 71 16 51 09. E-mail : [email protected] Advertising – Alexandra Bernard-Boudal. Tél. : + 33 (0)1 71 16 51 11. E-mail : [email protected] Site web : www.em-communication-sante.com Subscriptions – Tél. : + 33 (0)1 71 16 55 99. Fax : + 33 (0)1 71 16 55 77. http://www.em-consulte.com/infos Publisher – Freya Weise – Tél. : + 33 (0)1 71 16 53 58. Fax : + 33 (0)1 71 16 51 84. E-mail : [email protected] General manager and publishing director – Daniel Rodriguez Author enquiries For enquiries relating to the submission of articles (including electronic submission where available) please visit Elsevier’s Author Gateway at http://authors. elsevier.com. The Author Gateway also provides the facility to track accepted articles and set up e-mail alerts to inform you of when an article’s status has changed, as well as detailed artwork guidelines, copyright information, frequently asked questions and more. Contact details for questions arising after acceptance of an article, especially those relating to proofs, are provided after registration of an article for publication. Subscription conditions, instructions to authors, the contents of each issue as well as the abstracts of the articles published in clinical Research in hepatology and Gastroenterology are available on the website of Elsevier Masson: http://www.em-consulte.com/produit/clinre Printed in France by Jouve, 53101 Mayenne CPPAP : 0317 T 81435. Dépôt légal à parution ISSN 2210-7401 CLINICS AND RESEARCH IN HEPATOLOGY AND GASTROENTEROLOGY

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Editor in ChiefRaoul Poupon (Paris)

Associate EditorsLionel Arrivé (Paris), Charles Balabaud (Bordeaux), François Ballet (Paris), Yvon Calmus (Paris), Massimo Conio (Genoa), Dominique Debray (Paris), Christèle Desbois-Mouthon (Paris), Christian Florent (Paris), Chantal Housset (Paris), Mehdi Karoui (Paris), Patrick Marcellin (Clichy), Vlad Ratziu (Paris), Marika Rudler (Paris), Lawrence Serfaty (Paris), Julien Taieb (Paris), Christian Trépo (Lyon)

Consultant EditorsPaulette Bioulac-Sage (Bordeaux), Christian Bréchot (Paris), Fabrice Carrat (Paris), Daniel Dhumeaux (Paris), Serge Erlinger (Aix-en-Provence), Jean-Michel Pawlotsky (Créteil), Thierry Poynard (Paris), Kristina Schoonjans (Lausanne)

Editorial assistantYves Chrétien (Paris)

CopyeditingDale Roche-Lebrec (Paris), Gerald Pope (Rennes), Sharyn Wong (Little Rock)

Editorial BoardDomenico Alvaro (Rome), Tony Andréani (Paris), Mario Angelico (Rome), Vicente Arroyo (Barcelone), Tarik Asselah (Clichy), Adolfo F Attili (Rome), Johan Auwerx (Lausanne), Charles Balabaud (Bordeaux), Jean-Claude Barbare (Amiens), Véronique Barbu (Paris), Margaret Bassendine (Newcastle), Antonio Benedetti (Ancone), Ulrich Beuers (Amsterdam), Montgomery Bissell

(San Francisco), Jaime Bosch (Barcelone), James Boyer (New Haven), Jordi Bruix (Barcelone), Jean-François Cadranel (Creil), Jacqueline Capeau (Paris), Nicolas Carbonell (Paris), Martin C Carey (Boston), Xavier Causse (Orléans), Roger W Chapman (Oxford), Olivier Chazouillères (Paris), Nicolas Chignard (Paris), Christophe Corpechot (Paris), Aimery de Gramont (Paris), Françoise Degos (Clichy), Victor de Ledinghen (Bordeaux), Yves Deugnier (Rennes), Jacques Devière (Bruxelles), François Durand (Clichy), Shahin Emami (Paris), Claude Eugène (Poissy), Peter Ferenci (Vienne), Johan Fevery (Louvain), Peter Fickert (Graz), Jean-François Fléjou (Paris), Pere Gines (Barcelone), Pierre-Marie Girard (Paris), Odile Goria (Rouen), Olivier Goulet (Paris), Jérôme Guéchot (Paris), Antoine Hadengue (Genêve), Dieter Haüssinger (Düsseldorf), Alan F Hofmann (San Diego), Pietro Invernizzi (Milan), Harry LA Janssen (Rotterdam), Neil Kaplowitz (Los Angeles), François Lacaine (Paris), Karine Lacombe (Paris), Didier Lebrec (Clichy), Sophie Lotersztajn (Créteil), Ariane Mallat (Créteil), Marco Marzioni (Ancone), Andrew Mason (Edmonton), Michèle Maurice (Paris), Juan F Medina (Pamplone), Yves Menu (Paris), Giorgina Mieli-Vergani (Londres), Richard Moreau (Paris), Antonio Moschetta (Bari), Sylvie Naveau (Clamart), Albert Pares (Barcelone), Gustav Paumgartner (München), Arnaud Pauwels (Paris), Dominique Pessayre (Paris), Massimo Pinzani (Florence), Jesus Prieto (Pamplone), Eric Raymond (Clichy), Mario Rizetto (Turin), Cecilia Rodriguez (Lisbonne), Jean Rosenbaum (Bordeaux), Olivier Rosmorduc (Paris), Didier Samuel (Villejuif), Daniel Shouval (Jerusalem), Olivier Soubrane (Paris), Mario Strazzabosco (New Haven), Claudio Tiribelli (Trieste), Thierry Tordjmann (Orsay), Michaël Trauner (Graz), Christian Trautwein (Aachen), Christian Trepo (Lyon), Jean-Michel Tubiana (Paris), Henk van Buuren (Rotterdam), Karel J van Erpecum (Utrecht), Michel Vaubourdolle (Paris), Dominique Wendum (Paris), Fabien Zoulim (Lyon), Jessica Zucman-Rossi (Paris)

Clinics and Research in Hepatology and Gastroenterology (ISSN 2210-7401 ) 2014 ( volume 38 ), monthly publication: 6 issues. France : 526 euros(TTC). See complete rates on www.elsevier-masson.fr/revue/CLINRE.Address order and payment to Elsevier Masson SAS, Service Abonnements, 62, rue Camille-Desmoulins, 92442 Issy-les-Moulineaux cedex: payment by check or credit card (CB, EuroCard, MasterCard or Visa: indicate no, and expiration date); La Banque Postale, centre de Paris, n° RIB 20041 00001 1904540H020 95.Subscriptions begin 4 weeks after receipt of payment and start with the fi rst issue of the calendar year. Back issues and volumes are available from the publisher. Claims for missing issues should be made within 6 months of publication. Includes air delivery.

Journal manager – Pierre Vautherin. Tél. : + 33 (0)1 71 16 54 18. Fax : + 33 (0)1 71 16 51 67 - E-mail : [email protected] Director - Marie-Pierre Cancel. Tél. : +33 (0)1 71 16 51 09. E-mail : [email protected] – Alexandra Bernard-Boudal. Tél. : + 33 (0)1 71 16 51 11. E-mail : [email protected] web : www.em-communication-sante.comSubscriptions – Tél. : + 33 (0)1 71 16 55 99. Fax : + 33 (0)1 71 16 55 77. http://www.em-consulte.com/infos Publisher – Freya Weise – Tél. : + 33 (0)1 71 16 53 58. Fax : + 33 (0)1 71 16 51 84. E-mail : [email protected] manager and publishing director – Daniel Rodriguez

Author enquiriesFor enquiries relating to the submission of articles (including electronic submission where available) please visit Elsevier’s Author Gateway at http://authors.elsevier.com. The Author Gateway also provides the facility to track accepted articles and set up e-mail alerts to inform you of when an article’s status has changed, as well as detailed artwork guidelines, copyright information, frequently asked questions and more. Contact details for questions arising after acceptance of an article, especially those relating to proofs, are provided after registration of an article for publication.Subscription conditions, instructions to authors, the contents of each issue as well as the abstracts of the articles published in clinical Research in hepatology and Gastroenterology are available on the website of Elsevier Masson: http://www.em-consulte.com/produit/clinre

Printed in France by Jouve, 53101 MayenneCPPAP : 0317 T 81435. Dépôt légal à parutionISSN 2210-7401

CLINICS AND RESEARCH IN HEPATOLOGY AND GASTROENTEROLOGY