Checklist of Plants and Animals at the Bucknell University

Chillisquaque Creek Natural Area

Based on field observations, collections, and identifications by

Warren Abrahamson Donald Dearborn

David DeLuca Kerry Givens Duane Griffin Wayne Herr Ann Johnson Mark Larson

Wayne McDiffett James Mixon Ron Pursell

William Raschi DeeAnn Reeder

Allen Schweinsberg Cherri Smith

Tristan Stayton Karen Tarnow Lisa Withrow

Compiled by

Warren Abrahamson

Last update: September 5, 2006

- NOTE -

Habitat Key for location of mosses and herbaceous species: F better-drained field areas G grassy swale W woods P ponds (including temporary ponds) S streamside or drainage ditches

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Mosses Amblystegiaceae Amblystegium riparium Aquatic moss W Amblystegium varium Amblystegium moss W Brachytheciaceae Brachythecium oxycladon Cedar moss W Eurynchium hians Eurynchium moss W Climaciaceae Climacium americanum Tree moss W Dicraniaceae Dicranella heteromalla Silky forklet moss W Entodonaceae Entodon seductrix Seductive entodon moss W Fissidentraceae Fissidens taxifolius Flat fork moss W Hypnaceae Callicladium haldianum Callicladium moss W Hypnum curvifolium Curveleaf hypnum moss G Hypnum fertile Hypnum moss W Hypnum imponens Golden feather moss W Hypnum lindbergii Lindberg’s hypnum moss W Hypnum pallescens Hypnum moss W Platygyrium repens Platygyrium moss W Leskeaceae Anomodon attenuatus Tree apron moss W Hapocladium microphyllum W Leskea gracilescens Leskea moss W Leskea polycarpa Leakea moss W Thuidium delicatulum Common fern moss W Mniaceae Mnium cuspidatum Calcareous moss W Plagiotheciaceae Plagiothecium latebricola Plagiothecium moss W Polytrichaceae Atrichum angustatum Lesser smooth cap moss W Pogonatum pensylvanicum Hair cap moss S

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Herbaceous Plants Lemnaceae (Duckweed family) Lemna minor Duckweed P Sparganiaceae (Bur-reed family) Sparganium americanum Common bur-reed P Alismataceae (Arrowhead family) Alisma triviale Water plantain P Sagittaria latifolia Broad-leaved arrowhead P Gramineae or Poaceae (Grass family) Agrostis alba Redtop G Alopecurus spp. Foxtail G Bromus japonicus Japanese brome grass F Lolium perenne English rye grass F Panicum virgatum Switch grass G Cyperaceae (Sedge family) Carex flava Sedge G Carex lupulina Hop sedge G Carex scoparia Broom sedge G Carex vulpinoidea Sedge G Eleocharis acicularis Spike rush G Juncaceae (Rush family) Juncus effusus Soft rush C Juncus tenuis Path rush F Araceae (Arum family) Acorus calamus Sweetflag G,P Arisaema dracontium Green dragon W,S Arisaema triphyllum Jack-in-the-pulpit W Symplocarpis foetidus Skunk cabbage S Liliaceae (Lily family) Allium canadense Wild garlic W Allium tricoccum Wild leek W Clintonia umbellata White clintonia W Erythronium americanum Trout lily W Lilium canadense Canada lily F,G,W Polygonatum biflorum Solomon’s seal W Smilacina racemosa False Solomon’s seal W Trillium cernuum Nodding trillium W Uvularia sessilifolia Wild oats W Veratrum viride False hellebore W,S

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Iridaceae (Iris family) Iris versicolor Larger blue flag S Orchidaceae (Orchid family) Epipactus helleborine Helleborine S Aristolochiaceae (Birthwort family) Asarum canadense Wild ginger W Polygonaceae (Buckwheat family) Polygonum cespitosum Long-bristled smartwood G,S Polygonum sagittatum Arrow-leaved tear thumb G,S Tovara virginiana Virginia knotweed W Chenopodiaceae (Goosefoot family) Chenopodium album Lamb’s-quarters F Urticaceae (Nettle family) Boehmeria cylindrica False nettle G Laportea canadensis Wood nettle W,S Pilea pumila Clearweed W,S Urtica dioica Stinging nettle W Portulacaceae (Purslane family) Claytonia virginica Spring beauty W Plantaginaceae (Plantain family) Plantago lanceolata English plantain F Caryophyllaceae (Pink family) Dianthus ameria Deptford pink F Ranunculaceae (Buttercup family) Anemone quinquefolia Wood anemone W Clematis virginiana Virgin’s bower W Hepatica americana Round-leaved hepatica W Ranunculus abortivus Kidney-leaved buttercup W Ranunculus acris Common buttercup P Thalictrum polygamum Tall meadow-rue W Berberidaceae (Barberry family) Caulophyllum thalictroides Blue cohosh W Podophyllum peltatum May apple W Papaveraceae (Poppy family) Dicentra cucullaria Dutchman’s-breeches W Sanquinaria canadensis Bloodroot W

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Brassicaceae or Cruciferae (Mustard family) Capsella bursa-pastoris Shepherd’s purse F Dentaria laciniata Cut-leaved toothwort W Crassulaceae (Sedum family) Sedum telephium Orpine W Rosaceae (Rose family) Agrimonia sp. Agrimony W Fragaria vesca Wood strawberry F,W Fragaria virginiana Common strawberry F Potentilla recta Rough-fruited cinquefoil F Potentilla simplex Common cinquefoil F Fabaceae or Leguminosae (Legume family) Desmodium paniculatum Panicled tick-trefoil F Trifolium pratense Red clover F Oxalidaceae (Wood-sorrel family) Oxalis europaea Yellow wood-sorrel F Geraniaceae (Geranium family) Geranium maculatum Wild geranium W Polygalaceae (Milkwort family) Polygala sanquinea Field milkwort F Limnanthaceae (False Mermaid family) Floerka proserpinacoides False mermaid W Balsaminaceae (Touch-me-not family) Impatiens capensis Spotted jewelweed G,S Impatiens pallida Pale jewelweed S Malvaceae (Mallow family) Abutilon theophrasti Velvet leaf F Clusiaceae or Guttiferae (St. Johnswort family) Hypericum perforatum Common St. John’s wort F Violaceae (Violet family) Viola cucullata Marsh blue violet W Viola papilionea Common blue violet W Viola pubescens Downy yellow violet W Viola striata Cream violet W

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Lythraceae (Loosestrife family) Lysimachia nummularia Moneywort W,P Lysimachia quadrifolia Fringed loosestrife P,S Decodon verticillatus Swamp loosestrife P Cuphea petiolata Clammy cuphea F Onagraceae (Evening Primrose family) Circaea lutetiana Enchanter’s nightshade W Oenothera biennis Evening primrose F Araliaceae (Ginseng family) Panax trifolius Dwarf ginseng W Apiaceae or Umbelliferae (Carrot family) Daucus carota Queen Anne’s lace F Osmorhiza claytoni Sweet cicely W Sium suave Water parsnip P Zizia aurea Golden Alexanders W Apocynaceae (Dogbane family) Apocynum medium Intermediate dogbane F Asclepiadaceae (Milkweed family) Asclepias incarnata Swamp milkweed F,G Asclepias syriaca Common milkweed F Convolvulaceae (Morning Glory family) Convolvulus arvensis Field bindweed F Cuscuta gronovii Dodder G,S Ipomoea pandurata Wild potato vine F Polemoniaceae (Phlox family) Phlox divaricata Blue phlox W Phlox maculata Wild sweet William W Boraginaceae (Forget-me-not family)) Lythospermum arvense Corn gromwell F Mertensia virginica Virginia bluebells W,S Myostis scorpioides True forget-me-not W,S Verbenaceae (Vervian family) Verbena hastata Blue vervain F,S Verbena urticifolia White vervain S Lamiaceae or Labiatae (Mint family) Leonurus cardiaca Motherwort F Lycopus americanus Cut-leaved water horehound G

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Mentha arvenis Wild mint F,G Monarda clinopardia White bergamot S,W Prunella vulgaris Self-heal F Pycnanthemum tenuifolium Narrow-leaved mountain mint F Scutellaria lateriflora Mad-dog skullcap W Stachys sp. Hedge-needle sp. F,G Solanaceae (Nightshade family) Physalis subglabrata Smooth ground cherry S Solanum carolinense Horse nettle F Solanum dulcamara Deadly nightshade S Scrophulariaceae (Snapdragon family) Chelone glabra Turtlehead W,P Linaria vulgaris Butter-and-eggs W,P Mimulus ringens Square-stemmed monkey flower F,G Rubiaceae (Bedstraw family) Galium sp. Bedstraw G,W Campanulaceae (Bluebell family) Lobelia spicata Pale spiked lobelia F Asteraceae or Compositae (Sunflower family) Achillea millefolium Yarrow F Aster divaricatus White wood aster W Aster puniceus Purple-stemmed aster F,G Aster vimineus Small white aster F Bidens frondosa Beggar-tick W Eupatorium maculatum Spotted Joe-Pye-Weed F,G Eupatorium perfoliatum Boneset F Eupatorium purpureum Sweet Joe-Pye-Weed W Euthamia graminifolia Grass-leaved goldenrod F Galinsoga ciliate Galinsoga F Helenium autumnale Sneezewood F,G Helianthus decapetalus Thin-leaved sunflower S Senecio aureus Golden ragwort W Solidago altissima Tall goldenrod F Solidago canadensis Canada goldenrod F,G Solidago gigantea Late goldenrod F,G Solidago juncea Early goldenrod F Solidago rugosa Rough-stemmed goldenrod F Taraxacum officinale Common dandelion F,S

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Trees and Shrubs Pinaceae (Pine family) Pinus strobus White pine Magnoliaceae (Magnolia family) Liriodendron tulipifera Tulip-poplar Lauraceae (Laurel family) Lindera benzoin Spicebush Sassafras albidum Sassafras Ulmaceae (Elm family) Celtis occidentalis Hackberry Ulmus americana American elm Ulmus rubra Slippery elm Salicaceae (Willow family) Salix sp. Willow Juglandaceae (Walnut family) Carya cordiformis Bitternut hickory Carya laciniosa Shellbark hickory Carya ovata Shagbark hickory Juglans nigra Black walnut Fagaceae (Beech family) Fagus grandifolia American beech Quercus alba White oak Quercus bicolor Swamp white oak Quercus palustris Pin oak Quercus rubra Northern red oak Betulaceae (Birch family) Alnus sp. Alder Betula nigra River birch Carpinus caroliniana American hornbeam Ostrya virginiana Eastern hop hornbeam Tiliaceae (Linden family) Tilia americana American basswood Rosaceae (Rose family) Crataegus sp. Hawthorn Prunus serotina Wild cherry Rosa multiflora Multiflora rose Rosa palustris Swamp rose Rubus allegheniensis Blackberry

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Rubus hispidus Bristly dewberry Rubus occidentalis Black raspberry Spiraea latifolia Meadowsweet Fabaceae or Leguminosae (Pea family) Robinia pseudoacacia Black locust Cornaceae (Dogwood family) Cornus amomum Silky dogwood Cornus alternifolia Alternate-leaved dogwood Cornus florida Flowering dogwood Nyssaceae (Tupelo family) Nyssa sylvatica v. biflora Black or swamp tupelo Aceraceae (Maple family) Acer negundo Box elder Acer nigrum Black maple Acer rubrum Red maple Acer saccharinum Silver maple Anacardiaceae (Cashew family) Rhus typhina Staghorn sumac Toxicodendron radicans Poison ivy Oleaceae (Olive family) Fraxinus nigra Black ash Fraxinus pennsylvanica v. subintegerrima Green ash Rubiaceae (Madder family) Cephalanathus occidentalis Buttonbush Caprifoliaceae (Honeysuckle family) Sambucus canadensis Common elderberry Viburnum cassinoides Northern wild-raisin Viburnum prunifolium Smooth blackhaw Viburnum recognitum Northern arrowwood

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Butterflies Satyridae (Wood nymphs and satyrs) Cercyonis pegala Blue-eyed grayling Euptychia cymela Little wood satyr Lethe eurydice Eyed brown Danaidae (Monarchs) Danaus plexippus Monarch Nymphalidae (Brush-footed) Boloria toddi Meadow fritillary Euphydryas phaeton Baltimore checkerspot Euptoieta claudia Variegated fritillary Limentis archippus Viceroy Limentis arthemis Red-spotted purple Nymphalis antiopa Mourning cloak Phycoides tharos Pearl crescent Polygonia comma Comma butterfly Polygonia interrogationis Question-mark butterfly Speyeria cybele Great spangled fritillary Vanessa atalanta Red admiral Lycaenidae (Gossamer-winged) Everes comyntas Eastern tailed blue Lycaena thoe Bronze copper Lycaenopsis argiolus Spring azure Papilionidae (Swallowtails) Papilio glaucus Tiger swallow tail Papilio polyxenes Black swallow tail Pieridae (Whites and sulphurs) Colias eurytheme Alfalfa butterfly Colias philodice Clouded sulphur Pieris rapae Cabbage butterfly Hesperiidae (Skippers) Ancyloxipha numitor Least skipper Epargyreus clarus Silver spotted skipper Pyrgus communis Checkered skipper Thorybes pylades Northern cloudy wing

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Fishes Catostomidae (Suckers) Hypentelium nigricans Hogsucker Cyprinidae (Carp) Campostoma anomalum Stone roller Esox americanus Redfin pickerel Notemigonus crysoleucas Golden shiner Notropis atherinoides Emerald shiner Notropis cornutus Common shiner Notropis hudsonius Spottail shiner Notropis spilopterus Spotfin shiner Pimephales notatus Bluntnose minnow Semotilus atromaculatus Creek chub Centrarchidae (Bass and sunfish) Lepomis gibbosus Pumpkinseed sunfish Lepomis marcochirus Bluegill Percidae (Perch) Boleosama nigrum Johnny darter

Amphibians Ambystomidae (Mole salamanders) Ambystoma jeffersonianum Jefferson’s salamander Salamandridae (Newts) Notophthalamus viridescens Red-spotted newt Plethodontidae (Lungless salamanders) Eurycea bislineata Two-lined salamander Plethodon cinereus Red-backed salamander Plethodon wehrlei Wehrle’s salamander Bufonidae (Toads) Bufo americanus American toad Bufo fowleri Fowler’s toad Hylidae (Treefrogs, cricket frogs, and chorus frogs) Pseudacris crucifer Spring peeper Pseudacris triseriata feriarum Upland chorus frog Ranidae (True frogs) Rana catesbeiana Bullfrog Rana clamitans melanota Green frog

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Rana pipiens Northern leopard frog

Reptiles Chelydridae (Snapping turtles) Chelydra serpentine Snapping turtle Kinosternidae (Musk and mud turtles) Sternotherus odoratus Stinkpot Emydidae (Box and water turtles) Chrysemys picta picta Eastern painted turtle Clemmys guttata Spotted turtle Glyptemys insculpta Wood turtle Terrapene carolina Box turtle Colubridae (Colubrid snakes) Nerodia sipedon Northern water snake Thamnophis sirtalis sirtalis Eastern garter snake

Mammals Didelphidae (opossums) Didelphis virginiana Virginia opossum Leporidae (hares and cottontail rabbits) Sylvilagus floridanus Eastern cottontail Sciuridae (squirrels and chipmunks) Tamias striatus Eastern chipmunk Sciurus carolinensis Eastern gray squirrel Cricetidae (voles and other cricetine rodents) Peromyscus maniculatus North American deer mouse Peromyscus leucopus White footed mouse Microtus pennsylvanicus Meadow vole Ondatra zibethicus Common muskrat Procyonidae (Raccoons and coatis) Procyon lotor Raccoon Mustelidae (Weasels) Mustela frenata Longtail weasel Mephitidae (skunks) Mephitis mephitis Striped skunk

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Cervidae (Deer) Odocoileus virginianus White-tailed deer

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Birds Anatidae (Ducks, geese, and swans) Branta canadensis Canada goose Chen hyperborean Snow goose Aix sponsa Wood duck Anas crecca Green-winged teal Anas discors Blue-winged teal Anas platyrhynchos Mallard Anas rubripes American black duck Phasianidae (Partridges, grouse, turkeys, and old world quail) Meleagris gallopavo Wild turkey Phasianus colchicus Ring-necked pheasant Odontophoridae (New world quail) Colinus virginianus Northern bobwhite Ardeidae (Bitterns, heron and allies) Ardea herodias Great blue heron Butorides virescens Green heron Cathartidae (New world vultures) Cathartes aura Turkey vulture Accipitridae (Hawks, kites, eagles, and allies)

Accipiter striatus Sharp-shinned hawk Buteo jamaicensis Red-tailed hawk Buteo platypterus Broad-winged hawk Pandion haliaetus Osprey Falconidae (Caracaras and falcons) Falco sparverius American kestrel Rallidae (Rails, gallinules, and coots) Fulica americana American coot Porzana carolina Sora Rallus limicola Virginia rail Charadriidae (Lapwings and plovers) Charadruis vociferous Killdeer Scolopacidae (Sandpipers, phalaropes, and allies) Calidris melanotos Pectoral sandpiper Gallinago delicata Wilson’s snipe Scolopax minor American woodcock Tringa solitaria Solitary sandpiper

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Tringa flavipes Lesser yellow-legs Laridae (Skuas, gulls, terns, and skimmers) Larus philadelphia Bonaparte’s gull Columbidae (Pigeons and doves) Columba livia Rock pigeon Zenaida macroura Mourning dove Cuculidae (Cuckoos, roadrunners, and anis) Coccyzus americanus Yellow-billed cuckoo Coccyzus erythrophthalmus Black-billed cuckoo Strigidae (Typical owls) Bubo virginianus Great horned owl Apodidae (Swifts) Chaetura pelagica Chimney swift Trochilidae (Hummingbirds) Archilochus coubris Ruby-throated hummingbird Alcedinidae (Kingfishers) Ceryle alcyon Belted kingfisher Picidae (Woodpeckers and allies) Colaptes auratus Northern flicker Dendrocopus pubescens Downy woodpecker Dendrocopus villosus Hairy woodpecker Melanerpes carolinus Red-bellied woodpecker Melanerpus erythrocephalus Red-headed woodpecker Sphyrapicus varius Yellow-bellied sapsucker Tyrannidae (Tyrant flycatchers) Contopus virens Eastern wood-pewee Empidonax alnorum Alder flycatcher Empidonax traillii Willow flycatcher Empidonax minimus Least flycatcher Myiarchus crinitus Great crested flycatcher Sayornis phoebe Eastern phoebe Vireonidae (Vireos) Vireo gilvus Warbling vireo Vireo griseus White-eyed vireo Vireo olivaceus Red-eyed vireo Vireo solitarius Blue-headed vireo Corvidae (Jays and crows)

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Corvus brachyrhynchos American crow Corvus ossifragus Fish crow Cyanocitta cristata Blue jay Hirundinidae (Swallows) Hirundo rustica Barn swallow Stelgidopteryx serripennis Northern rough-winged swallow Tachycineta bicolor Tree swallow Paridae (Chickadees and Titmice) Poecile atricapillus Black-capped chickadee Baeolophus bicolor Tufted titmouse Sittidae (Nuthatches) Sitta carolinensis White-breasted nuthatch Certhiidae (Creepers) Certhia americana Brown creeper Troglodytidae (Wrens) Thyrothorus ludovicianus Carolina wren Troglodytes aedon House wren Regulidae (Kinglets) Regulus calendula Ruby-crowned kinglet Sylviidae (Old world warblers and gnatcatchers)

Poliotila caerulea Blue-gray gnatcatcher Turdidae (Thrushes) Catharus fuscescens Veery Hylocichla mustelina Wood thrush Sialia sialis Eastern bluebird Turdus migratorius American robin Mimidae (Mockingbirds and thrashers) Dumetella carolinensis Gray catbird Mimus polyglottos Northern mockingbird Toxostoma rufum Brown thrasher Sturnidae (Starlings) Sturnus vulgaris European starling Bombycillidae (Waxwings) Bombycilla cedrorum Cedar waxwing Parulidae (Wood warblers)

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Dendroica castanea Bay-breasted warbler Dendroica coronota Yellow-rumped warbler Dendroica fusca Blackburnian warbler Dendroica magnolia Magnolia warbler Dendroica palmarum Palm warbler Dendroica pensylvanica Chestnut-sided warbler Dendroica petechia Yellow warbler Dendroica striata Blackpoll warbler Dendroica tigrina Cape May warbler Dendroica virens Black-throated green warbler Geothlypis trichas Common yellowthroat Helmitheros vermivorum Worm-eating warbler Icteria virens Yellow-breasted chat Mniotilta varia Black-and-white warbler Parula americana Northern parula Seiurus aurocapilla Ovenbird Seiurus motacilla Louisiana waterthrush Seiurus noveboracensis Northern waterthrush Setophaga ruticilla American redstart Vermivora ‘lawrencei’ (hybrid: V. chrysoptera x V. pinus)

“Lawrence’s” warbler

Vermivora chrystoptera Golden-winged warbler Vermivora peregrina Tennessee warbler Vermivora pinus Blue-winged warbler Vermivora ruficapilla Nashville warbler Thaupidae (Tanagers) Piranga olivacea Scarlet tanager Emberizidae (Emberizids – towhees, sparrows, buntings, juncos, and longspurs) Junco hyemalis Dark-eyed junco Melospiza georgiana Swamp sparrow Melospiza melodia Song sparrow Passerella illiaca Fox sparrow Pipilo erythrophthalmus Eastern towhee Spizella passerina Chipping sparrow Spizella pusilla Field sparrow Zonotrichia albicollis White-throated sparrow Zonotrichia leucophrys White-crowned sparrow Cardinalidae (Cardinals) Cardinalis cardinalis Northern cardinal Passerina cyanea Indigo bunting Pheucticus ludovicianus Rose-breasted grosbeak Icteridae (Blackbirds) Agelaius phoeniceus Red-winged blackbird Euphagus carolinus Rusty blackbird

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Icterus galbula Baltimore oriole Icterus spurius Orchard oriole Molothrus ater Brown-headed cowbird Quiscalus quiscula Common grackle Fringillidae (Fringilline and cardueline finches and allies) Carduelis tristis American goldfinch Carpodacus mexicanus House finch Carpodacus purpureus Purple finch Passerudae (Old world sparrows) Passer domesticus House sparrow